(DRAFT) - Taxonomy
                        Species MUSSEL, PEARLY, HIGGINS' EYE
                                Species Id ESIS403001
                                   Date 13 MAR 96



TAXONOMY

NAME - MUSSEL, PEARLY, HIGGINS' EYE OTHER COMMON NAMES - MUSSEL, PEARLY, HIGGINS' EYE; MUSSEL and HIGGINS' EYE ELEMENT CODE - CATEGORY - Aquatic Molluscs PHYLUM AND SUBPHYLUM - MOLLUSCA, CLASS AND SUBCLASS - BIVALVIA, ORDER AND SUBORDER - UNIONOIDA, FAMILY AND SUBFAMILY - UNIONIDAE, GENUS AND SUBGENUS - LAMPSILIS, SPECIES AND SSP - HIGGINSI, SCIENTIFIC NAME - LAMPSILIS HIGGINSI AUTHORITY - TAXONOMY REFERENCES - COMMENTS ON TAXONOMY - Higgins' Eye Pearly Mussel Lampsilis higginsi (Lea, 1857) KINGDOM: Animal GROUP: Clam PHYLUM: Mollusca CLASS: Bivalvia ORDER: Unionoida FAMILY: Unionidae The shell is oval, elliptical, or rhomboid, thick and gaping at the anterior base. The species is sexually dimorphic. The anterior end is rounded and posterior end is bluntly pointed about midway up from the base in the male. The female shell is truncated and has a well-developed post-basal swelling. In both sexes the surface is usually shiny and marked with irregular growth lines. The epidermis is yellowish or olive with faint but distinct green rays. Internally the nacre is silvery-white and often tinged with pink or salmon, and is irridescent posteriorly (01,02). Photographs are available in 04. The species is most commonly called Higgins' eye mussel and is sometimes confused with Lampsilis orbiculata (04). However, L. higginsi and L. orbiculata are allopatric, so they are not likely to be confused in nature. Taxonomy - 1
                                  (DRAFT) - Status
                        Species MUSSEL, PEARLY, HIGGINS' EYE
                                Species Id ESIS403001
                                   Date 13 MAR 96



STATUS

Coded Status E: Federal Endangered Commercial COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The Higgins' eye pearly mussel (Lampsilis higginsi) has been designated an Endangered species pursuant to the Endangered Species Act of 1973 (50 CFR 17.11; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The species has this status wherever found including the States of IL, IA, MN, MO, NE, and WI. The present known distribution is encompassed by four States, IL, IA, MN, and WI. This species is protected by the Lacey Act (P.L. 97-79, as amended; 16 U.S.C. 3371 et seq.) which makes it unlawful to import, export, transport, sell, receive, acquire, or purchase any wild animal (alive or dead including parts, products, eggs, or offspring): (1) in interstate or foreign commerce if taken, possessed, transported or sold in violation of any State law or regulation; or (2) if taken or possessed in violation of any U.S. law, treaty, or regulation or in violation of Indian tribal law. It is also unlawful to possess any wild animal (alive or dead including parts, products, eggs, and offspring) within the U.S. territorial or special maritime jurisdiction (as defined in 18 U.S.C. 7) that is taken, possessed, transported, or sold in violation of any State law or regulation, foreign law, or Indian tribal law. RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely to jeopardize the continued existence of the species or result in the destruction or adverse modification of Critical Habitat (50 CFR 402), and to utilize their authorities to carry out programs for the Status - 1 (DRAFT) - Status Species MUSSEL, PEARLY, HIGGINS' EYE Species Id ESIS403001 Date 13 MAR 96 conservation of the species. STATE STATUSES AND LAWS: STATE: Illinois, Iowa, Minnesota, and Wisconsin DESIGNATED STATUS: Endangered ADMINISTRATIVE AGENCY: (IL) Department of Conservation; (IA) Conservation Commission; (MN) Department of Natural Resources; (WI) Department of Natural Resources. STATE STATUTES: (IL) Illinois Endangered Species Act, Ch. 8, Sec. 331-340, Illinois Revised Statutes; (IA) Management and Protection of Endangered Plants and Wildlife, Ch. 109A; (MN) Minnesota Law 1983, Ch. 97.488, Protection of Threatened and Endangered Species. (WI) Enabling Legislatin of Wisconsin Administrative Codes, Section 29.415, Endangered and Threatened Species Protected. STATE: Missouri DESIGNATED STATUS: Recognized Endangered ADMINISTRATIVE AGENCY: Department of Conservation STATE STATUTE: Wildlife Code of Missouri: Rule 3 CSR 10-4.111. INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: This species is listed in Appendix I of the Convention on International Trade in Endangered Species of Wild Fauna and Flora. ECONOMIC STATUSES: The Higgins' eye pearly mussel was at one time valued as a button shell and commercially harvested for this purpose. 74/10/17:39 FR 37078/37079 - Notice of review 75/09/26:40 FR 44329/44333 - Proposed rule 76/06/14:41 FR 24062/24067 - Final rule - listed as Endangered 77/02/22:42 FR 10462/10488 - Implementation of CITES 78/03/06:43 FR 09168/09172 - Advance notice of proposed rule on CITES 78/05/03:43 FR 21338/21339 - Advance notice of potential rule on CITES 81/02/27:46 FR 14652/14658 - Five year review 87/07/07:52 FR 25523/25528 - Notice of review Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC INLAND AQUATIC LAND USE - Streams and Canals NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Riverine, upper perennial UB2 Riverine, upper perennial UB1 Riverine, upper perennial OW0 COMMENTS ON HABITAT ASSOCIATIONS - L. higginsi prefers deep water habitat in large rivers (06). The species has been found in 8-15 feet of water in mud with a mixture of gravel and stones (03). They are usually found in beds bordering main or side channels and prefer areas of high turbulence and oxygen content (03). Little information is available about the specific habitats required by this species. The habitat of the glochidia is initially in the gills of the female then in the water column, and finally attached to a suitable host fish. Habitat associations or requirements for the juvenile stage are unknown Habitat Associations - 1
                                (DRAFT) - Food Habits
                        Species MUSSEL, PEARLY, HIGGINS' EYE
                                Species Id ESIS403001
                                   Date 13 MAR 96



FOOD HABITS

TROPHIC LEVEL - OMNIVORE LIFESTAGE FOOD FOOD PART General Detritus General Bacteria General Phytoplankton General Diatoms General Zooplankton General Fish Food Habits - 1
                         (DRAFT) - Environment Associations
                        Species MUSSEL, PEARLY, HIGGINS' EYE
                                Species Id ESIS403001
                                   Date 13 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Aquatic Features: Pool areas Environment Associations - 1
                               (DRAFT) - Life History
                        Species MUSSEL, PEARLY, HIGGINS' EYE
                                Species Id ESIS403001
                                   Date 13 MAR 96



LIFE HISTORY

FOOD HABITS: There is a lack of information on the feeding behavior of L. higginsi and for most of the bivalves of the Upper Mississippi River (04). However, there is no known interspecific differences in feeding among freshwater mussels (18). The glochidia are obligate parasites on the gills or fins of fish. Adult mussels are filter-feeders and consume particulate matter in the water column. Identifiable stomach contents almost invariably include desmids, diatoms, algae, protozoa, and zooplankters (12,19). HOME RANGE/TERRITORY: No information is available. PERIODICITY: No information is available. MIGRATION PATTERNS: Non-migratory. COVER/SHELTER REQUIREMENTS: The species seems to prefer deep water pools (8-15 ft. deep) with fast moving water (03). REPRODUCTIVE SITE REQUIREMENTS: The reproductive site requirements for this species appear to be the same as its general habitat requirements. They prefer deep water rivers with fast moving water (03,04). REPRODUCTIVE CHARACTERISTICS: The reproductive cycle of L. higginsi is similar to that of most unionids (04). No data are available for this particular species, but sexual maturity is not attained for several years in most related species. Lampsilis luteola were observed to be sexually mature by the third year (12). L. higginsi is sexually dimorphic and the posteriorly tapered males produce sperm that are discharged into the surrounding water (04). The posteriorly truncated females obtain these sperm during the normal process of siphoning water (04). It is unknown whether sexually mature individuals are able to detect the presence of one another during periods of reproductive activity (04). The breeding season is between May and September (06) and the species apparently has one breeding season per year. Fertilization of eggs occurs in the female's gills and fertilized eggs are retained within the gills where development takes place (04). Female L. higginsi are bradytitic (i.e., long-term breeders that retain the developing larvae within their marsupia throughout most of the year, except during early summer). A single female may brood and release several hundred thousand glochidia each year, but few reach sexual maturity (04). PARENTAL CARE: Eggs are fertilized within the gills of the female and are Life History - 1 (DRAFT) - Life History Species MUSSEL, PEARLY, HIGGINS' EYE Species Id ESIS403001 Date 13 MAR 96 retained until he eggs develop into the parasitic stage (glochidia) at which time they are released into the water. No other parental care occurs. POPULATION BIOLOGY: L. higginsi has historically occurred in low to medium densities and has never been abundant (04,15,16). Little is known about the population biology of the species. A combination of factors is probably responsible for its decline (04) (See N-REASONS). SPECIES INTERRELATIONSHIPS: Since the known host species of L. higginsi, sauger, freshwater drum, green sunfish, largemouth bass, small mouth bass, walleye, yellow perch, and white bass, are abundant and widespread. It is doubtful that the hosts are limiting or affecting the population of the species (17). Also, the possible competition of L. higginsi with the introduced Asiatic clam, Corbicula fluminea, is only speculative (10,11). OTHER LIFE HISTORY DESCRIPTORS: None. Life History - 2
                           (DRAFT) - Management Practices
                        Species MUSSEL, PEARLY, HIGGINS' EYE
                                Species Id ESIS403001
                                   Date 13 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Controlling/Restricting Boating Activities Beneficial Developing fishways Beneficial Developing/maintaining stream structures Beneficial Maintaining/Controlling Water Flow Beneficial Maintaining undisturbed/undeveloped areas Beneficial Controlling pollution [thermal, chemical, physical] Beneficial Transplanting wild animals Beneficial Regulating commercial harvest levels Adverse Commercial Exploitation Existing Commercial Exploitation Adverse Boating Existing Boating Adverse Inherent Reproductive Characteristics Existing Inherent Reproductive Characteristics Adverse Disease Existing Disease Adverse Recreational development Existing Recreational development Adverse Siltation Existing Siltation Adverse Water Level Stabilization Existing Water Level Stabilization Adverse Migration barriers Existing Migration barriers Adverse Dredging Existing Dredging Adverse Environmental Contamination/Pollution Existing Environmental Contamination/Pollution Adverse Competition Existing Competition Adverse Exotic/Feral/Introducted Species Existing Exotic/Feral/Introducted Species COMMENTS ON MANAGEMENT PRACTICES - Although L. higginsi was once widespread in the Upper Mississippi River and in some of its major tributaries, it was probably never abundant (04) because of its specific habitat requirements (05). Factors that have led to the species' decline are diverse and while some have substantial backing, others are subject to speculation. One factor leading the species' decline is commercial harvesting for a variety of decorative functions (03,04,05). Although it was not abundant, L. higginsi was considered a good button shell (06,07) and was harvested commercially when found (03). The general alteration of the Upper Mississippi River from riverine to an impounded system and changes in water quality as a result of municipal, industrial, and agricultural effluents have also contributed to the decline of this species (03,04,08). Channel dredging for maintenance and/or construction projects has specifically Management Practices - 1 (DRAFT) - Management Practices Species MUSSEL, PEARLY, HIGGINS' EYE Species Id ESIS403001 Date 13 MAR 96 been identified as a problem (04). Aside from the direct physical removal of L. higginsi from its substrate, dredging poses a number of other potential problems (04). These are alteration and destruction of habitat and increases turbidity and subsequent sedimentation (03,04). Prolonged and increased turbidity caused by boat prop wash has the same detrimental effects (05). Other possible factors for the decline of the Higgins' eye pearly mussel are: unavailability of appropriate glochidial hosts (09), disease induced by bacteria and protozoans (09), and competition by the introduced Asiatic clam Corbicula fluminea (03,10,11). APPROVED PLAN: U.S. Fish and Wildlife Service. 1983. Higgins' Eye Mussel Recovery Plan. U.S. Fish and Wildlife Service, Twin Cities, MN. 69 pp. The primary objective of the recovery plan is to remove L. higginsi from the Federal Endangered Species List. This involves the following actions: 1. Preserve the species and its essential habitat by determining habitat requirements and the effects of habitat alterations such as dredging, runoff, various discharges (pollutants), and drastic changes in the water level. 2. Habitat improvements, such as fish runways around locks and dams, may be needed to aid in dispersal of the mussels. 3. Study potential detrimental and or beneficial effects of various commercial mussel collection methods and establish reasonable regulatory measures. 4. Control/restrict barging operations and power boating activities. 5. Determine the best transplanting methods and restore the species to unoccupied habitat. Management Practices - 2
                                   (DRAFT) - References
                           Species MUSSEL, PEARLY, HIGGINS' EYE
                                  Species Id ESIS403001
                                      Date 13 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 Burch, J.B. 1975. Freshwater Unionacean clams (Mollusca:Pelecypoda) of North America. Malacological Pub., Hamburg, Michigan. 204 pp. 02 Parmalee, P.W. 1967. The freshwater mussels of Illinois. Ill. State Mus. Pop. Sci. Ser. 8:1-108. 03 Fuller, S.L. 1978. Fresh-water mussels of the upper Mississippi River. Report to U.S. Army Corps of Engineers. 04 U.S. Fish and Wildlife Service. 1983. Higgins' Eye Pearly Mussel Recovery Plan. U.S. Fish and Wildlife Service. 69 pp. 05 Wisconsin Dept. Nat. Resour. 1980. Petition to the Director, U.S. Fish and Wildlife Service to designate critical habitats. Wisc. DNR Report. 06 Baker, F.C. 1928. The freshwater mollusca of Wisconsin, Part II: Pelecypoda. Bull. Wis. Geol. and Nat. Hist. Surv. No. 70:1-495. 07 Knott, M.J. 1980. The pearl button industry and its impacts on Mississippi River mussel fauna. Pages 11-16. IN: Proceedings of the UMRCC Symp. on Upper Mississippi River Bivalve Mollusks. Rock Island, IL. 08 Engel, J.M. 1980. The Endangered Species Act, 1979. IN: Proceedings of the UMRCC Symp. on Upper Mississippi River Bivalve Mollusks. Rock Island, IL. 09 Ellis, M.M. 1931. Some factors affecting the replacement of the commercial freshwater mussels. U.S. Bur. Fish. Circ. 7:1-10. 10 Fuller, S.L.H. and M.J. Imlay. 1976. Spatial competition between Corbicula manilensis (Philippi), the Chinese clam (Corbiculidae), and freshwater mussels (Unionidae) in the Waccamaw River basin of the Carolinas (Mollusca:Bivalvia). ASB Bull. 23:60. 11 Fuller, S.L.H. and J.W. Richardson. 1977. Amensalistic competition between Corbicula manilensis (Philippi), the Asiatic Clam (Corbiculidae) and freshwater mussels (Unionidae) in the Savannah River of Georgia and South Carolina (Mollusca:Bivalvia). ASB Bull. 24:25. 12 Coker, R.E., A.F. Shira, H.W. Clark, and A.D. Howard. 1921. Natural history and propagation of fresh water mussels. Bull. Bur. Fish. 36:77-181. 13 Wilson, C.B. 1916. Copepod parasites of freshwater fishes and their economic relations to mussel glochidia. Bull. Bur. Fish. 34:331-374. 14 Witter, F.M. 1883. The Mollusca of Muscatine County and Vicinity. The Muscatine Conchological Club, Muscatine, Iowa. S.E. Cassino and Company, Boston. 24 pp. 15 Surber, T. 1913. Notes on the natural hosts of freshwater mussels. Bull. Bur. Fish. Doc. 771:1-10. 16 Pratt, W.H. 1876. List of land and freshwater shells found at Davenport, Iowa. Proc. Dav. Acad. Nat. Sci. 1:165-167. 17 Rasmussen, J.L. 1979. Distribution and relative abundance of Upper Mississippi River fishes. IN: A compendium of fishery information on the Upper Mississippi River, 2nd ed. J.L. Rasmussen, editor. Upper Mississippi River Cons. Comm., Rock References - 1 (DRAFT) - References Species MUSSEL, PEARLY, HIGGINS' EYE Species Id ESIS403001 Date 13 MAR 96 Island, IL. 18 Fuller, S.L.H. 1974. Clams and mussels (Mollusca:Bivalvia). Pages 215-273. IN: Pollution Ecology of Freshwater Invetebrates. C.W. Hart and S.L.H. Fuller, eds. Academic Press, NY. 19 Churchill, E.P., Jr., and S.I. Lewis. 1924. Food and feeding in freshwater mussels. Bull. U.S. Bur. Fish. 39:439-471. ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 U.S. Fish and Wildlife Service. 1983. Higgins' Eye Pearly Mussel Recovery Plan. U.S. Fish and Wildlife Service. 69 pp. 02 Havlick, M.E. and L.L. Marking. 1981. A quantitative analysis of naiad mollusks from the Prarie du Chien, Wisconsin dredge material site on the Mississippi River. Bull. Am. Malacolog. Union 1980:30-34. References - 2