(DRAFT) - Taxonomy
                              Species BARBERRY, TRUCKEE
                                Species Id ESIS701014
                                   Date 13 MAR 96



TAXONOMY

NAME - BARBERRY, TRUCKEE OTHER COMMON NAMES - BARBERRY, TRUCKEE; BARBERRY, SONNE'S; MAHONIA, SONNE'S; MAHONIA and TRUCKEE ELEMENT CODE - CATEGORY - Angiosperm PHYLUM AND SUBPHYLUM - MAGNOLIOPHYTA, CLASS AND SUBCLASS - MAGNOLIOPSIDA, ORDER AND SUBORDER - RANUNCULALES, FAMILY AND SUBFAMILY - BERBERIDACEAE, GENUS AND SUBGENUS - MAHONIA, SPECIES AND SSP - SONNEI, SCIENTIFIC NAME - MAHONIA SONNEI AUTHORITY - TAXONOMY REFERENCES - COMMENTS ON TAXONOMY - Truckee Barberry Mahonia sonnei Abrams KINGDOM: Plant GROUP: Angiosperm DIVISION: Magnoliophyta CLASS: Magnoliopsida ORDER: Ranunculales FAMILY: Berberidaceae The following is a revised description by M. P. Yoder-Williams prepared from all available herbarium species from the type locality. This description will be published in a forthcoming taxonomic review by M. P. Yoder-Williams. Berberis sonnei (Abrams) McMinn, Ill. Manual Calif. Shrubs, 123. 1939. (Mahonia sonnei Abrams, Phytologia 1:89-94, 1934.) Creeping or ascending low shrub, 10-40 cm tall; leaves odd-pinnately compound, 10.5-26.5 cm long and 7.0-13.5 cm wide, with (3-) 5-7 (-9) leaflets; leaflets ovate-elliptic to broadly lanceolate in outline, primarily pinnately to sub-palmately veined, 4.0-8.5 (-11) cm long and 2.2-6.5 cm wide, acute to obtuse at tip and obliquely cuneate to obtuse at base, margins serrate and undulate with (7-) 12-24 (-26) short spines per margin, abaxial surface microscopically (60X) papillate, adaxially glossy to dull-glossy and dull to slightly Taxonomy - 1 (DRAFT) - Taxonomy Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 glossy below; racemes terminal or axillary, 15-45 (-60) mm long, densely yellow flowered; flowers borne on bracteate pedicels 7-8 mm long with an early deciduous prophyll at mid-length (approx. 5 pct. of the pedicels bearing prophylls at anthesis); sepals 6 in two series of three, the outer 3-3.5 mm long, the inner 4.5-5.0 mm long; petals 6 in two series of three, the outer longer, 6-7 mm long, with the inner 5-6 mm long, both petals and sepals deciduous upon maturation of the fruit; stamens 6 opposite the petals; berries blue-black, glaucous, ellipsoid to 6 mm long; flowering mid-April to late May. Due to considerable taxonomic confusion regarding this species (02,03,06), distribution of this species is difficult to determine. Given our present knowledge and the taxonomic methodology attempted to date, the characters typically used to differentiate Mahonia (=Berberis) sonnei from B. repens do not consistently separate these entities when all the variation in the latter species is considered (20). B. repens is a widespread species exhibiting considerable morphological variation throughout its extensive range (California east to Arizona, north to Montana and Alberta, and west to British Columbia and Washington, then south to Oregon and California tending to occur mostly east of the Cascades and Sierran crest). No precise boundaries can be drawn between B. repens and B. aquifolium of the Pacific Northwest. B. repens recently has been reduced to varietal status under B. aquifolium (18,19). B. sonnei (Mahonia sonnei) may represent the intergradation between the two above-mentioned widespread species (06). Such intergradation is also seen in northern Idaho where clear delimination of B. repens or B. aquifolium is not possible. The latest Federal listing (50 CFR 17.12) listed this species as Mahonia sonnei (=Berberis s.). Location of Specimens: Type specimen, C. F. Sonne 11, Rocky banks [of] Truckee River, Nevada Co., California, April 1885 & August 11, 1884, DS 95828, now at CAS. Additional Specimens: J.L. Thompson s.n., Truckee, Nevada Co., CA, CAS 317107. C. Kramer 73-2, near Truckee River from river 50 ft among rocks, 150 yards W of Hwy 267 and West River St., CAS 565539. G.H. True, [ditto location], CAS 673175, CAS 673176. C.F. Sonne s.n., Truckee, banks south of factory, Nevada Co., CA, UC 1992553. C.F. Sonne s.n., Truckee, dry stony ground, Nevada Co., CA, UC 9742. C.F. Sonne, Truckee River at Truckee, Nevada Co., CA, UC 118110. B. Trowbridge 5956, Bank of the Truckee River, Nevada Co., CA, SFSU. T. Anderson, Truckee, banks of the river 150 yards west of Hwy 267, Nevada Co., CA, HSU. J.B. Roof s.n., [ditto location], JEPS 73273. M.E. Jones s.n., Soda Springs [probably at Truckee], Nevada Co., CA, POM 97706. Other descriptions of the Truckee barberry can be found in 01 and Taxonomy - 2 (DRAFT) - Taxonomy Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 02 (illustrated), 03 (without keys), 05, and 16. Photographs are located at the U.S. Fish and Wildlife Service, Endangered Species Field Station, Sacramento; at the California Department of Fish and Game, Endangered Plant Program, Sacramento, CA; and with M. P. Yoder-Williams, Department of Botany, University of Washington, Seattle, WA 98195. Living plant material may be seen at the type locality on the banks of the Truckee River west of State Route 267 and south of Riverside Drive and in cultivation at Tilden Botanic Garden, East Bay Regional Parks, Berkeley, CA. Cornflower Farms continues to successfully propagate cuttings of the Truckee barberry. Other common names for the Truckee barberry are Sonne's barberry, Sonne's mahonia, and Truckee mahonia. Taxonomy - 3
                                  (DRAFT) - Status
                              Species BARBERRY, TRUCKEE
                                Species Id ESIS701014
                                   Date 13 MAR 96



STATUS

Coded Status E: Federal Endangered COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The Truckee barberry (Mahonia sonnei) has been designated an Endangered species pursuant to the Endangered Species Act of 1973 (50 CFR 17.12; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The species has this status wherever found including the State of California. Removal and reduction to possession of any Federally listed plant from an area under Federal jurisdiction is unlawful (50 CFR 17.61 and 17.71). RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. USFS -Responsible for the law enforcement/protection of this species with applicable State and Federal laws on public lands under their control. Also responsible for management/recovery on Forest Service lands. The Forest Service is responsible for integrating management, protection, and conservation of Federally listed species into the Forest Planning process (36 CFR 219.19 and 219.20). This species has possible occurrence on the Modoc National Forest, CA. All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely to jeopardize the continued existence of the species or result in the destruction or adverse modification of Critical Habitat (50 CFR 402), and to utilize their authorities to carry out programs for the conservation of the species. STATE STATUSES AND LAWS: STATE: California DESIGNATED STATUS: Endangered ADMINISTRATIVE AGENCY: California Department of Fish and Game STATE STATUTE: Calif. Endangered Species Act, Fish and Game Code, Chapter 1.5, Article 1, Sec. 2050. INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: None. ECONOMIC STATUSES: Status - 1 (DRAFT) - Status Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 None. 75/07/01:40 FR 27823/27924 - Smithsonian report, review of status 76/06/16:41 FR 24524/24572 - Proposed rule, Endangered 79/11/06:44 FR 64246/ - Final rule, listed as Endangered 85/07/22:50 FR 29901/29909 - Five year review Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - TERRESTRIAL INLAND AQUATIC SOCIETY OF AMERICAN FORESTRY TYPES SAF TYPE STAGE CLOSURE Interior Ponderosa Pine shrub--seedling Interior Ponderosa Pine young tree Interior Ponderosa Pine mature tree Interior Ponderosa Pine Old Growth LAND USE - Evergreen Forest Land Residential Cropland and Pasture Shrub and Brush Rangeland Streams and Canals Forested Wetland Nonforested Wetland NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Palustrine SS1 Palustrine FO1 COMMENTS ON HABITAT ASSOCIATIONS - The only known habitat of the Truckee barberry is a disturbed streamside forest area behind (within 15 m of) houses. The species appears to be restricted to the flood plain of the river, although this may be an artifact of the degree of disturbance above the flood plain. The canopy is fairly open and sporadic in cover, composed of a number of large Populus trichocarpa and less so of Prunus virginiana var. demissa and Salix lutea. One planted apple tree (Malus) contributes to the canopy cover of one of the five patches of the Truckee barberry. Two of the patches have little canopy overstory and are scattered in open exposed areas among grasses and weedy herbaceous species. Rosa woodsii var. ultramontana is common in association with the Truckee barberry. The soil throughout the Truckee population is a rocky sandy loam and exhibits poorly defined horizons (13). It is derived from granitic and volcanic alluvium (06,13). Annual alluvial deposition onto the site can be considerable depending on the spring snowmelt-induced flooding. The site is highly disturbed from domestic animal burrowing, clearing for garden sites, rubbish dumping, residential development, and recreational use. Intense past human use of the area has likely altered the natural vegetation of the site from one originally dominated by yellow pine/riparian forest to one now consisting of ruderal species with a remnant wetland forest canopy. Contrary to Roof and Howard (10), the species does appear to require a close association with abundant subsurface water. Their discussion to the contrary was based on the so-called "xeric" site of a transplanted individual of Berberis sonnei (Mahonia sonnei) at the Tilden Botanical Garden in Berkeley, CA. This site is situated downslope from a watered lawn area and likely has abundant subsurface water available throughout the year (06). It in no way approaches the xeric sites of the eastern Sierra Nevada and provides no information towards an assessment of the tolerance of the Truckee barberry to Habitat Associations - 1 drought stress (06). B. repens, the more xeric barberry of the western U.S., is always topographically associated with drainages of subsurface moisture. A similar situation would be expected for the Truckee barberry. The species could be (and probably has been in the past) adversely impacted by pasture for livestock (especially horses) if such use was made of the area behind the houses and by the river's edge in Truckee. Such use would result in rapid loss of the plants in the area immediately affected. Considering the intensity of disturbance on this site over the past 130 years, it is difficult to discuss the natural habitat associated with this species. Present disturbance to the site continues and is likely to increase in intensity and degree as economic growth of the Truckee area continues at a rapid pace. New home construction along Riverside Drive has quickened the last 5 years with larger, more elaborate homes being built on the sites of previous small houses (06). Regarding possible other sites in the Truckee River drainage, it seems likely that any other populations of the species were lost in the recent past. With the development of the Central Pacific (subsequently the Southern Pacific) Railroad in the 1860-70's, followed by large-scale logging flume systems, irrigation systems, highway and then freeway construction down the narrow Truckee River canyon, large-scale extirpation of the species, if it occurred there, would have resulted. Extensive surveys of the side canyons of the Truckee River from Deer Creek downstream to Verdi in 1985 failed to turn up any other populations of the species. One noticeable character of this watershed observed during the surveys was the degree of disturbance throughout the area, both from direct anthropogenic causes as well as indirectly from man-introduced grazers. Specific grazing by the introduced and successfully naturalized beaver population on shrub and tree species is extremely high. During the early fluctuation period characteristic of introduced animal population, the beaver population of the Truckee and Feather Rivers rose quite high followed by rapid crashes, then rising again. Heavy grazing during this time may have removed all palatable shrubs from within 150-250 m of the river banks. A plant such as B. sonnei which appears to be restricted to river banks or stream systems would have been rapidly decimated. Its only known occurrence in the backyards of a dense housing site, where beavers may have been less likely to forage far from the safety of the water, may support the idea that grazing was the overall factor in its loss in the watershed. Other possible habitat associations could include seepage areas or arid riparian sites in the eastern Sierra Nevada and adjacent Great Basin ranges. This is especially true for isolated wet areas in the yellow-pine sagebrush belt extending north into Modoc County, CA, and possible adjacent Oregon and Nevada. Habitat Associations - 2
                                (DRAFT) - Food Habits
                              Species BARBERRY, TRUCKEE
                                Species Id ESIS701014
                                   Date 13 MAR 96



FOOD HABITS

TROPHIC LEVEL - AUTOTROPH Food Habits - 1
                         (DRAFT) - Environment Associations
                              Species BARBERRY, TRUCKEE
                                Species Id ESIS701014
                                   Date 13 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G G Aquatic Features: Springs [flowing] G Environment Associations - 1
                               (DRAFT) - Life History
                              Species BARBERRY, TRUCKEE
                                Species Id ESIS701014
                                   Date 13 MAR 96



LIFE HISTORY

HABIT: The Truckee barberry is a shrub (01,02,05). LIFE CYCLE: Perennial (01,02,05). The species is a low-growing (less than 1 m tall), rhizomatous shrub and can persist on a site vegetatively for likely 50+ years (06). At the present time, no sexual reproduction has been observed as no viable seeds have been documented. No seedling establishment has been observed (06,10). TYPE OF REPRODUCTION: Sexual reproduction appears possible if outcrossing is made. Viable pollen was determined in 1985 (06), but racemes were few (2) and flowers aborted or were grazed off before the end of the growing season in 1985. No viable fruit have been documented (06,10). Vegetative spread does occur via rhizomes. However, degree of spreading vegetatively at present seems minimal due to small shoots and continued disturbance of the site (06). REPRODUCTIVE PHENOLOGY: Phenology varies considerably with snowpack melt-off. Little is known of fruit dispersal as viable fruit are rarely, if at all, produced. Germination dates are unknown but likely follow snowmelt in the spring. Leafing and budding occurs from April 1 to May 15. Anthesis dates range from April 15 to May 30 and fruiting from May 30 to July 1. Dispersal of fruit occurs after May 30. Suggestion of mature fruit (03,14) may be in error, and may actually refer to enlarged but abortive ovaries. Past fruit collections for seed have yielded no viable seed (06,10). SEX OR SPORE STATUS: Monoclinous. (01,02,05,06,12,16). The species is believed to require out-crossing for viable seed (06). Plants at Truckee may represent as few as two genetic individuals, and possibly as many as five. However, this may represent a very low gene pool and may therefore represent a population that is no longer capable of producing viable seed. POLLINATION, SPORE AND SEED DISSEMINATION: Pollen is disseminated by insects of the genus Bombus and other large insect visitors including Apis melifera (domestic honey bee) (06). Viable pollen tests were made in 1985. Pollen viability exceeded 95% (06). Human collecting of berries is likely and wasting of the fruit would result. Bird foraging on the berries is possible although the low fruit production probably prevents selective foraging by birds to any extent. Rodent foraging is also likely (03,06). SEED BIOLOGY: At present, nothing is known of the seed biology of this species. Viable looking seed was obtained in 1981 (14), but no germination resulted. Early seed collections gave the same results (10). Life History - 1 (DRAFT) - Life History Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 Information of seed biology from the related species, B. repens, is likely similar for the Truckee barberry. Viability of seed for B. repens has been determined to be at least 5 years (09). Germination for B. repens is undertaken as follows: Clean seed from fruit and dry; store in sealed containers at slightly above freezing; pretreat with successive cold, warm, cold stratification periods; germinate at 1.1 degrees C (09). POPULATION BIOLOGY: Due to the extensive and unprotected use of the remaining site at Truckee (for river access, fishing access, gardening, feral or domestic animal activity, housing construction, dumping, etc.) the population there appears to be declining. Detailed mapping in 1985 (06) will allow for periodic resampling and inventory of the individual stems. The location of two additional patches in 1985 has expanded the areal extent of the population to approximately 200 meters long by 15 meters wide with five localized patches present. Individual patch size does not exceed 10 meters in diameter. Considering the ability of this species to spread vegetatively, it is likely that five or fewer genotypes exist and therefore sexual reproduction may be limited by insufficient out-crossing or low pollinator activity between distinct patches. Pollinating extremes of the population should be attempted if flowers are available. ECOLOGICAL/EDAPHIC FACTORS: Gentle, south-facing slopeof river flood plain in open or shaded situations under a reparian canopy (Populus trichocarpa-Salix-Prunus) at 1770 m elevation; subsurface soil water is available throughout most of the year; soils derived from quaternary nonmarine terrace deposits (11) being primarily recently deposited alluvium to soils of the Jorge-Tahoma Series, very stoney sandy loams of the Alfisol Order and Typic Cryoboralfs (13). Site temperatures range from -6 to 35 degrees C during the growing season with mean temperatures approaching 10-15 degrees C. Winter extremes (5 to -35 degrees C are moderated by the snowpack (+200 cm) where subnivean temperatures remain near freezing (06). Disturbance is considerable at the site expecially in the form of foot traffic and garden site clearing. Natural disturbance in the form of periodic flooding and alluvial deposition is a characteristic of the riparian vegatation and it is likely that the Truckee barberry evolved under such disturbance. TROPHIC STATUS: Phototrophic (01,02,05,16). Mycorrhizal root symbionts are suspected, especially vesicular-arbuscular types, but remain unknown at the present (06). CHARACTERISTIC DOMINANCE: The Truckee barberry is a minor component (less than 1% cover) of the understory of the riparian vegetation. This is especially true due to the poor condition of the surviving shoots. Plantings or introduction of rye grass and other exotics may have further reduced the condition or spread of the remaining shoots via decreased Life History - 2 (DRAFT) - Life History Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 availability of nutrients. Foliar fertilization in 1985 by the U.S. Fish and Wildlife Service appeared to enhance the growth of the stems and a greening of the leaves (06).] Information relating to the cover of B. repens in the Great Basin may shed light on the original role of the Truckee barberry in the eastern Sierra Nevada. COMMUNITY ECOLOGY: The Truckee site represents a highly disturbed, riparian forest site. Once probably closely associated with the yellow pine forest, but due to residential development since the 1880's, the riparian habitat has become isolated and further fragmented. Regeneration of the Populus is unlikely, as only mature individuals remain on the site with little or no replacement (06). Further disruption of the understory is likely. Early photographs of this area in the 1880-1890's (10,15) show the site much more disturbed than at the present with no canopy existing on this particular area and buildings built right to the high water mark. Future disturbance to the site again seems likely as development of the area expands. Transplanting attempts would only succeed if sites of similar characteristics could be found (without the man-caused disturbance) and in the absence of grazing by beavers. SPECIES INTERRELATIONSHIPS: Beaver (Castor canadensis) may selectively graze the Truckee barberry when compared to other less palatable shrubs common in this area. Introduction of the beaver in the 1930's (08) likely brought about loss of some populations of the plant elsewhere. Beaver can cause complete death of aspen patches by repeated and intense cutting over an extended period of time (06). With added grazing by ungulates, loss of a patch could have resulted within a ten year period (06). OTHER LIFE HISTORY DESCRIPTORS: The western barberries are not a host for the wheat rust (Puccinia graminis) (02,03), although a unidentified leaf rust was observed in 1985 on the Truckee barberry at Truckee (06). Life History - 3
                           (DRAFT) - Management Practices
                              Species BARBERRY, TRUCKEE
                                Species Id ESIS701014
                                   Date 13 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Land Acquisition Beneficial Controlling/Removing Nonnative Vegetation Beneficial Controlling/Removing Native Vegetation Beneficial Stocking captive-reared wild-strain animals Beneficial Transplanting wild animals Beneficial Transplanting Wild Eggs/Wild Seeds Beneficial Controlling/Removing Feral Animals Beneficial Controlling/Removing Domestic Animals Adverse Collecting Existing Collecting Adverse Low Gene Pool Existing Low Gene Pool Adverse Predation Existing Predation Adverse Rural Residential/Industrial Areas Existing Rural Residential/Industrial Areas Adverse Highway/Railroads Existing Highway/Railroads Adverse Soil compaction by heavy equipment in mine areas Existing Soil compaction by heavy equipment in mine areas Adverse Dredging Existing Dredging Adverse Developing/maintaining stream bank vegetation Existing Developing/maintaining stream bank vegetation Adverse Exotic/Feral/Introducted Species Existing Exotic/Feral/Introducted Species Adverse Grazing Existing Grazing Adverse Harvesting Existing Harvesting COMMENTS ON MANAGEMENT PRACTICES - The Truckee barberry has likely disappeared from the Tuckee watershed and possibly elsewhere due to intense grazing by introduced ungulates (sheep, cattle, and horses) as well as the introduced beaver (Castor canadensis), which was brought into the watershed (and the upper Feather River watershed) as a game animal in the 1930's (08). Thoughout the area, grazing impacts by both beaver and cattle/horses were common, (e.g. eroded banks, high degree of siltation, dead willow clumps, trampling of the vegetation, etc). Considerable modification of the riparian areas of these drainages has occurred over the last 130 years as the logging industry grew, especially in the 1880-1920's. All portions of the Truckee drainage experienced considerable site modification with clearcutting practices common throughout the area. The entire riparian area of the main channel of the Truckee River was highly modified as flume systems, mill sites, irrigation systems, and ice production plants were developed before the turn of the century (06). Management Practices - 1 (DRAFT) - Management Practices Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 With the development of the Central Pacific (subsequently the Southern Pacific) Railroad in the 1860-70's, followed by large-scale logging flume systems, irrigation systems, highway and then freeway construction down the narrow Truckee River canyon, large-scale extirpation of the species (if it occurred there), would have resulted. Extensive surveys of the side canyons of the Truckee River from Deer Creek downstream to Verdi in 1985 failed to turn up any other populations of the species. One noticeable character of this watershed observed during the surveys was the degree of disturbance throughout the area, both from direct anthropogenic causes (i.e., changes to the river channel) as well as indirectly from man-introduced grazers (e.g., beaver, cattle and/or horses). Specific grazing by the introduced and successfully naturalized beaver population on shrub and tree species is extremely high. During the early fluctuation period characteristic of introduced animal population, the beaver population of the Truckee and Feather Rivers rose quite high followed by rapid crashes, then rising again. Heavy grazing during this time may have removed all palatable shrubs from within 150-250 m of the river banks. A plant such as the Truckee barberry which appears to be restricted to river banks or stream systems would have been rapidly decimated. Its only known occurrence in the backyards of a dense housing site, where beavers may have been less likely to forage far from the safety of the water, may support the idea that grazing was the overall factor in its loss in the watershed. In addition, possible overcollecting by amateur gardeners or by botanists may have further impacted the remaining plants of this species. The development of small garden plots or landscaping projects have resulted in loss of stems at Truckee even in the last few years (06). At present, the small gene pool of the remaining population may prevent production of viable seeds. The plants at Truckee may include only two to five genetic individuals and represent a population no longer capable of producing seed. Due to the extensive and unprotected use of the remaining site at Truckee (for river access, fishing access, gardening, feral or domestic animal grazing, housing construction, dumping, etc.) the population there appears to be declining. Disturbance is considerable at the site expecially in the form of foot traffic and garden site clearing. Early photographs of this area in the 1880-1890's (10,15) show the site much more disturbed than at the present with no canopy existing on this particular area and buildings built right to the high water mark. Future disturbance to the site again seems likely as development of the area expands. APPROVED PLAN: U.S. Fish and Wildlife Service. 1984. Recovery Plan for Truckee Barberry [Berberis (=Mahonia) sonnei (Abrams) McMinn]. U.S. Fish and Wildlife Service, Portland, OR. 68 pp. Actions recommended in the Truckee barberry Recovery Plan include: Management Practices - 2 (DRAFT) - Management Practices Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 1) Maintain and protect the single population which occurs on private land. 2) Assure access, and obtain cooperation through agreements with adjacent land owner to insure the safety of the population. 3) Monitor population, and manage habitat to reduce competition from native and/or non-native plants, maintain vigor, and encourage regeneration. Erect a fence to protect the habitat from domestic and/or feral animals. 4) Establish additional populations through propagation from revetative materials and seeds. Determine safe collecting levels, and store at nurseries or repositories for holding plant materials, and storing seeds. 5) Increase seed production, and identify optimum habitat. 6) Transplant revegetative stock or seed to establish at least five new populations, monitor and protect new populations. 7) Conduct searches for undiscovered populations. Management Practices - 3
                                   (DRAFT) - References
                                Species BARBERRY, TRUCKEE
                                  Species Id ESIS701014
                                      Date 13 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 Abrams, L.E. 1944. An illustrated flora of the Pacific States, Volume 2. Stanford University Press, Stanford. 218 pp. 02 McMinn, H.E. 1951. An illustrated manual of California shrubs. University of California Press, Berkeley. 663 pp. 03 Roof, J.B. 1974. Found alive: the Truckee barberry. Four Seasons 4:1-18. 04 U.S. Government. 1979. Determination that Berberis sonnei (the Truckee barberry) is and Endangered species. Federal Register 44(216):64246/64247. 05 Abrams, L.E. 1934. The mahonias of the Pacific states. Phytologia 1:89-94. 06 Yoder-Williams, M.P. Unpublished data. Department of Botany, KB-15, University of Washington, Seattle, WA 98103. 07 Takhtajan, A.L. 1980. Outline of the classification of flowering flowering plants (Magnoliophyta). Bot. Rev. 46:225-359. 08 Tappe, D.T. 1942. The status of beavers in California. Calif. Dept. Fish Game, Game Bull. No. 3. 59 pp. 09 Vories, K.C. 1980. Growing Colorado plants from seed: a state of the art. Volume 1: Shrubs. USDA Forest Service Gen. Tech. Rept. INT-103. 80 pp. [Available from: Intermountain Forest and Range Exper. Stn., Ogden, UT 84401.] 10 U.S. Fish and Wildlife Service. 1984. Recovery Plan for the Truckee Barberry [Berberis (=Mahonia) sonnei (Abrams) McMinn]. U.S. Fish and Wildlife Service, Portland, OR. 68 pp. 11 Burnett, J.L. and C.W. Jennings. 1962. Geological map of California, Chico Sheet. Calif. Div. Mines and Geology, Sacramento, CA. 12 Lawrence, G.H.M. 1951. The taxonomy of vascular plants. MacMillian Co., New York. 823 pp. 13 Rogers, J.H. 1974. Soil survey of the Tahoe Basin area, California and Nevada. USDA, Soil Conservation Service. 14 Demming, B.E. 1985. Phenological notes and personal communication. P.O. Box 963, Kings Beach, CA 95719. 15 Meschery, J. 1978. Truckee: An illustrated history of the town and its surroundings. Rocking Stone Press, Truckee, CA 95734. 114 pp. 16 Munz, P.A. 1959. A California flora. University of California Press, Berkeley. 108 pp. 17 Jepson, W.L. 1922. A flora of California, Part VII. Associated Students Store, Univ. Calif., Berkeley. 18 Scoggan, H.J. 1978. The flora of Canada, Part 1, General survey-nomenclatural changes. National Museum of Natural Sciences, National Museums of Canada, Ottawa. 19 Biovin, B. 1966. Enumeration des plantes du Canada. Naturaliste Can. 93:642. 20 Yoder-Williams, M. [In Prep.] Taxonomy of Truckee barberry, Berberis sonnei (Berberidaceae). Available from: California Department of Fish and Game, Sacramento. References - 1 (DRAFT) - References Species BARBERRY, TRUCKEE Species Id ESIS701014 Date 13 MAR 96 ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 Yoder-Williams, M.P. Unpublished data. Department of Botany, KB-15, University of Washington, Seattle, WA 98103. 02 Abrams, L.E. 1934. The mahonias of the Pacific states. Phytologia 1:89-94. 03 U.S. Fish and Wildlife Service. 1984. Recovery Plan for the Truckee Barberry [Berberis (=Mahonia) sonei (Abrams) McMinn]. U.S. Fish and Wildlife Service, Portland, OR. 68 pp. 04 Lenz, L. July 2, 1985. Personal communication. Rancho Santa Ana Botanic Garden, Clarement, CA. 05 Roof, J.B. 1974. Found alive: the Truckee barberry. Four Seasons 4:1-18. References - 2