(DRAFT) - Taxonomy
                               Species CATFISH, YAQUI
                                Species Id ESIS252013
                                   Date 14 MAR 96



TAXONOMY

NAME - CATFISH, YAQUI OTHER COMMON NAMES - CATFISH and YAQUI ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - CHORDATA, CLASS AND SUBCLASS - OSTEICHTHYES, ORDER AND SUBORDER - SILURIFORMES, FAMILY AND SUBFAMILY - ICTALURIDAE, GENUS AND SUBGENUS - ICTALURUS, SPECIES AND SSP - PRICEI, SCIENTIFIC NAME - ICTALURUS PRICEI AUTHORITY - TAXONOMY REFERENCES - Taxonomy - 1 (DRAFT) - Taxonomy Species CATFISH, YAQUI Species Id ESIS252013 Date 14 MAR 96 COMMENTS ON TAXONOMY - Yaqui Catfish Ictalurus pricei (Rutter, 1896) KINGDOM: Animal GROUP: Fish PHYLUM: Chordata CLASS: Osteichthyes ORDER: Siluriformes FAMILY: Ictaluridae Very little is known about the taxonomic status of this species. J. Humphries and R.R. Miller (01) are now synthesizing data on the Mexican ictalurids. The Yaqui catfish (Ictalurus pricei) is a moderately large (to 50 cm TL), fork-tailed species characterized in part by the presence of "villia-like" rugosities on the skin (01). Minckley (01) points-out that this condition is found in catfish from headwater regions of streams but not in channel catfish. Coloration dark gray to black dorsally, white to grayish beneath; far darker over-all than either the channel or blue catfishes. Barbels jet-black except on chin, where gray to whitened. Body usually profusely speckled (03). This species was first described by Rutter (1896) as Villarius pricei. A possible synonym according to Miller (04) may be Ictalurus meeki from the Rio Yaqui. Taxonomy - 2
                                  (DRAFT) - Status
                               Species CATFISH, YAQUI
                                Species Id ESIS252013
                                   Date 14 MAR 96



STATUS

Coded Status T: Federal Threatened Game (Consumptive Recreational) COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The Yaqui catfish (Ictalurus pricei) has been designated a Threatened species pursuant to the Endangered Species Act of 1973 (50 CFR 17.11; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The species has this status wherever found including the States of Arizona, USA, and Chihuahua and Sonora, Mexico.. Critical Habitat has been designated in aquatic habitats of San Bernardino NWR in the S 1/2 Sec. 11; Sec. 14; S 1/2 and NE 1/4 Sec. 15 T245, R30E in Cochise County, AZ (50 CFR 17.95(e)). Special rules concerning "take" for educational, scientific, or conservation purposes in accordance with applicable Arizona State laws and regulations for this species can be found in 50 CFR 17.44(h). This species is protected by the Lacey Act (P.L. 97-79, as amended; 16 U.S.C. 3371 et seq.) which makes it unlawful to import, export, transport, sell, receive, acquire, or purchase any wild animal (alive or dead including parts, products, eggs, or offspring): (1) in interstate or foreign commerce if taken, possessed, transported or sold in violation of any State law or regulation, or foreign law; or (2) if taken or possessed in violation of any U.S. law, treaty, or regulation or in violation of Indian tribal law. It is also unlawful to possess any wild animal (alive or dead including parts, products, eggs, and offspring) within the U.S. territorial or special maritime jurisdiction (as defined in 18 U.S.C. 7) that is taken, possessed, transported, or sold in violation of any State law or regulation, foreign law, or Indian tribal law. RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely to jeopardize the continued existence of the species or result in the destruction or adverse modification of Critical Habitat (50 CFR 402), and to utilize their authorities to carry out programs for the conservation of the species. Status - 1 (DRAFT) - Status Species CATFISH, YAQUI Species Id ESIS252013 Date 14 MAR 96 STATE STATUSES AND LAWS: STATE: Arizona DESIGNATED STATUS: "Game Fish", Group 1, species is extirpated from the state. ADMINISTRATIVE AGENCY: Arizona Department of Game and Fish STATE STATUTE: AZ Rev. Stat. 17-102 and 17-101 A 21; and AZ Admin. Rules and Regs. 12-4-210 (permits). INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: None. ECONOMIC STATUSES: This species has value as part of the cultural value of Arizona's natural heritage of biotic diversity, and it also served in a minor way, as a recreational outlet for the fisherman. 82/12/30:47 FR 58454/58460 - Notice of Review (vertebrates). 83/07/15:48 FR 32527/32534 - Proposed rule. 84/08/31:49 FR 34490/34497 - Listed as Endangered. Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC INLAND AQUATIC COMMENTS ON HABITAT ASSOCIATIONS - The Yaqui catfish ranges from an elevation of about 2100 m downward in relatively small creeks and rivers to the Rio Yaqui mainstream and backwater habitats on the delta of that river. This species exhibits broad ecological tolerance (01). Hendrickson et al. (02) indicates that no fish of this species were captured from artificial reservoirs although the species occurred in substantial numbers above and below the reservoirs. "The species was most common in larger river areas of medium to slow currents over sand/rock bottoms. In small streams at higher elevations, however, it lived in quiet, clear pools with abundant macrophytes, over gravel/sand substrate. The absence of the species from collections in the upper Papigochic system in recent years...may indicate a reduction in its range..." Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species CATFISH, YAQUI
                                Species Id ESIS252013
                                   Date 14 MAR 96



FOOD HABITS

TROPHIC LEVEL - OMNIVORE LIFESTAGE FOOD FOOD PART General Detritus General Carrion General Fish Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species CATFISH, YAQUI
                                Species Id ESIS252013
                                   Date 14 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Inland wetlands: swamp G Inland Wetland: vegetated stream banks G G Aquatic Features: Pool areas G Aquatic Features: Springs [flowing] G Terrestrial Features: Downed logs Environment Associations - 1
                               (DRAFT) - Life History
                               Species CATFISH, YAQUI
                                Species Id ESIS252013
                                   Date 14 MAR 96



LIFE HISTORY

FOOD HABITS: The stomach contents of four individuals revealed the remains of other fishes and detrital/carrion materials (01). HOME RANGE/TERRITORY: No information on home range/territory. PERIODICITY: No information on periodicity. MIGRATION PATTERNS: No information on migration patterns. COVER/SHELTER REQUIREMENTS: No information on cover/shelter requirements. REPRODUCTIVE SITE REQUIREMENTS: No information on reproductive site requirements. REPRODUCTIVE CHARACTERISTICS: There is little information on the reproductive biology of the Yaqui catfish. Males from the Rio Sirupa, Chihuahua developed swollen heads in breeding condition and females had ripe ova in mid-June 1978. PARENTAL CARE: No information on parental care. POPULATION BIOLOGY: No information on population biology. SPECIES INTERRELATIONSHIPS: Associated species include most large-river fishes of the Rio Yaqui system especially the Yaqui sucker, and roundtail chub in the channels of larger streams. Smaller fishes taken with the Yaqui catfish in large, mainstream habitats included the young of the Yaqui sucker and roundtail chub as well as the beautiful shiner, while Mexican stoneroller, longfin dace, and others lived along the stream margins (02). OTHER LIFE HISTORY DESCRIPTORS: No information on other life history descriptors. Life History - 1
                           (DRAFT) - Management Practices
                               Species CATFISH, YAQUI
                                Species Id ESIS252013
                                   Date 14 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Restricting/regulating human disturbance of populations Beneficial Developing/maintaining stream structures Beneficial Maintaining/Controlling Water Flow Beneficial Developing/maintaining/protecting wetlands Beneficial Controlling water levels Beneficial Controlling/Restricting Pesticide Use Beneficial Controlling/Restricting Herbicide Use Beneficial Controlling/Restricting Agricultural Practice Beneficial Creating Artificial Habitat/Nesting Structure Beneficial Reforestation Beneficial Stocking captive-reared wild-strain animals Beneficial Controlling/Removing Exotic Vertebrates Adverse Surface Mines Existing Surface Mines Adverse Siltation Existing Siltation Adverse Water Temperature Alteration Existing Water Temperature Alteration Adverse Water Level Fluctuation Existing Water Level Fluctuation Adverse Irrigating Existing Irrigating Adverse Reservoirs Existing Reservoirs Adverse Migration barriers Existing Migration barriers Adverse Dredging Existing Dredging Adverse Competition Existing Competition Adverse Erosion Existing Erosion Adverse Grazing Existing Grazing Adverse Existing COMMENTS ON MANAGEMENT PRACTICES - This species was introduced into the upper Santa Cruz River basin, Arizona in the late 1800's but was extirpated by the 1950's (02). As for reasons for this species extirpation from the Santa Cruz River, suitable habitat conditions (i.e. water flow, temperatures, dissolved oxygen, cover, etc.) have been altered in a way that was detrimental to this species survival within this drainage. As for the populations within Mexico, "major modifications in the Rio Yaqui basin in Mexico consisted of three mainstream reservoirs, principally for irrigation storage and secondarilly for hydroelectric power, that regulate discharges and have had local Management Practices - 1 (DRAFT) - Management Practices Species CATFISH, YAQUI Species Id ESIS252013 Date 14 MAR 96 impacts on the system. Development of pumping and other facilities for irrigation and munning water supplies are associated with these installations. Highlands have been heaviliy overgrazed and subject to extensive logging in times past. Sawmill operations polluted upland streams, as did outwash of silt from slopes denuded by timber removal and spoil from local minning" (these actions have caused: channal modifications detrimental to the Yaqui fishes [i.e. passage barriers, water diversion drawdown, water level fluctuations, water temperature alterations, and siltation]) (01). Much of these effected areas are under improvement at this time, and will aid in preservation of this habitat area. As for the expanding irrigated agriculture, this still remains a prim factor in survival of the Rio Yaqui fishes (01). UNAPPROVED PLAN: U.S. Fish and Wildlife Service. 1984. Yaqui Fishes Recovery Plan. Technical Review Draft. U.S. Fish and Wildlife Service, Albuquerque, NM. v + 43 pp. The recovery of the Yaqui catfish is dependent upon the successful implementation of the master plan for the San Bernadino National Wildlife Refuge (NWR). Delisting is recommended when four stable populations have been established there. Since the Yaqui catfish does not presently exist within the U.S., it is necessary to obtain stocks of these fish from Mexico, where they are still available. The first step is to contact appropriate Mexican Government officials to request permits to obtain Yaqui catfish. A minimum of 200 individuals (400 is more desirable) would then be captured and transported to Dexter National Fish Hatchery, New Mexico. After obtaining fish from Mexico, other recommended actions include: 1. maintaining the stock of Yaqui catfish at Dexter NFH, where it probably can be successfully cultured because of its similarity with channel catfish, 2. development of cultural techniques for Yaqui catfish, 3. maintain the genetic integrity of hatchery reared Yaqui catfish, 4. provide a refugium for Yaqui catfish at Dexter NFH, 5. reintroduction of Yaqui catfish into suitable habitat on San Bernardino NWR (actions that may be required to create or maintain suitable habitat are: control or removal of exotic vertebrates, habitat revegetation/land rehabilitation, creation of artificial habitat or nesting areas, control of agricultural practices that may affect the habitat, restriction of use of pesticides and herbicides within the habitat area, maintaining the water table and flow rate, wetland management, stream bank or channel modification, and to limit human impact by limiting their access, 6. monitoring of reintroductions, 7. enforcement of all Federal laws protecting the species, and 8. development of public support through an information and education program through: a. development of an interpretive program at San Bernardino NWR, Management Practices - 2 (DRAFT) - Management Practices Species CATFISH, YAQUI Species Id ESIS252013 Date 14 MAR 96 b. preparation of an information pamphlet, c. production of a video for schools and public TV stations, and d. news releases to the media. The plan also calls for studies on this and other native fish species, Yaqui chub, Yaqui topminnow, and the Beautiful shiner (the other listed fishes present or destined for reintroduction at San Bernardino NWR), including: 1. basic studies on the biology and ecology of the species, 2. species habitat utilization, 3. competition and predation among listed species and other historically present species that are slated for reintroduction at San Bernardino NWR (roundtail chub, Mexican stoneroller, longfin dace, and Yaqui sucker). Ongoing recovery efforts: Management of the San Bernardino NWR. Management Practices - 3
                                   (DRAFT) - References
                                  Species CATFISH, YAQUI
                                  Species Id ESIS252013
                                      Date 14 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 Anon. [n.d.] Draft Recovery Plan for the Yaqui chub, beautiful shiner, Yaqui catfish, Sonoran (Yaqui) topminnow. Unpubl. info. on file with the U.S.D.I., Fish and Wildlife Service, Division of Endangered Species, 1000 N. Glebe Road, Arlington, VA. 02 Hendrickson, D., W.L. Minckley, R.R. Miller, D.J. Siebert, and P.H. Minckley. 1981. Fishes of the Rio Yaqui basin, Mexico and United States. Journ. Arizona-Nevada Acad. of Sci. 15(1980):66-106. 03 Minckley, W.L. 1973. Fishes of Arizona. AZ. Game and Fish Dept. 293 pp. 04 Miller, R.R. 1978. In: Trans. Symp. Biol. Res. Chihuahua Desert Reg. Pp. 365-81. ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 Smith, M.L. and R.R. Miller. 1986. The evolution of the Rio Grande basin as inferred from its fish fauna. pp. 457-486. In: C.H. Hocutt and E.O. Wiley (eds.) Zoogeography of North American Freshwater Fishes. John Wiley and Sons, Inc., New York, N.Y. 02 Minckley, W.L. 1985. Native fishes and natural aquatic habitats in the U.S. Fish and Wildlife Service Region II west of the Continental Divide. Unpubl. rep. on file with the U.S. Fish Wildl. Serv., Albuquerque, New Mexico. 03 Hendrickson, D. 1984. Distribution records of native exotic fishes in Pacific drainages of northern Mexico. Journ. Arizona-Nevada Acad. Sci. 18:33-38. 04 Miller, R.R. and C.H. Lowe. 1964. Pt. 2. Annotated check-list of the fishes of Arizona. pp. 133-151. In: C.H. Lowe, (ed.) The Vertebrates of Arizona. Univ. Ariz. Press, Tucson. 05 Minckley, W.L., D. Hendrickson, and C.E. Bond. 1986. Geography of western North American freshwater fishes: description and relationships to intra-continental tectonism. Pp. 519-613. IN: C.H. Hocutt and E.O. Wiley, eds. Zoogeography of North American Freshwater Fishes. John Wiley and Sons, Inc., New York, N.Y. 06 Minckley, W.L. 1973. Fishes of Arizona. Arizona Game Fish Dept. 293 pp. References - 1