(DRAFT) - Taxonomy
                          Species CACTUS, FISHHOOK, TOBUSCH
                                Species Id ESIS702006
                                   Date 13 MAR 96



TAXONOMY

NAME - CACTUS, FISHHOOK, TOBUSCH OTHER COMMON NAMES - CACTUS, FISHHOOK, TOBUSCH; ANCISTROCACTUS and BANDERA COUNTY ELEMENT CODE - CATEGORY - Angiosperm PHYLUM AND SUBPHYLUM - MAGNOLIOPHYTA, CLASS AND SUBCLASS - MAGNOLIOPSIDA, ORDER AND SUBORDER - CARYOPHYLLALES, FAMILY AND SUBFAMILY - CACTACEAE, GENUS AND SUBGENUS - ANCISTROCACTUS, SPECIES AND SSP - TOBUSCHII, SCIENTIFIC NAME - ANCISTROCACTUS TOBUSCHII AUTHORITY - TAXONOMY REFERENCES - COMMENTS ON TAXONOMY - Tobusch Fishhook Cactus Ancistrocactus tobuschii (W.T. Marsh.) W.T. Marsh. ex Backeberg KINGDOM: Plant GROUP: Angiosperm DIVISION: Magnoliophyta CLASS: Magnoliopsida ORDER: Caryophyllales FAMILY: Cactaceae The Tobusch fishook cactus has fibrous roots and short taproots, (01). The stems are normally solitary, sometimes branching due to injury, dark green, obconical or turbinate, 5 to 8 cm long, and 2.5 to 8.5 cm in diameter. The ribs are usually 8 in number but poorly defined, consisting of slightly confluent tubercles 6 to 9 (10 to 12) mm long (02), 6 mm broad, and protruding 5 to 12 mm. The areoles are elliptical to linear, the spine-clusters are about 4.5 mm in basal diameter and 9 to 12 mm apart. The spines are rather sparse, finely scaberulous-canescent. The central spines number 3 to 5, are light yellow with red tips fading to gray, the upper 2 to 3 are straight, erect or ascending; the lower one is hooked and spreading, the upper ones are longer, 2.2 to 3.75 cm long and somewhat flattened. The radial spines number 7 to 12, are slender and spreading irregularly, 12 to 19 mm long and are articular. The flowers are 3 to 4 cm long and wide. The outer floral bracts show brownish-red midribs and pale Taxonomy - 1 (DRAFT) - Taxonomy Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 yellow margins, they are elliptic-oblong, 12 to 15 mm long, about 6 mm broad, apically rounded and entire. The inner floral bracts are yellow to cream, nearly spatulate, about 15 mm long, and about 4.5 mm broad, entire or irregularly dentate. The filaments are cream to yellowish, 6 mm long. The anthers are a golden-yellow, about 0.7 mm long. The style is green, about 17 mm long. Stigmas are yellowish, 5 to 9 lobed and the lobes are about 1 mm long. The fruit is green, flushing pink or a pinkish brown when ripe, ovid or ellipsoid, 25 to 30 mm long, and about 9 to 15 mm wide, with small scales on the fruit, these broad, membranous, and irregularly lobed. The seeds are blackish, finely papillate, 1.5 mm long, 1.5 mm or more broad and 1 mm thick (01,02,03,04,05,06). Ancistrocactus tobuschii is closely releated to A. brevihamatus, which replaces tobuschii farther south and west; these two taxa intergrade in Kinney County, TX (09). The westernmost population of tobuschii slightly approaches A. brevihamatus in spine morphology, but has typical tobuschii flowers (09). The typical differences between A. tobuschii and A. brevihamatus are: 1) flower color (petaloid bracts dull golden-yellow, or golden- tan, or greenish-cream-colored in tobuschii, as opposed to pinkish brown or pink and green in brevihamatus) and 2) flower shape (funnelform-salverform in tobuschii, but funnelform-cylindroid in A. brevihamatus) (08). However, tobuschii flowers go through a funnelform-cylindroid period as they open and close, so their characteristic shape pertains only to fully open flowers (09). The vegetative differences supposedly characteristic of tobuschii (relatively small size of all parts, and greener stem surface) are not useful (09). Most tobuschii plants in the vicinity of the type locality are indeed very small relative to the average A. brevihamatus that one sees farther west, but this is due only to population structure (09). The tobuschii populations near the City of Vanderpool, TX, consist predominantly of seedlings with a few individuals living to old age; most brevihamatus populations, in contrast, are reproducing very poorly and consist of widely separated old adults (09). The few immatures seen in brevihamatus populations appear almost identical to typical plants of tobuschii but sometimes have gray-green (instead of dark green) stem surfaces (09). The color of the stem surface depends on the environment in which the plant is growing; i.e., increasingly gray with increasing isolation and decreasing moisture (09). The tobuschii all grow in a relatively humid region, and so they are relatively green; in cultivation, the differences in stem color are much weaker than they are in the wild (09). In the original description, Marshall published the name as Mammillaria (Ancistrocactus) tobuschii (02). Although the inclusion of Ancistrocactus in parentheses would indicate a subgenus, Marshall intended the names to be alternative (03). Backeberg (07) later published the species in Ancistrocactus. Weniger placed the species in the genus Echinocactus (03); however, this combination is invalid. A report by Wauer from the Dead Horse Mountains in Brewster County, TX (08), is a misidentification according to Weniger (01), Warnock (01), and Zimmerman (09). Taxonomy - 2 (DRAFT) - Taxonomy Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 The holotype (Marshall B1501) is on deposit at the Desert Botanical Garden in Tucson, AZ. Isotypes are at DES and TEX. Other specimens may be found at DES and UNM. Several references contain descriptions, photographs, and/or drawings; (01,02,03,04,05,06,10). The IUCN Red Data Book uses the common name "Bandera County Ancistrocactus" for the Tobusch fishhook cactus. Taxonomy - 3
                                  (DRAFT) - Status
                          Species CACTUS, FISHHOOK, TOBUSCH
                                Species Id ESIS702006
                                   Date 13 MAR 96



STATUS

Coded Status E: Federal Endangered Commercial Non-consumptive recreational Ornamental COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The Tobusch fishhook cactus (Ancistrocactus tobushii) has been designated an Endangered species pursuant to the Endangered Species Act of 1973 (50 CFR 17.12; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The species has this status wherever found including the State of Texas. This species is protected by the Lacey Act (P.L. 97-79, as amended; 16 U.S.C. 3371 et seq.) which makes it unlawful to possess any wild plant (including roots, seeds, and other parts) within U.S. territorial or special maritime jurisdiction (as defined in 18 U.S.C. 7); or to import, export, transport, sell, receive, acquire, or purchase in interstate or foreign commerce any wild plant (including roots, seeds, and other parts) taken, possessed, transported, or sold in violation of any State law or regulation. It is also unlawful to import, export, transport, sell, receive, acquire, or purchase any wild plant (including roots, seeds, and other parts) taken or possessed in violation of any U.S. law, treaty, or regulation or in violation of Indian tribal law. RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely to jeopardize the continued existence of the species or result in the destruction or adverse modification of Critical Habitat (50 CFR 402), and to utilize their authorities to carry out programs for the conservation of the species. STATE STATUSES AND LAWS: STATE: Texas DESIGNATED STATUS: Endangered ADMINISTRATIVE AGENCY: Texas Parks and Wildlife Department STATE STATUTE: Chapter 88 of the Texas Parks and Wildlife Code, Texas Parks and Wildlife Rules for Protected, Status - 1 (DRAFT) - Status Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 Threatened and Endangered Native Plant Species, Sec 57.401-57.413 INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: This species is listed in Appendix I of CITES and as Endangered under the common name Bandera County Ancistrocactus in the IUCN Red Data Book (1978). ECONOMIC STATUSES: This species has horticultural value and is commercially available from cactus and succulent dealers. It's rarity makes it particularly desirable to collectors. It has value as a component of Texas's natural heritage of biotic diversity. 75/07/01:40 FR 27823/27924 - Notice of review 76/06/16:41 FR 24523/24572 - Proposed Endangered 79/11/07:44 FR 64736/64738 - Listing as Endangered 85/07/22:50 FR 29901/29909 - Five year review Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - TERRESTRIAL TERRESTRIAL INLAND AQUATIC LAND USE - Mixed Rangeland Streams and Canals Bare Exposed Rock NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Riverine, intermittent SB1 Riverine, intermittent BB1 COMMENTS ON HABITAT ASSOCIATIONS - Ancistrocactus tobuschii occurs in sunlit rocky sites such as gravel bars or adjacent low limestone ledges and ridges in or near rocky, intermittent streams (01); and on open rocky hilltops and mesa- tops (09). The habitat is entirely within the Juniper-Oak Association of the Edwards Plateau, a mixed rangeland within the Texas Savanna. Platanus occidentalis and Juglans microcarpa are found in or near the streambed, while Juniperus ashei, Berberis trifoliolata, and Quercus fusiformis occur in association with Ancistrocactus tobuschii at all of the non-riparian sites (09). The cactus is usually found in full sun, although it will tolerate light shade (01). Although Weniger states that there is little grass in the immediate vicinity (01), this is true only of a fraction of the populations (09). Marshall, in the type description, said that it was usually growing in clumps of Aristida (02). Other cacti found in association with Ancistrocactus tobuschii are; Coryphantha sulcata, Coryphantha vivipara, Echinocereus enneacanthus, and Epithelantha micromeris (09,10). The topographic elevation varies between 400 and 500 meters (01, 02). The mean annual total precipitation is about 22 to 30 inches and the mean annual temperature is about 65 to 69 degrees F (11). Habitat Associations - 1
                                (DRAFT) - Food Habits
                          Species CACTUS, FISHHOOK, TOBUSCH
                                Species Id ESIS702006
                                   Date 13 MAR 96



FOOD HABITS

TROPHIC LEVEL - AUTOTROPH Food Habits - 1
                         (DRAFT) - Environment Associations
                          Species CACTUS, FISHHOOK, TOBUSCH
                                Species Id ESIS702006
                                   Date 13 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G G Terrestrial Features: Cliffs/ledges G Terrestrial Features: Rock outcrops G G G G G G G G G G Environment Associations - 1
                               (DRAFT) - Life History
                          Species CACTUS, FISHHOOK, TOBUSCH
                                Species Id ESIS702006
                                   Date 13 MAR 96



LIFE HISTORY

HABIT: Succulent (01,02,03,05,06,07,08). LIFE CYCLE: Perennial (01,02,05,06,09). Ancistrocactus tobuschii has a relatively short life-cycle (09). In cultivation they reach sexual maturity in as little as 3 years, and they give the appearance of perhaps equally rapid maturation in the field (as evidenced by adult plants having very soft underground parts, with very few nodes/internodes; in contrast to wild adults of most other species of cacti) (09). The life expectancy of wild Ancistrocactus tobuschii plants, beyond attainment of sexual maturity, probably is less than ten years (09). TYPE OF REPRODUCTION: Sexual (01,09). These plants are obligate outcrossers, reproducing exclusively by seeds (09). Although the plants branch when injured (01), the branches are sessile and are not a mechanism for disperal or establishment of new plants (09). REPRODUCTIVE PHENOLOGY: Germination and leafing dates are unknown. Budding and anthesis occur in February to early April (09). Fruiting occurs in late spring to early summer (09). Timing of Seed/fruit dispersal is unknown (09). According to Weniger (01), Ancistrocactus tobuschii flowers from late April until early June and produces ripe fruits by late summer. However, field research by Zimmerman (12) indicates that blooming starts in mid-February (early March if the weather is cold) and is over by mid-March, rarely as late as early April. The fruits ripen in late spring or early summer, although sometimes they persist until late summer (12). SEX OR SPORE STATUS: Monoclinous (03,04,05,06,07). POLLINATION, SPORE AND SEED DISSEMINATION: Pollen dissemination agents are small native bees, including Dialictus sp. (09). The seed dissemination agents are possibly birds or rodents. Concentrations of seedlings close to the adults suggest that seed dispersal often is passive (09). However, a few of the fruits disappear from the plants shortly after ripening, strongly indicative of fruit consumption (and possibly seed dispersal) by either birds or rodents (09). SEED BIOLOGY: In cultivation, the percentage of germination is high (02). POPULATION BIOLOGY: In June 1952, Marshall observed seedlings immediately around Life History - 1 (DRAFT) - Life History Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 established plants and scattered throughout the area (02). Weniger (01) stated that seedlings have not been found in nature since then. However, the populations seen by Zimmerman (09) range from those in which seedlings are very few to those in which the majority of the plants are immatures, representing every possible age-class. In 1963, about 100 individuals were observed along the Sabinal River near Vanderpool, at a density of up to 3 individuals per hundred square feet (01). But in 1979 Weniger found no Ancistrocactus tobuschii there (01). One or two dozen plants were observed along a ten mile strip in western Kerr County, TX, and a couple of dozen individuals were found on a few acres in northern Bandera County, TX, on the Upper Medina River (01). The known population has greatly declined over the past 15 years (01). The life expectancy of wild Ancistrocactus tobuschii plants, beyond attainment of sexual maturity, probably is less than ten years (09). This means that a population consisting largely of adult plants, with little evidence of recent reproduction, could completely disappear as a result of natural die-off in the few years following its discovery (09). This tendency for Ancistrocactus tobuschii populations to be ephemeral might render this taxon very difficult to manage on a single-population basis (09). At least near Vanderpool, Ancistrocactus tobuschii seems to simultaneously experience both castastrophic die-off and/or destruction of some populations, and very rapid growth of other populations (09). If Ancistrocactus tobuschii is indeed a fugitive species then very large tracts of habitat containing many emphemeral Ancistrocactus tobuschii populations would be necessary for the long-term survival of Ancistrocactus tobuschii at a given site (09). ECOLOGICAL/EDAPHIC FACTORS: Ancistrocactus tobuschii grows in open, sunny, rocky sites, although it will tolerate some shade (01,09). It can be found on gravel bars or low ledges and ridges in or near intermittent streams (01) as well as on rocky hilltops and mesa-tops (09). When on open mesa-tops, the cactus is usually in the most stable available microhabitats, such as mats of Selaginella among rock outcrops on relatively level sites where over-all drainage is good (09). The soil texture varies from very cobbly and gravelly to deep clays and loams (14,15,16,17). Most of the sites are xeric, but the sites in the intermittent streams could be considered mesic at some times. The topographic elevation varies between 400 and 500 meters (01, 02). The mean annual total precipitation is about 22 to 30 inches and the mean annual temperature is about 65 to 69 degrees F (11). Geologic formations in the area of the cactus' occurrence are the Glen Rose Formation, the Edwards Limestone, the Salmon Peak Limestone, and the Hensell Sand, all are lower Cretaceous (13). Also within the Ancistrocactus tobuschii range are the following upper Cretaceous Formations; the Del Rio Clay, the Buda Limestone, and the Cretaceous Igneous Rocks (13). Pleistocene Formations in the habitat are the Leona Formation, the Quaternary Deposits, and the Fluviatile Terrace Deposits (13). Holocene Formations found in the area of Ancistrocactus tobuschii are Low Terrace Deposits and Alluvium (13). Soil orders within the range of Ancistrocactus tobuschii are Life History - 2 (DRAFT) - Life History Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 primarily mollisols, with some vertisols and entisols (14,15,16,17). The soil series are; Anhalt, Denton, Dev, Ector, Frio, Knippa, Montell, Nuvalde, Olmos, Orif, Tarrant, and Uvalde (14,15,16,17). TROPHIC STATUS: Phototrophic (03,04,05,06). CHARACTERISTIC DOMINANCE: Component (09). COMMUNITY ECOLOGY: Ancistrocactus tobuschii occurs in the Ash Juniper-Live Oak Savannah. Most of this vegetation consists of woodland or scrub to dense to allow the build-up of extensive dense populations of Ancistrocactus tobuschii (09). Either Ancistrocactus tobuschii has always managed to survive under these conditions, implying that it has always gotten by with small scattered populations (half of which are in rocky flood-plains kept clear of trees only by catastrophic disturbance); or the dense scrub cover of the southern Edwards Plateau is a new phenomenon (09). SPECIES INTERRELATIONSHIPS: Ancistrocactus tobuschii is cross-pollinated by Dialictus (09). The cactus is utilized by larval Gerstaeckeria sp., a cactus- specialist weevil, which usually causes the death of the plant (09). Ancistrocactus tobuschii is also utilized by Chelinidea sp., a cactus- specialist bug whose feeding-punctures leave yellow scars on the plant but seem to cause no serious damage (09). OTHER LIFE HISTORY DESCRIPTORS: None. Life History - 3
                           (DRAFT) - Management Practices
                          Species CACTUS, FISHHOOK, TOBUSCH
                                Species Id ESIS702006
                                   Date 13 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Restricting/regulating human disturbance of populations Beneficial Maintaining/Controlling Water Flow Beneficial Maintaining undisturbed/undeveloped areas Beneficial Land Acquisition Beneficial Maintaining Sperm/Seed Banks Beneficial Stocking captive-reared wild-strain animals Beneficial Restricting Poaching Beneficial Controlling/Restricting Noncommercial Harvest Beneficial Regulating commercial harvest levels Beneficial Controlling/Removing Feral Animals Beneficial Controlling/Removing Domestic Animals Adverse Collecting Existing Collecting Adverse Commercial Exploitation Existing Commercial Exploitation Adverse Inherent Reproductive Characteristics Existing Inherent Reproductive Characteristics Adverse Low Gene Pool Existing Low Gene Pool Adverse Hybridization Existing Hybridization Adverse Parasites Existing Parasites Adverse Predation Existing Predation Adverse Rural Residential/Industrial Areas Existing Rural Residential/Industrial Areas Adverse Soil compaction by heavy equipment in mine areas Existing Soil compaction by heavy equipment in mine areas Adverse Flooding Existing Flooding Adverse Reservoirs Existing Reservoirs Adverse Dredging Existing Dredging Adverse Developing/maintaining stream bank vegetation Existing Developing/maintaining stream bank vegetation Adverse Applying herbicides Existing Applying herbicides Adverse Exotic/Feral/Introducted Species Existing Exotic/Feral/Introducted Species Adverse Erosion Existing Erosion Adverse Grazing Existing Grazing Adverse Vegetation Composition Changes Existing Vegetation Composition Changes Management Practices - 1 (DRAFT) - Management Practices Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 COMMENTS ON MANAGEMENT PRACTICES - Because many of Ancistrocactus tobuschii populations occur in areas prone to yearly flooding, these populations often undergo a great deal of stress (01,09). Such flooding also contributes to erosion along the stream and further modifying of the habitat of the cactus. Feral goats abound in central TX, and it is thought that their trampling of the cacti causes damage to the apical meristem (01). The greatest threat to the species is from goats (some are feral), sheep (domestic), and land development (i.e., housing) (09). Most of the otherwise suitable mesa-tops on the Edwards Plateau are nearly devoid of all species of dwarf cacti due to extreme alteration of the soil surface and a non-arborescent plant community. This situation is due to utilization of the plants by livestock, which eat the nurse plants (01) and mostly grasses and perennial herbs on which cactus seedlings depend for shelter, and livestock possibly eat Ancistrocactus tobuschii itself. Another threat is the destruction of habitat for land development (09). The best sites for housing are natural openings on bluffs and alluvial terraces, and these sites are the sites most often occupied by new houses (09). A large house, with the associated habitat disturbance, was placed within 100 meters or so of a dense Ancistrocactus tobuschii population, a short distance north of the City of Vanderpool, TX (09). Ancistrocactus tobuschii is cross-pollinated by Dialictus (09). The cactus is utilized by larval Gerstaeckeria sp., a cactus- specialist weevil, which usually causes the death of the plant (09). Ancistrocactus tobuschii is also utilized by Chelinidea sp., a cactus- specialist bug whose feeding-punctures leave yellow scars on the plant but seem to cause no serious damage (09). Ancistrocactus tobuschii is closely related to A. brevihamatus, which replaces tobuschii farther south and west; these two taxa intergrade in Kinney County, TX (09). The westernmost population of tobuschii slightly approaches A. brevihamatus in spine morphology, but has typical tobuschii flowers (09). The life expectancy of wild Ancistrocactus tobuschii plants, beyond attainment of sexual maturity, probably is less than ten years (09). This means that a population consisting largely of adult plants, with little evidence of recent reproduction, could completely disappear as a result of natural die-off in the few years following its discovery (09). This tendency for Ancistrocactus tobuschii populations to be ephemeral might render this taxon very difficult to manage on a single-population basis (09). At least near Vanderpool, Ancistrocactus tobuschii seems to simultaneously experience both castastrophic die-off and/or destruction of some populations, and very rapid growth of other populations (09). If Ancistrocactus tobuschii is indeed a fugitive species then very large tracts of habitat containing many emphemeral Ancistrocactus tobuschii populations would be necessary for the long-term survival of Ancistrocactus tobuschii at a given site (09). Private and commercial collectors are a very serious threat to any Ancistrocactus tobuschii population that becomes known to cactus collectors (09). The plant is a rare show specimen (01). Seeds were offered for sale in mid-1979 by the New Mexico Cactus Research Management Practices - 2 (DRAFT) - Management Practices Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 Foundation (01), alleviating the demand for field-collected plants, but collecting continues (09). Perhaps an equally important concern is the low gene pool. According to Weniger (01), less than 200 individuals are currently known; even if this is a great underestimate of the numbers, the fact remains that most Ancistrocactus tobuschii populations are small and widely separated (09). Other present threats to the plant include dams of any size (this means flooding because of a dam or reservoir), brush control by clearing or herbicides, and channelization or bank modification of any sort. UNAPPROVED PLAN: U.S. Fish and Wildlife Service. 1985. Technical Draft: Recovery Plan for the Tobusch Fishhook Cactus. U.S. Fish and Wildlife Service, Albuquerque, New Mexico. 30 pp. The prime objective of the Recovery Plan is to remove immediate human threats to presently known populations of Tobusch fishhook cactus and its essential habitat. The four known populations all occur on private land in widely separated areas of the Edwards Plateau and are threatened by real estate development and collecting. Removal of threats to Tobusch fishhook cactus populations are recommened through: 1) Enforcement of existing collecting and trade regulations under the Endangered Species Act, CITES, the Lacey Act and Texas State Law. 2) Establishment of safe sites at each known population site on private lands utilizing cooperative agreements, easements and/or purchases. 3) Development and implementation of habitat management plans for each safe site. 4) Efforts to ensure that the safe sites, once they are estabilished, are secure from possible impacts from the surrounding area (such as impacts from grazing and trampling by sheep and feral goats, development, etc.) by; a) maintaining the drainage systems at the four sites, b) controlling human access, and c) establishing buffer zones around each population for protection from foreseable threats. Ecological and population biology studies of Tobusch fishhook cactus are needed. Information required for better management include the determination of; 1) pollination vectors, 2) seed dormancy and longevity, 3) seed germination requirements and percentage of germination, 4) microhabitat factors involved in population establishment (i.e., periodic flooding), 5) the relationship between Tobusch fishhook cactus and associated flora, and 6) the role of animals in seed dispersal. The plan also calls for a long term survey program in Sabinal and Management Practices - 3 (DRAFT) - Management Practices Species CACTUS, FISHHOOK, TOBUSCH Species Id ESIS702006 Date 13 MAR 96 Frio Canyons, Texas, to locate new habitat sites and determine the number of mature plants and seedlings at each site. Other management needs addressed by the Recovery Plan include: 1) Development of a comprehensive trade management plan for all cacti. 2) Development of propagation techniques to provide nursery stocks and seeds to reduce collecting pressure and possibily developing a seed bank. 3) Development of public awareness, appreciation and support for the preservation of Tobusch fishhook cactus. Management Practices - 4
                                   (DRAFT) - References
                            Species CACTUS, FISHHOOK, TOBUSCH
                                  Species Id ESIS702006
                                      Date 13 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 Weniger, D. 1979. Status report on Ancistrocactus tobuschii. On file at: U.S. Fish and Wildlife Service, Albuquerque, NM. 02 Marshall, W.T. 1952. A new and interesting cactus from Texas. Saguaroland Bull. 6:78-81. 03 Benson, L. 1982. The cacti of the United States and Canada. Stanford Univ. Press, Stanford. 04 Weniger, D. 1970. Cacti of the southwest. Univ. of TX Press, Austin. 05 Benson, L. 1969. "Cactaceae" In: Flora of Texas, Vol. II. TX Res. Fond., Renner, TX. 06 Correll, D.S. and M.C. Johnston. 1970. Manual of the vascular plants of Texas. Texas Research Foundation. Renner, TX. 07 Backeberg, C. 1961. Die Cactaceae 5:2967. 08 Wauer, R. 1973. Naturalist's Big Bend. Peregrine Prodctions, Santa Fe, NM. 09 Zimmerman, A.D. 1985. Pers. comm. Department of Biology, University of Texas, Austin, TX 78712. 10 Sabo, K. 1978. Assignment: Ancistrocactus tobuschii. Cact. and Succ. J. (US) 50:214-215. 11 Hunt, D.R. 1978. More on Escobaria. Cact. Succ. J. (Great Britan) 40:30. 12 Zimmerman, A.D. 1981. The floral biology of Ancistrocactus tobuschii. Unpubl. manus. 13 Bureau of Economic Geology. 1965-1983. Geologic atlas of Texas. Univ. of TX, Austin. 14 Brand, R. 1985. Pers. comm. 15 Hensell, J.L., G.W. Dittmar, and F. Taylor. 1977. Soil survey of Bandera County, TX. USDA/SCS. 16 Stevens, J.W. and D.L. Richmond. 1970. Soil survey of Uvalde County, TX. USDA/SCS. 17 Blum, E.L. 1982. Soil survey of Kimble County, TX. USDA/SCS. ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 Zimmerman, A.D. 1985. Pers. comm. Department of Biology, University of Texas, Austin, TX 78712. 02 Weniger, D. 1979. Status report on Ancistrocactus tobuschii. On file at: U.S. Fish and Wildlife Service, Albuquerque, NM. 03 Brack, S. 1983. Field trip report on Endangered cacti to the U.S. Fish and Wildlife Service. On file at: U.S. Fish and Wildlife Service, Albuquerque, NM. References - 1