(DRAFT) - Taxonomy
                               Species DARTER, NIANGUA
                                Species Id ESIS253001
                                   Date 14 MAR 96



TAXONOMY

NAME - DARTER, NIANGUA OTHER COMMON NAMES - DARTER and NIANGUA ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - CHORDATA, CLASS AND SUBCLASS - OSTEICHTHYES, ORDER AND SUBORDER - PERCIFORMES, FAMILY AND SUBFAMILY - PERCIDAE, GENUS AND SUBGENUS - ETHEOSTOMA, SPECIES AND SSP - NIANGUAE, SCIENTIFIC NAME - ETHEOSTOMA NIANGUAE AUTHORITY - TAXONOMY REFERENCES - COMMENTS ON TAXONOMY - Niangua Darter Etheostoma nianguae Gilbert and Meek in Gilbert, 1888 KINGDOM: Animal GROUP: Fish PHYLUM: Chordata CLASS: Osteichthyes ORDER: Perciformes FAMILY: Percidae Etheostoma nianguae is a large, slender darter with a long head that tapers into a slender, pointed snout. The background coloration is yellowish-olive, with seven or eight (04) prominent saddle bars along the back, and orange spots scattered over the upper sides. A series of U-shaped greenish blotches alternate with narrow orange bars along the midside. Two small, jet-black spots are present at the base of the caudal fin. Breeding males are more brilliantly colored than other individuals, and have an orange-red belly and a series of iridescent blue-green bars along the sides. The head length of the Niangua darter is a little more than 1/4 the standard length. There are 72 to 81 scales in the lateral line, which is complete. The anal fin has 2 stiff spines and 11 or 12 rays. The maximum total length is about 111 mm (05). The only near-relative of the Niangua darter is the arrow darter, Etheostoma sagitta (Jordan and Swain), which has a localized Taxonomy - 1 (DRAFT) - Taxonomy Species DARTER, NIANGUA Species Id ESIS253001 Date 14 MAR 96 distribution in the upper Cumberland and Kentucky stream systems of eastern Kentucky (05,08). These two species form a species group of the subgenus Oligocephalus. They share many characters, but the Niangua darter differs in having smaller scales, a more complete lateral line, a more completely scaled cheek and two discrete jet-black circular spots at the base of the caudal fin. Etheostoma nianguae was described by Gilbert and Meek (01) from specimens they collected from the Niangua River near Marshfield, MO, in the summer of 1884. The only synonym for the Niangua darter was Poecilichthys nianguae used by Bailey (08). The lectotype of this species is deposited at the U.S. National Museum (USNM 36214), and a paralectotype is deposited in the collections of Stanford University (SU 5299). The type material of E. nianguae is discussed by Collette and Knapp (02). For photographs, illustrations and identification keys to the Niangua darter, see Pflieger (03), Kuehne and Barbour (04), and Page (05). Taxonomy - 2
                                  (DRAFT) - Status
                               Species DARTER, NIANGUA
                                Species Id ESIS253001
                                   Date 14 MAR 96



STATUS

Coded Status T: Federal Threatened COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The Niangua darter (Etheostoma nianguae) has been designated a Threatened species pursuant to the Endangered Species Act of 1973 (50 CFR 17.11; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The species has this status wherever found including the State of MO. Critical Habitat for this species has been designated in Miller, Dallas, Greene, Cedar, St. Clair, Camden and Hickory Counties in Missouri (50 CFR, Sec. 17.95 (e)). Special rules (50 CFR, Sec. 17.44 (k)) address the subject of "take" for this species with the intent of enhancement of propagation and survival of the darter consistent with the Endangered Species Act. This species is protected by the Lacey Act (P.L. 97-79, as amended; 16 U.S.C. 3371 et seq.) which makes it unlawful to import, export, transport, sell, receive, acquire, or purchase any wild animal (alive or dead including parts, products, eggs, or offspring): (1) in interstate or foreign commerce if taken, possessed, transported or sold in violation of any State law or regulation; or (2) if taken or possessed in violation of any U.S. law, treaty, or regulation or in violation of Indian tribal law. It is also unlawful to possess any wild animal (alive or dead including parts, products, eggs, and offspring) within the U.S. territorial or special maritime jurisdiction (as defined in 18 U.S.C. 7) that is taken, possessed, transported, or sold in violation of any State law or regulation, foreign law, or Indian tribal law. RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. DOD (COE) -Responsible for the law enforcement/protection of this species with applicable State and Federal laws on public land under their control. Also responsible for management/recovery on Department of Defense lands. All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely to jeopardize the continued existence of the species or result in the destruction or adverse modification of Critical Habitat (50 CFR 402), and to utilize their authorities to carry out programs for the Status - 1 (DRAFT) - Status Species DARTER, NIANGUA Species Id ESIS253001 Date 14 MAR 96 conservation of the species. STATE STATUSES AND LAWS: STATE: Missouri DESIGNATED STATUS: Endangered ADMINISTRATIVE AGENCY: Missouri Department of Conservation STATE STATUTE: Chapter 252, Revised Statutes of Missouri Wildlife Code of Missouri (3 Code of State Regulations-4.111). INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: The Niangua darter is listed in the IUCN Red Data Book, Vol. 4, 1977. ECONOMIC STATUSES: None. 82/12/30:47 FR 58454/58460 - Vertebrate notice of review 84/04/17:49 GR 15102/15109 - Proposed Threatened w/ CH and Spec. Rules 85/06/12:50 FR 24649/24654 - Listing as Threat w/ CH and Spec. Rules Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC INLAND AQUATIC LAND USE - Transportation, communications, and Util Cropland and Pasture Deciduous Forest Land Streams and Canals Reservoirs NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Riverine, upper perennial UB1 Riverine, upper perennial SB1 Riverine, upper perennial FL1 COMMENTS ON HABITAT ASSOCIATIONS - All populations of the Niangua darter are in streams of the Salem Plateau, of order 3, 4 and 5, having gradients of 3 to 21 feet/ mile (06). In these streams the species was not collected closer to the ultimate headwater divide than 8 miles, or farther from the headwater divide than 49 miles. Most collections were from localities where the local relief was between 150 and 250 feet, and the elevation of the streambed was between 550 and 1,050 feet. Streams in which the Niangua darter occurred may be characterized as small to moderate sized (05), moderately clear upland creeks, draining hilly topography underlain by bedrocks consisting principally of chert-bearing dolomites. Streams conforming to this description are widespread and common in the Ozark uplands. The Niangua darter was not found in small intermittent headwater creeks. Except when spawning, Niangua darters were most often found in shallow pools or "runs" having slight to moderate current and clean, gravelly or rocky bottoms. Occasionally, Niangua darters were found in riffles as early as mid-March, and by mid-April when spawning occurred most adults were in or near that habitat. By mid-May spawning was completed and Niangua darters were again found in pools. In early fall they were found only in pools and slow "runs." The winter habitat was not determined, but may be similar to that in which the species was found in the fall. Throughout the year the Niangua darter exhibited a preference for silt-free substrates. The substrate was generally gravel with scattered rubble and occasional boulders or bedrock. Stream sections where this species occurred commonly had thick growths of water willow (Justicia americana) but the Niangua darter did not exhibit any particular affinity for this vegetation. Land use/land cover types associated with the Niangua darter habitat include State highways and right-of-ways, pasture areas, second growth oak-hickory forests and reservoirs [i.e., Harry S. Truman Reservoir (COE)]. Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species DARTER, NIANGUA
                                Species Id ESIS253001
                                   Date 14 MAR 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE LIFESTAGE FOOD FOOD PART General Crustaceans General Arthropods Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species DARTER, NIANGUA
                                Species Id ESIS253001
                                   Date 14 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Aquatic Features: Pool areas Environment Associations - 1
                               (DRAFT) - Life History
                               Species DARTER, NIANGUA
                                Species Id ESIS253001
                                   Date 14 MAR 96



LIFE HISTORY

FOOD HABITS: In a food analysis study the food of 34 Niangua darters consisted almost entirely of the immature stages of aquatic insects, along with occasional cladocerans, copepods and amphipods (06). Only a single item, a hymenopteran insect, was of terrestrial origin. Nymphs of mayflies (Ephemeroptera) and stoneflies (Plecoptera) comprised most of the food items in Niangua darter stomachs. Microcrustaceans and amphipods occurred in three stomachs. Sand and fine gravel which occurred in 10 (29.4 percent) of the stomachs was probably ingested accidentally. The same may also be true of plant fragments that occurred in two (5.9 percent) of the stomachs. Little evidence was found of change in food habits with size over the length range (34.0 to 94.0 mm SL) of the specimens available for analysis. Microcrustaceans occurred only in two of the smaller specimens (42.5 and 44.5 mm SL). Stonefly nymphs occurred only in spring samples, but in that season they were more important volumetrically than any other taxonomic group. Isoperla, which was the most abundant stonefly genus, occurred in 7 stomachs and comprised 84.0 percent by volume of all stoneflies. Nymphs of the genus Nemoura occurred in two stomachs and nymphs of the genus Isogenus and the families Capniidae and Chloroperlidae each occurred in one stomach. Mayfly nymphs were numerous in both spring and fall stomachs. The family Heptageniidae, including Heptagenia, Rhithrogena, Stenonema and Stenacron was the dominant mayfly group. Stenonema comprised 85.3 percent (spring) and 80.9 percent (fall) by volume of the heptageniid mayflies in Niangua darter stomachs. Stenonema tripunctatum, S. pulchellum and S. nepotellum were identified. Heptagenia occurred only in spring stomachs and Stenacron occurred only in fall stomachs. The mayfly genus Baetis (Baetidae) comprised 12.1 percent by volume of all mayfly nymphs in the spring, but did not occur in fall stomachs. Leptophlebiid nymphs comprised 3.0 percent by volume of mayfly nymphs in spring stomachs and 7.4 percent in fall stomachs. Chironomidae were the only Diptera identified. They occurred in seven (20.6 percent) of the stomachs and comprised 1.8 percent by volume of all insects. A higher proportion of stomachs (39.3 percent) contained no food items in fall as compared with spring (16.7 percent), suggesting more feeding activity in spring. Two specimens collected Febraury 28, 1977, contained food, indicating some food intake even in winter. Identifiable items in their stomachs included the only occurrences of Cladocera and the mayfly Ephemerella and one of two occurrences of Copepoda. HOME RANGE/TERRITORY: Niangua darters exhibited a clumped or non-random distribution (06). The majority of the pools had none, but when one individual was found, there were usually others nearby. As many as nine individuals were observed and expected frequencies revealed a significant departure from randomness in the distribution of this species. Evidently the Niangua darter is gregarious or exhibits an affinity for Life History - 1 (DRAFT) - Life History Species DARTER, NIANGUA Species Id ESIS253001 Date 14 MAR 96 certain stream pools. On several occasions the same individual was observed in the same general areas on different days, suggesting that the Niangua darter may occupy a home range. This species does not appear to be territorial, except perhaps briefly during the spawning season, when males may occupy a small breeding territory. PERIODICITY: Periodicity has not been observed in the Niangua darter. It is thought to be diurnal and relatively inactive in winter. MIGRATION PATTERNS: Migration does not appear to be an important aspect of the life history of the Niangua darter. The species appears to occupy the same stream reaches throughout the year. COVER/SHELTER REQUIREMENTS: Water depths in pools and runs, where Niangua darters were observed in early spring (March to early April), ranged from 8 to 24 inches, averaging about 16 inches (06). Current velocities were sometimes to slight to measure, but more often were between 0.7 and 2.0 feet/second. Riffles where Niangua darters were found during the spawning season in mid-April ranged in depth from 5 to 18 inches and averaged 11.6 inches. Current velocities were generally 3.1 to 40 feet/second. By mid-May spawning was completed and Niangua darters were again found in pools and slow "runs". Niangua darters were found in September, October and November in pools and slow "runs" that had minimal current and depths ranging from 7 to 46 inches (average 22.4 inches). Throughout the year the Niangua darter exhibited a preference for silt-free substrates. The substrate was generally gravel with scattered rubble and occasional boulders or bedrock. Stream sections where this species occurred commonly had thick growths of water willow (Justicia americana) but the Niangua darter did not exhibit any particular affinity for this vegetation. One Niangua darter took refuge beneath a large stone when an attempt was made to capture it with a dip net and another was observed emerging from beneath a stone. On one occasion when a stream section was snorkeled twice on the same day, more individuals were observed near noon than in early morning. Perhaps the Niangua darter spends periods of inactivity beneath stones. REPRODUCTIVE SITE REQUIREMENTS: Each year adults move from pools and "runs" to swift gravelly riffles just prior to spawning (06). Swift current and gravel substrate appear to be critical factors in the selection of spawning sites by the Niangua darter. Riffles floored by bedrock or coarse rubble are not used. Spawning riffles are in most cases places where the stream is actively cutting gravel during floods. Due to sorting action of the swift current, the substrate is largely free of silt. During spawning the female Niangua darter burrows into the gravel where the eggs are deposited. In Big Tavern Creek the Niangua darter stopped using a riffle in response to changes in its configuration Life History - 2 (DRAFT) - Life History Species DARTER, NIANGUA Species Id ESIS253001 Date 14 MAR 96 from one year to the next. The change involved channel widening which resulted in less current. Current velocity, depth and substrate composition were quite similar in three spawning riffles. Current velocities on other spawning riffles never measured less than 3.1 feet/second. Dry weight does not provide an adequate portrayal of substrate composition, since a few large stones can dominate the analysis. The flow rates are for some of the smallest streams where the Niangua darter was found and perhaps indicate the minimum for Niangua darter spawning streams. A base flow of 243.1 cf/s was measured in April at Station 32 on the Niangua River, the largest stream where this species was found. Each year, daily maximum temperatures (degrees F) were in the upper 50's when males began congregating on spawning riffles, and were near 65 degrees F when spawning began. In 1971 and 1975 the initiation of spawning was accompanied by an abrupt rise in stream temperatures as a warm front moved through. The following is a table of physical characteristics of three Niangua darter spawning riffles (06). Substrate composition is the total dry weight of particles in the indicated size ranges. PHYSICAL CHARACTERISTIC - RIFFLE # 1 - RIFFLE # 2 - RIFFLE # 3 Current velocity (surface) 4 ft/sec 4 ft/sec 4 ft/sec Volume of flow 13.8 cf/s 13.8 cf/s 9.9 cf/s Riffle depth (max) 10.5 in 11.5 in 5.7 in Riffle width (max) 8 ft 6 ft 3 ft Riffle length 40 ft 33 ft 13 ft Substrate particle size: 1/8 inch or less 7.2 pct. 6.3 pct. 2.5 pct. 1/8 inch to 1/4 inch 8.8 pct. 8.7 pct. 4.6 pct. 1/4 inch to 1/2 inch 9.8 pct. 15.8 pct. 10.2 pct. 1/2 inch to 3/4 inch 13.9 pct. 13.8 pct. 13.5 pct. 3/4 inch or more 60.3 pct. 55.4 pct. 69.2 pct. REPRODUCTIVE CHARACTERISTICS: The length of the spawning period of the Niangua darter has not been adequately documented (06). In 1971, males were still on the spawning riffles April 19, but had deserted the riffles by April 30. Spawning was evidently completed over a period of 2 or 3 weeks in April. In 1975 heavy rains, April 24, may have interrupted spawning. In 1976, males were still on the riffles April 15. No further observations were made that year until May 26, by which time spawning was completed. Males precede females to the spawning riffles and there is an obvious predominance of females in adjacent pools at that time. Females seem to enter the riffles only to spawn and males generally outnumber females in these areas throughout the spawning period. Male Niangua darters follow and attempt to mount females while still in pools. Females respond by engaging in head-bobbing. Encounters between males result in threat displays characterized by erection of the spinous dorsal. Encounters between a male and other males or mature females are accompanied by a change in color pattern Life History - 3 (DRAFT) - Life History Species DARTER, NIANGUA Species Id ESIS253001 Date 14 MAR 96 that heightens the contrast between his light background color and darker markings. The anterior saddle bar extending downward behind the pectoral fin is especially noticeable during these encounters. This color change does not occur in females. Observations of spawning behavior are difficult because the spawning riffles are shallow and the darters are easily frightened. The spawning act was observed in its entirety only once. This observation and other less complete observations indicate that spawning occurs as the female lies buried in the gravel with the male above. The eggs are mixed in the gravel and left to hatch without further attention from the adults. A Males and females may become sexually mature at one year of age and most reproduction is accomplished by adults one or two years of age. The number of mature ova produced by females averaged 189.8 for age-group-I and 3379.8 for age-group-II. An age group IV females produced 748 mature ova. PARENTAL CARE: The Niangua darter does not practice parental care. The eggs are abandoned soon after spawning. POPULATION BIOLOGY: Populations of the Niangua darter consist mostly of individuals one or two years of age with maximum longevity of about four years. The sexes are approximately equally represented (06). Nine species of darters occur in association with the Niangua darter and most are more abundant than E. nianguae. Competition for food with these species is probably minimized by different feeding strategies. The Niangua darter obtains its food by probing crevices with its slender snout. Potential predators of the Niangua darter are all Centrarchids, of which seven species are present. Two of these (spotted bass and rock bass) are not native within the range of the Niangua darter. No heavy infestations of parasites were noted for the Niangua darter. The population biology of the Niangua darter is not notably different from that of other darters providing few insights into the reasons for its scarcity and limited distribution. SPECIES INTERRELATIONSHIPS: The Niangua darter is not known to have a direct causal relationship with any other species. OTHER LIFE HISTORY DESCRIPTORS: No other life history descriptors can be added at this time for the Niangua darter. Life History - 4
                           (DRAFT) - Management Practices
                               Species DARTER, NIANGUA
                                Species Id ESIS253001
                                   Date 14 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Restricting/regulating human disturbance of populations Beneficial Developing fishways Beneficial Developing/maintaining stream structures Beneficial Controlling/Restricting Road Maintainance Actions Beneficial Land Acquisition Beneficial Controlling pollution [thermal, chemical, physical] Beneficial Controlling/Restricting Pesticide Use Beneficial Controlling/Restricting Herbicide Use Beneficial Controlling/Restricting Agricultural Practice Beneficial Stocking captive-reared wild-strain animals Beneficial Transplanting wild animals Beneficial Controlling/Removing Native Vertebrates Adverse Predation Existing Predation Adverse Siltation Existing Siltation Adverse Flooding Existing Flooding Adverse Reservoirs Existing Reservoirs Adverse Migration barriers Existing Migration barriers Adverse Dredging Existing Dredging Adverse Developing/maintaining stream bank vegetation Existing Developing/maintaining stream bank vegetation Adverse Environmental Contamination/Pollution Existing Environmental Contamination/Pollution Adverse Competition Existing Competition Adverse Exotic/Feral/Introducted Species Existing Exotic/Feral/Introducted Species Adverse Erosion Existing Erosion Adverse Existing Adverse Harvesting Existing Harvesting COMMENTS ON MANAGEMENT PRACTICES - The Niangua darter is rare, localized in occurrence and vulnerable to extinction (06). The extent to which man is responsible for the present status of this species is uncertain because the distribution of this species was not documented until recently. However, there is considerable circumstantial evidence that its range has been reduced by man's activities and further reduction can be expected if present trends continue. Reservoir construction may be the principal threat to survival of Management Practices - 1 (DRAFT) - Management Practices Species DARTER, NIANGUA Species Id ESIS253001 Date 14 MAR 96 the Niangua darter. Four major reservoirs (Lake of the Ozarks, Pomme de Terre Reservoir, Stockton Reservoir and Truman Reservoir) have been constructed within its range. These reservoirs have destroyed Niangua darter habitat. Taber and Wilkinson (07) reported the collection of Niangua darters from Maze (Arbell) Creek, 50 yards upstream from the rising waters of Stockton Reservoir, at a locality subsequently covered by 15 feet of water. These specimens likely were upstream migrants retreating ahead of the lake as it filled. Only 6 miles of Maze Creek are not inundated by Stockton Reservoir, and this stream does not support a population of the species. All known habitat for the Niangua darter in the Little Pomme de Terre River, Benton County, MO, is within the flood pool of Truman Reservoir. Habitat changes associated with periodic inundation may have eliminated this population. The lower 2 miles of Brush Creek, St. Clair County, MO, are also within the flood pool of this reservoir. Reservoir construction could be detrimental to the Niangua darter in ways unrelated to the physical destruction or degradation of habitat. The influx of species favored by the reservoir into tributary streams may increase competition and/or predation. The logperch, a potential competitor with the Niangua darter, is often favored by reservoir construction. The largemouth bass and the spotted bass are favored by reservoirs and are potential predators. Reservoir construction creates insurmountable barriers to the dispersal of the Niangua darter between suitable habitats. Such movements are essential for maintaining populations in streams where local extinctions occur as a result of environmental extremes or other factors. Studies of insular biogeography have indicated that repeated local extinction, followed by dispersal and recolonization from other inhabitated areas, is characteristic of all species that occur in discontinuous habitats. The introduction of fishes not native to the Osage Basin is another change that could be detrimental to the Niangua darter. The spotted bass and the rock bass were introduced by 1940, and are now widely distributed in streams where Niangua darters occur. Both are to some extent piscivorous and thus are potential predators of the Niangua darter. The general deterioration of stream habitats in the Osage Basin, resulting from a variety of factors is, less obvious but no less important than reservoir construction in determining the prospects for survival of the Niangua darter. The accelerated conversion of woodlands to pasture in recent years is one factor. Increased sedimentation and nutrient enrichment are likely results of this activity. Stream channelization is not as extensive within the range of the species as in some areas of Missouri, but is still a factor in habitat destruction. It has been common practice to channelize streams for a short distance above and below new road bridges. Landowners also channelize streams to control local flooding. Another common practice that is detrimental to stream habitat is the removal of willows and other woody vegetation from the stream channel, on the assumption that this increases water carrying capacity and reduces bank erosion. This results in greater instability of the substrate. Management Practices - 2 (DRAFT) - Management Practices Species DARTER, NIANGUA Species Id ESIS253001 Date 14 MAR 96 There is little doubt that all of the factors discussed above contribute to a general reduction of the quality of stream habitat. A comment heard from long-time residents throughout the Ozarks and specifically from along some streams inhabited by the Niangua darter is that the streams are "graveling in." By this these observers mean that the channel is becoming choked with deposits of unstable gravel. This eliminates pools and reduces surface flow across riffles. The Niangua darter has persisted in spite of these changes, but it cannot be assumed that it will continue to do so indefinitely. The general deterioration of habitat and other factors may be exerting stresses on existing Niangua darter populations so that recruitment fails to completely compensate for mortality. This increases the likelihood of local extirpation of populations, and once eliminated, these populations will not be reestabilished because the reservoirs which isolate them are barriers to dispersal. This could lead the species slowly and almost imperceptably to extinction. UNAPPROVED PLAN: As of 1986, no target date for recovery plan completion has been estabilished. A Recovery Plan should consider the following: 1) long term protection of riparian habitat including; land protection (acquisition, management agreements, etc.); reducing adverse agricultural practices on adjacent lands, stream bank modification; controlling the use of pesticides/herbicides and other pollutants (i.e., on roads near stream habitats, on adjacent agricultural lands, etc.); and controlling stream channel modification and channelization for water level stabilization (i.e., near agricultural lands and roads/highways). 2) Establish procedures to permit and increase the movement of the species between areas of suitable habitat. These procedures may involve stream channel modification by eliminating passage barriers or providing fish passage devices. 3) The possibility of captive breeding and reintroduction; and 4) methods of controlling the influx of predator fish from existing impoundments. Management Practices - 3
                                   (DRAFT) - References
                                 Species DARTER, NIANGUA
                                  Species Id ESIS253001
                                      Date 14 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 Gilbert, C.H. 1888. Descriptions of new and little known etheostomids. Proc. U.S. Nat. Mus. 10:47-64. 02 Collette, B.B. and L.W. Knapp. 1966. Catalog of type specimens of the darters (Pisces, Percidae, Etheostomatini). Proc. U.S. Nat. Mus. 119(3550):1-88. 03 Pflieger, W.L. 1975. The fishes of Missouri. Missouri Dept. Conserv. 342 pp. 04 Kuehne, R.A. and R.W. Barbour. 1983. The American darters. Univ. of Kentucky Press, Lexington. 177 pp. 05 Page, L.M. 1983. Handbook of darters. TFH Pub., Inc., Neptune City, NJ. 271 pp. 06 Pflieger, W.L. 1978. Distribution, status, and life history of the Niangua darter, Etheostoma nianguae. Missouri Dept. Conserv., Aquatic Series No. 16. 25 pp. 07 Taber, C.A. and R.F. Wilkinson, Jr. 1973. Etheostoma nianguae in the Sac River drainage, Missouri. Amer. Midl. Nat. 89(1):251-252. 08 Bailey, R.M. 1948. Status, relationships, and characters of the Percid fish Poecilichtys sagitta (Jordon and Smith). Copeia 1948, No. 2, Pp. 77-85. ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 Pflieger, W.L. 1971. A distributional study of Missouri fishes. Mus. Nat. Hist., Univ. of Kansas Publ. 20(3):2225-570. 02 Pflieger, W.L. 1978. Distribution, status, and life history of the Niangua darter, Etheostoma nianguae. Missouri Dept. Conserv., Aquatic Series No. 16. 25 pp. References - 1