(DRAFT) - Taxonomy
                              Species CHUB, TUI, HUTTON
                                Species Id ESIS251014
                                   Date 14 MAR 96



TAXONOMY

NAME - CHUB, TUI, HUTTON OTHER COMMON NAMES - CHUB, TUI, HUTTON; ROACH; CHUB, HUTTON SPRING;CHUB, TUI, HUTTON SPRING; CHUB, TUI, HUTTON SPRINGS; OREGON TUI CHUB OF HUTTON SPRING; CHUB, TUI, OREGON and HUTTON SPRING ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - CHORDATA, CLASS AND SUBCLASS - OSTEICHTHYES, ORDER AND SUBORDER - CYPRINIFORMES, FAMILY AND SUBFAMILY - CYPRINIDAE, GENUS AND SUBGENUS - GILA, SPECIES AND SSP - BICOLOR, SSP. SCIENTIFIC NAME - GILA BICOLOR SSP. AUTHORITY - TAXONOMY REFERENCES - COMMENTS ON TAXONOMY - Hutton Tui Chub Gila bicolor ssp. (Girard, 1857) KINGDOM: Animal GROUP: Fish PHYLUM: Chordata CLASS: Osteichthyes ORDER: Cypriniformes FAMILY: Cyprinidae The morphological description below is taken from Bills (03) and the laboratory and field notes of C.E. Bond (18). The Hutton tui chub is an undescribed subspecies of Gila bicolor (Girard) 1857. The Hutton tui chub is robust, with the greatest depth of body immediately behind the head. The head is large, its length 0.30 to 0.31 m standard length (SL) and the depth 0.20 m (SL). The eye appears large and is usually slightly more than 0.06 m (SL). Fin rays are as follows: dorsal, 8; anal, 8; pelvic, 9-10; pectoral, 15-18. Lateral line scales are about 54. Other counts are: gill rakers, 10-16; vertebrae, 37-42 (usually 41); pharyngeal teeth, 5-4. The subspecies has the heaviest, strongest pharyngeal teeth of any Gila bicolor known. Pigmentation usually dark, brassy. Mouth lining and tongue purplish. Although the synonymy of Gila bicolor is complex, the Hutton tui chub has been known by only a few names so no attempt will be made to Taxonomy - 1 (DRAFT) - Taxonomy Species CHUB, TUI, HUTTON Species Id ESIS251014 Date 14 MAR 96 give an extended Gila bicolor synonymy. Following is a chronology of the names that have applied, directly or indirectly, to the Hutton tui chub. Snyder, in 1908 (14), described the formes from XL Spring, Chewaucan River, Ana River, Bridge Creek, Warner Creek, and the spring at Alkali Lake (=Hutton Spring) and named them as Rutilus oregonensis. In 1913 Fowler (07) applied the name Rutilus formosus to some of the forms, but not, apparently to those from Alkali Lake. Snyder (15) subsequently in 1917 used the name Siphateles oregonensis, and in 1935 Schultz and DeLacy (13) synonymized oregonensis with bicolor and regarded it to have subspecific status, so the name became Siphateles bicolor oregonensis. Hubbs and Miller (08) considered bicolor to be synonym of obesus and used the name Siphateles obesus oregonensis in 1942. Bailey and Uyeno, in 1964 (01), merged Siphateles with Gila so that the name became Gila bicolor oregonensis. Bills (03) showed in 1977 that the Hutton tui chub is distinct on at least the subspecific level from the other forms of the nominal G. b. oregonsis. The form has no subspecific name as yet, so it is called Gila bicolor ssp. in current works. Thus the most significant synonyms used in the past 50 years are the following: 1. for the species: Siphateles obesus Girard, 1857 Siphateles bicolor Girard, 1857 Gila bicolor Girard, 1857 Siphateles oregonensis Snyder, 1908 2. for the subspecies: Siphateles bicolor oregonensis Snyder, 1908 S. obesus oregonensis Snyder, 1908 Gila bicolor oregonensis Snyder, 1908 Other common names include; roach, Hutton Spring chub, Oregon tui chub of Hutton Spring, Hutton Spring tui chub, and Hutton Springs tui chub. Specimens in the fish collection, Dept. of Fisheries and Wildlife, Oregon State University bear the following catalog numbers: 4208, 10203, 7756, 4209, 5005, 5158, 5136. Photographs are in F. Bills' thesis (03) and in Deacon, et al. 1979 (05). Taxonomy - 2
                                  (DRAFT) - Status
                              Species CHUB, TUI, HUTTON
                                Species Id ESIS251014
                                   Date 14 MAR 96



STATUS

Coded Status T: Federal Threatened COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The Hutton tui chub (Gila bicolor ssp.) has been designated as Threatened pursuant to the Endangered Species Act of 1973 (50 CFR 17.11; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The subspecies has this status wherever found including the State of Oregon. Special rules concerning "take" for this subspecies can be found in 50 CFR 17.44(j). This subspecies is protected by the Lacey Act (P.L. 97-79, as amended; 16 U.S.C. 3371 et seq.) which makes it unlawful to import, export, transport, sell, receive, acquire, or purchase any wild animal (alive or dead including parts, products, eggs, or offspring): (1) in interstate or foreign commerce if taken, possessed, transported or sold in violation of any State law or regulation, or foreign law; or (2) if taken or possessed in violation of any U.S. law, treaty, or regulation or in violation of Indian tribal law. It is also unlawful to possess any wild animal (alive or dead including parts, products, eggs, and offspring) within the U.S. territorial or special maritime jurisdiction (as defined in 18 U.S.C. 7) that is taken, possessed, transported, or sold in violation of any State law or regulation, foreign law, or Indian tribal law. RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely to jeopardize the continued existence of the species or result in the destruction or adverse modification of Critical Habitat (50 CFR 402), and to utilize their authorities to carry out programs for the conservation of the species. STATE STATUSES AND LAWS: STATE: Oregon DESIGNATED STATUS: Threatened ADMINISTRATIVE AGENCY: Oregon Department of Fish and Wildlife STATE STATUTE: OR Admin. Rules, Chapt. 635, Div. 7; OR Rev. Status - 1 (DRAFT) - Status Species CHUB, TUI, HUTTON Species Id ESIS251014 Date 14 MAR 96 Stat. 496.002-498.029, 506.001-506.518, and 610.002-610.060. INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: The tui chub (Gila bicolor (subspecies)) is listed as threatened in the IUCN Red List of Threatened Animals (1986). ECONOMIC STATUSES: None. 84/05/17:49 FR 15099/15102 - Proposed Listing 85/03/28:50 FR 12302/12306 - Final Rule: Listed as Threatened Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC INLAND AQUATIC LAND USE - Herbaceous Rangeland Streams and Canals Nonforested Wetland Dry Salt Flats NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Riverine, upper perennial SB2 Riverine, upper perennial FL2 Palustrine OW0 Palustrine EM1 COMMENTS ON HABITAT ASSOCIATIONS - The Hutton tui chub lives in clear springs and outflow channels. The size of the springhole of Hutton Spring varies with excavations made by the owner. It has ranged from 20 feet to nearly 40 feet in diameter (03), and is about 15 feet deep in the center (17). The smaller spring hole is about 10 feet across and 2 feet deep (03). Hutton Spring is occupied in part by tules (Scirpus americanus) and the small spring is surrounded by water parsley (Oenanthe sp.). Other vegetation present includes sedge (Carex sp.), saltgrass (Distichlus sp.), and squirreltail (Sitanion hystrix) (11). The fish use the vegetation and whatever debris is present for cover. Some of the larger individuals use the deep spring hole as cover (19). The recorded water temperature is 64 deg. F. (17.7 deg. C.), during the summer (May - October) (12). The outflow from the springs form a small area of wetland adjacent to the sources. This is occupied by grasses, water parsley, and sedges. The springs are in a grassy rangeland bordered to the north and west by shrubby rangeland (See 09, for photograph) and to the east and south by the lake bed of Pluvial Alkali Lake. A low dry ridge with sagebrush is immediately south of the spring area. Elevation at the site is 4500 feet. Habitat Associations - 1
                                (DRAFT) - Food Habits
                              Species CHUB, TUI, HUTTON
                                Species Id ESIS251014
                                   Date 14 MAR 96



FOOD HABITS

TROPHIC LEVEL - OMNIVORE LIFESTAGE FOOD FOOD PART General Algae General Zooplankton General Arthropods General Molluscs Food Habits - 1
                         (DRAFT) - Environment Associations
                              Species CHUB, TUI, HUTTON
                                Species Id ESIS251014
                                   Date 14 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G G Aquatic Features: Springs [flowing] Environment Associations - 1
                               (DRAFT) - Life History
                              Species CHUB, TUI, HUTTON
                                Species Id ESIS251014
                                   Date 14 MAR 96



LIFE HISTORY

FOOD HABITS: Juvenile Hutton tui chubs feed on zooplankton of appropriate size, and apparently feed on small insects as well. At 50 to 60 mm in length, the fishes diet shifts toward plant material and they remain omnivorous, feeding on vegetation, insects, and gastropods (03,19). HOME RANGE/TERRITORY: Territoriality is not known in this subspecies. The habitat is so restricted that the concept of home range is probably not applicable (19). PERIODICITY: This fish has been observed as active diurnally (19, 02). MIGRATION PATTERNS: No migrations known or suspected because of restricted habitat (19). COVER/SHELTER REQUIREMENTS: Cover utilized by the Hutton tui chub includes Scirpus, Carex and Oenanthe as well as other aquatic or phreatophytic vegetation and floating boards (02,03). Some of the larger individuals use the deep spring hole as cover (19). REPRODUCTIVE SITE REQUIREMENTS: Reproduction has not been studied in this subspecies, but vegetation is the spawning substrate for typical Gila bicolor (10). Juveniles appear to use shallow margins as nursery areas (02). Spawning temperature is suspected to be 64 deg. F. (17.7 deg. C.) as that is the temperature of the spring water (12). REPRODUCTIVE CHARACTERISTICS: No details of reproduction are known, except that fish of 17 mm have been taken in late July (16), which infers that spawning probably takes place in May or June. See reference (10) for life history data for Gila bicolor. PARENTAL CARE: No parental care known or suspected. POPULATION BIOLOGY: Little is known of the population biology of this subspecies. The population has been estimated at about 450 individuals (07). Adult fish are predominantly females: 7 males to 33 females in Bills' (03) study. The largest specimen in the Oregon State University Fish Collection (055158) is 152 mm (SL) (12). SPECIES INTERRELATIONSHIPS: The main interrelationship with other species would be the use of the spring area SE of Hutton Spring by cattle. The fish feed Life History - 1 (DRAFT) - Life History Species CHUB, TUI, HUTTON Species Id ESIS251014 Date 14 MAR 96 on zooplankton, insects, gastropods, and plants. No instances of predation or parasitism have been observed (19). OTHER LIFE HISTORY DESCRIPTORS: None. Life History - 2
                           (DRAFT) - Management Practices
                              Species CHUB, TUI, HUTTON
                                Species Id ESIS251014
                                   Date 14 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Restricting/regulating human disturbance of populations Beneficial Developing/maintaining stream structures Beneficial Land Acquisition Beneficial Water Right Acquisition Beneficial Controlling pollution [thermal, chemical, physical] Beneficial Reforestation Beneficial Controlling/Removing Domestic Animals Adverse Climate Alteration Existing Climate Alteration Adverse Soil compaction by heavy equipment in mine areas Existing Soil compaction by heavy equipment in mine areas Adverse Siltation Existing Siltation Adverse Irrigating Existing Irrigating Adverse Dredging Existing Dredging Adverse Developing/maintaining stream bank vegetation Existing Developing/maintaining stream bank vegetation Adverse Environmental Contamination/Pollution Existing Environmental Contamination/Pollution Adverse Erosion Existing Erosion Adverse Grazing Existing Grazing COMMENTS ON MANAGEMENT PRACTICES - The isolation of the Hutton tui chub is due to the desiccation of pluvial Alkali Lake (14,09). Present status is in part a result of unrestricted access by cattle to "Three-eights" Spring and past access by cattle to Hutton Spring (07). Threats to the populations of tui chubs includes pumping of water from the springs which occurred in the past but is not occurring now (04,07), and contamination of ground water by dispersal of chemicals from a herbicide-manufacturing residue disposal site 1.75 miles south of Hutton Spring (07). Modification of the springs (via heavy equipment - thus in turn causing other problems such as siltation, erosion, vegetaion cover loss, water diversion and drawdown, etc.) had detrimental effects on the chub population. A possible threat is the establishment of a bombing range on Federal lands in the vicinity of Alkali Lake, but this is not considered a serious threat (07) because of the distance from the habitat. UNAPPROVED PLAN: At present there is no formal recovery plan for the Hutton tui chub. Management Practices - 1 (DRAFT) - Management Practices Species CHUB, TUI, HUTTON Species Id ESIS251014 Date 14 MAR 96 Recovery actions that are needed at present are: 1. Development of a land management agreement (including water right acqusition) with the land owner to protect the habitat and spring area. One problem is to prevent any further disturbance (i.e, siltation, erosion, water drawdown, and vegetation destruction) of the habitat, which occurs on a periodic basis, by the landowner and his back-hoe work in the spring for water control. Also, another problem is the vegetative damage, erosion, and sedimentation caused by cattle using the surrounding the habitat (other than removal of cattle from the area, a program to rehabilitate and revegetate the area damaged by cattle and back-hoe work). 2. Research is needed on: a) development of a formal taxonomic description for this species, b) food requirements, c) cover and shelter requirements, d) reproductive/spawning requirements, and e) general interactions with habitat and other species within this habitat. *3. Prevention of groundwater contamination (at present the major threat to this species is the contamination of surface and ground waters from a nearby toxic material site. The site contains thousands of 50 gallon drums with residue from DDE manufacture that were improperly disposed of. Signs of both groundwater and surface water contamination has occurred and also aerial contamination is occurring. Residues from the drums have entered the dry alkali soils of Alkali Lake and are easily distributed during high winds in which dust and the residue are dispersed through the air, speeding-up surface contamination. Management Practices - 2
                                   (DRAFT) - References
                                Species CHUB, TUI, HUTTON
                                  Species Id ESIS251014
                                      Date 14 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 Bailey, R.M., and T. Uyeno. 1964. Nomenclature of the blue chub and the tui chub, cyprinid fishes from western United States. Copiea 1964:238-239. 02 Bills, F.T. Field Notes. FTB75-13 and FTB76-3, On file at Dept. of Fisheries and Wildl., OR. State Univ. 03 Bills, F.T. 1978. Taxonomic status of the isolated populations of tui chub referred to as Gila bicolor oregonensis (Snyder). MS Thesis, OR State Univ. 121 pp. 04 Bond, C.E. 1974. Endangered plants and animals of Oregon: I, Fishes., OR Agric. Exp. Sta. Spec. Rept. 205. Pp. 1-9. 05 Deacon, J.E., G. Kobetich, J.D. Williams, S. Contreras, et al. 1979. Fishes of North America endangered, threatened, or of special concern: 1979. Fisheries 4(2):30-44. 06 Fowler, H.W. 1913. Some type-specimens of the American cyprinoid fishes of the genus Rutilus. Proc. Acad. Nat. Sci. Phil. Pp. 66-71. 07 Franzreb, K.E. 1985. Endangered and threatened wildlife and plants; determination of threatened status for Hutton tui chub and Foskett speckled dace. Fed. Reg. 50:12302-12306. 08 Hubbs, C.L., and R.R. Miller. 1942. Fish of Catlow Valley. Pg. 152, In: L.S. Cressman, Archeological researches of the northern Great Basin., Carn. Inst. Wash. Publ. No. 538. 09 Hubbs, C.L. and R.R. Miller. 1948. The zoological evidence/ correlation between fish distribution and hydrographic history in the desert basins of western United States. Bull. Univ. Utah 30(2) Pp. 17-166. 10 Moyle, P.B. 1976. Inland fishes of California., Univ. of Calif. Press. 405 pp. 11 Oregon Natural Heritage Program. 1985. Fish data. Portland, OR. 12 Oregon State University. 1986. Oregon fish data base; Hutton tui chub record. OR State Univ., Dept. of Fish. and Wildl., Corvallis. 13 Schultz, L.P., and A.C. DeLacy. 1935-36. Fishes of the American Northwest. Jour. Pan Pac. Res. Inst. X(4) Mid Pacific Magazine. 14 Snyder, J.O. 1908. Relationships of the fish fauna of the lakes of southeastern Oregon. Bull. U.S. Bur. Fish., vol. XXVII (1907): 69-102. 15 Snyder, J.O. 1917. The fishes of the Lahontan system of Nevada and northestern California. Bull. U.S. Bur. Fish. 35:33-86. 16 University of Michigan Museum of Zoology. Card catalog file. File No.: 130507, Ann Arbor, MI. 17 Waring, G.A. 1908. Geology and water resources of a portion of south-central Oregon. U.S. Geol. Surv. Water Supp. Pop. 220:1-86. 18 Bond, C.E. [n.d.]. Field notes. Oregon State Univ., Corvallis. 19 Bond, C.E. [n.d.]. Pers. obs. Oregon State Univ., Corvallis. ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 Bills, F.T. 1978. Taxonomic status of the isolated populations of References - 1 (DRAFT) - References Species CHUB, TUI, HUTTON Species Id ESIS251014 Date 14 MAR 96 tui chub referred to as Gila bicolor oregonensis (Snyder). MS Thesis, OR State Univ. 121 pp. 02 Bond, C.E. 1974. Endangered plants and animals of Oregon: I, Fishes., OR Agric. Exp. Sta. Spec. Rept. 205. Pp. 1-9. 03 Franzreb, K.E. 1985. Endangered and threatened wildlife and plants; determination of threatened status for Hutton tui chub and Foskett speckled dace. Fed. Reg. 50:12302-12306. 04 Hubbs, C.L., and R.R. Miller. 1948. The zoological evidence/ correlation between fish distribution and hydrographic history in the desert basins of western United States. Bull. Univ. Utah 30(2) Pp. 17-166. 05 Minckley, W.L., D.A. Hendrickson, and C.E. Bond. 1986. Geography of western North American freshwater fishes: description and relationships to intracontinental tectonism. Pp. 519-613. In: C.H. Hocutt and E.O. Wiley (eds). The zoogeography of North American freshwater fishes. John Wiley and Sons, Inc. 06 Oregon State University. 1986. Oregon fish data base; Hutton tui chub record. OR State Univ., Dept. of Fish. and Wildl., Corvallis. 07 Snyder, J.O. 1908. Relationships of the fish fauna of the lakes of southeastern Oregon. Bull. U.S. Bur. Fish., vol. XXVII (1907): 69-102. References - 2