(DRAFT) - Taxonomy
                               Species DARTER, BOULDER
                                Species Id ESIS254017
                                   Date 13 MAR 96



TAXONOMY

NAME - DARTER, BOULDER OTHER COMMON NAMES - DARTER, BOULDER;DARTER and ELK RIVER ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - CHORDATA, CLASS AND SUBCLASS - OSTEICHTHYES, ORDER AND SUBORDER - PERCIFORMES, FAMILY AND SUBFAMILY - PERCIDAE, GENUS AND SUBGENUS - ETHEOSTOMA, SPECIES AND SSP - SP., SCIENTIFIC NAME - ETHEOSTOMA SP. AUTHORITY - TAXONOMY REFERENCES - COMMENTS ON TAXONOMY - Boulder Darter Etheostoma sp. KINGDOM: Animal GROUP: Fish PHYLUM: Chordata CLASS: Osteichthyes ORDER: Perciformes FAMILY: Percidae The boulder darter (Etheostoma sp.), sometimes referred to as the Elk River darter, is an undescribed species in the subgenus Nothonutus (a manuscript describing it is in preparation) (01). It attains a maximum length of about 3 inches (01). The body of males is olive to gray, and they lack the red spots that are characteristic of closely related species. The female's color is similar but lighter. Both sexes have a gray to black bar located below the eye and a similar colored spot behind the eye. Type specimens are located at Illinois Natural History Survey, Tulane University, University of Alabama, Florida State Museum, U.S. National Museum, University of Michigan, and the University of Tennessee. Photographs will be available in the species' description being prepared (01). Taxonomy - 1
                                  (DRAFT) - Status
                               Species DARTER, BOULDER
                                Species Id ESIS254017
                                   Date 13 MAR 96



STATUS

Coded Status Alabama; Federal Endangered Tennessee; Federal Endangered Tennessee; State Recognized E: Federal Endangered COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The boulder darter (Etheostoma sp.) has been designated an Endangered species pursuant to the Endangered Species Act of 1973 (50 CFR 17.11; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The species has this status wherever found including the States of Alabama and Tennessee. This species is protected by the Lacey Act (P.L. 97-79, as amended; 16 U.S.C. 3371 et seq.) which makes it unlawful to import, export, transport, sell, receive, acquire, or purchase any wild animal (alive or dead including parts, products, eggs, or offspring): (1) in interstate or foreign commerce if taken, possessed, transported or sold in violation of any State law or regulation; or (2) if taken or possessed in violation of any U.S. law, treaty, or regulation or in violation of Indian tribal law. It is also unlawful to possess any wild animal (alive or dead including parts, products, eggs, and offspring) within the U.S. territorial or special maritime jurisdiction (as defined in 18 U.S.C. 7) that is taken, possessed, transported, or sold in violation of any State law or regulation, foreign law, or Indian tribal law. RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely to jeopardize the continued existence of the species or result in the destruction or adverse modification of Critical Habitat (50 CFR 402), and to utilize their authorities to carry out programs for the conservation of the species. STATE STATUSES AND LAWS: STATE: Alabama DESIGNATED STATUS: None STATE: Tennessee DESIGNATED STATUS: Recognized endandered Status - 1 (DRAFT) - Status Species DARTER, BOULDER Species Id ESIS254017 Date 13 MAR 96 ADMINISTRATING AGENCY: Tennessee Wildlife Resources Agency STATE STATUTES: Proclamations 75-15, 76-4 and amendments. INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: None ECONOMIC STATUSES: This species has no cultural or commercial value. The species may have aesthetic or scientific value. 85/09/18:50 FR 37959/ - Notice of Review 87/11/17:52 FR 43921/ - Proposed Rule 88/09/01: FR - Final Rule; listed as Endangered Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC INLAND AQUATIC LAND USE - Streams and Canals Cropland and Pasture Deciduous Forest Land NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Riverine, upper perennial RB2 Riverine, upper perennial RB1 COMMENTS ON HABITAT ASSOCIATIONS - The boulder darter inhabits a warn water riverine environment and has been found only in moderate- to fast-moving water over boulder/slab rock habitat in water over 2 feet deep. The fish has not been taken in pool habitat or in gravel runs lacking in boulder habitat. The species' preferred habitat is limited (01,02). Most of the habitat associated with the lower watershed where the fish exists is rural with only scattered housing. However, a tributary to the Elk River, Richland Creek, flows through a small city. Pasture and grazing are the prime use of cleared farm lands, but some row crops, primarily corn, are grown. Forest parallel most river sections and some second and third growth forests exist. Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species DARTER, BOULDER
                                Species Id ESIS254017
                                   Date 13 MAR 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE LIFESTAGE FOOD FOOD PART General Arthropods Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species DARTER, BOULDER
                                Species Id ESIS254017
                                   Date 13 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Inland wetlands: swamp G Aquatic Features: Pool areas G Coastal Features: Rocky offshore islands G Coastal Features: Sandy offshore islands G Coastal Features: Vegetated offshore islands G Terrestrial Features: Cliffs/ledges G Terrestrial Features: Rock outcrops G G Environment Associations - 1
                               (DRAFT) - Life History
                               Species DARTER, BOULDER
                                Species Id ESIS254017
                                   Date 13 MAR 96



LIFE HISTORY

Life History - 1 (DRAFT) - Life History Species DARTER, BOULDER Species Id ESIS254017 Date 13 MAR 96 FOOD HABITS: The food habits of this species are unknown. However, other members of the subgenus Nothonotus feed primarily on immature insects (01). HOME RANGE/TERRITORY: Based on behavior of related species, the adults probably are territorial and stay close to a home range associated with their boulder habitat (03,04). PERIODICITY: The species is likely a sight feeder and therefore probably diurnally active (04). MIGRATION PATTERNS: There are some indications that the juveniles may move into tributaries and also move downstream (02,04). COVER/SHELTER REQUIREMENTS: Adults appear to require large boulder/slabrock habitat in runs and riffles with moderate to fast water flow in water generally over 2 feet deep (01,02). Juveniles can likely exist in areas with smaller substrate size (02,04). REPRODUCTIVE SITE REQUIREMENTS: Spawning likely occurs in large boulder/slabrock habitat in areas with moderate to fast currents (01,02,04). The eggs of other members of the subgenus attach their eggs to the underside of rocks (03). REPRODUCTIVE CHARACTERISTICS: Based on behavior of related species, the boulder darter likely spawns from late May to early August (03,04). PARENTAL CARE: Based on data from related species, the males of this species likely guard the eggs until hatching (03,04). POPULATION BIOLOGY: The species is limited by the lack of boulder habitat in the Elk River (01,02). Recovery potential is low because much of its former habitat is impounded, but reclassification to threatened may be possible if a population can be established in Shoal Creek, Lawrence County, Tennessee. SPECIES INTERRELATIONSHIPS: The boulder darter is likely preyed upon by the banded sculpin and the smallmouth bass (04). OTHER LIFE HISTORY DESCRIPTORS: No additional data available. Life History - 2
                           (DRAFT) - Management Practices
                               Species DARTER, BOULDER
                                Species Id ESIS254017
                                   Date 13 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Maintaining/Controlling Water Flow Beneficial Regulating Water Temperature Beneficial Controlling pollution [thermal, chemical, physical] Beneficial Maintaining undisturbed/undeveloped areas Beneficial Controlling/Restricting Pesticide Use Beneficial Controlling/Restricting Herbicide Use Beneficial Creating Artificial Habitat/Nesting Structure Beneficial Stocking captive-reared wild-strain animals Beneficial Transplanting wild animals Beneficial Transplanting Wild Eggs/Wild Seeds Adverse Water Temperature Alteration Existing Water Temperature Alteration Adverse Reservoirs Existing Reservoirs Adverse Applying herbicides Existing Applying herbicides Adverse Applying pesticides Existing Applying pesticides Adverse Environmental Contamination/Pollution Existing Environmental Contamination/Pollution Adverse Existing COMMENTS ON MANAGEMENT PRACTICES - Historically, the boulder darter likely inhabited the southern bend area of the Tennessee River and the lower portion of its tributary from the Paint Rock River to at least Shoal Creek (02). The species was extirpated from most of this area by the impoundment behind Wheeler and Wilson Dams. The species' range in the Elk River was also reduced by Tims Ford Reservoir, which releases cold water into the Elk River. Because of the species' present restricted range (about 23 river miles) in the Elk River system, any factor that could impact water and substrate quality could adversely affect the species (01). UNAPPROVED PLAN: Utilize existing regulations to protect the species and its habitat. Develop propagation and transplantation techniques. Describe life history and evaluate threats. Except for enforcement of existing regulations, no recovery work is presently underway. Management Practices - 1
                                   (DRAFT) - References
                                 Species DARTER, BOULDER
                                  Species Id ESIS254017
                                      Date 13 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 O'Bara, C.J. and D.A. Etnier. 1987. Status survey of the boulder darter. Final report submitted to U.S. Fish and Wildlife Service, Asheville Field Office, 100 Otis Street, Room 224, Asheville, NC. May 1987. 13 pp. 02 Etnier, D.A. 1987. Personal communication, University of Tennessee, Knoxville, TN. 03 Stiles, R.A. 1972. The comparative ecology of three species of Nothonotus (Percidae-Etheostoma) in Tennessee's Little River. Ph.D. Diss., Univ. Tennessee. Knoxville, Tn. 97 pp. 04 Saylor, C.F. 1988. Personal communication. Tennessee Valley Authority, Norris, TN. ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 O'Bara, C.J., and D.A. Etnier. 1987. Status survey of the boulder darter. Final report submitted to U.S. Fish and Wildlife Service, Asheville Field Office, 100 Otis Street, Room 224, Asheville, NC. May 1987. 13 pp. 02 Etnier, David. 1987. University of Tennessee, personal communication with Richard G. Biggins, U.S. Fish and Wildlife Service, Asheville, NC. References - 1