(DRAFT) - Taxonomy
                           Species GOOSEBERRY, MICCOSUKEE
                                Species Id ESIS704018
                                   Date 13 MAR 96



TAXONOMY

NAME - GOOSEBERRY, MICCOSUKEE OTHER COMMON NAMES - GOOSEBERRY, MICCOSUKEE; GOOSEBERRY, FLORIDA;GOOSEBERRY and SPINEY ELEMENT CODE - CATEGORY - Angiosperm PHYLUM AND SUBPHYLUM - MAGNOLIOPHYTA, CLASS AND SUBCLASS - MAGNOLIOPSIDA, ORDER AND SUBORDER - ROSALES, FAMILY AND SUBFAMILY - SAXIFRAGACEAE, GENUS AND SUBGENUS - RIBES, SPECIES AND SSP - ECHINELLUM, SCIENTIFIC NAME - RIBES ECHINELLUM AUTHORITY - TAXONOMY REFERENCES - COMMENTS ON TAXONOMY - Miccosukee Gooseberry Ribes echinellum (Coville) Rehder KINGDOM: Plant GROUP: Angiosperm DIVISION: Magnoliophyta CLASS: Magnoliopsida ORDER: Rosales FAMILY: Saxifragaceae Ribes echinellum (Miccosukee gooseberry) is a shrub 0.5 to 1.0 meter in height, often forming patches several meters in diameter, the branches spreading and recurved. The stems with one, two or three stout, dark reddish brown spines at the nodes, the spines reaching a length of 1.5 cm. The outer bark of 1-year-old branches is dull white to almost buff, splitting with growth of the twig and exposing the inner dark reddish-brown bark. Petioles pubescent, to 3 cm long; the leafblade 1 to 2 cm long, sometimes reaching 3 cm, orbicular in outline, truncate to rounded at the base, 3 lobed, sparingly pubescent or nearly glabrous on both surfaces. The peduncles 5 to 12 cm long, pubescent and usually with gland-tipped hairs, pendulous one-flowered or occasionally two-flowered. The pedicels 1 to 6 mm long, densely pubescent and glandular, subtended by 2 small bracts. The flower 15 to 20 mm in length, green to greenish-white. Ovary densely covered with long glands; calyx tube green, pubescent, 3 to 4 mm long, calyx Taxonomy - 1 (DRAFT) - Taxonomy Species GOOSEBERRY, MICCOSUKEE Species Id ESIS704018 Date 13 MAR 96 lobes 5, linear, pubescent, 4 to 6 mm long, reflexed; petals 2 to 3 mm long, inrolled and appearing tubular. The filaments 9 to 12 mm long, scattered pilose, exserting the reddish-pink pilose anthers out from the petals. The pistil has a shallowly divided style, 10 to 15 mm long. The fruit is a green berry, 2 to 3 cm in diameter (including spines), densely covered with gland-tipped spines up to 8 mm long (01, 02). The species was first named Grossularia echinella by Coville in 1924 (02), this was later revised to the present nomenclature in 1926 by Rehder (03). In 1985, Sinnott (02) noted that the species should not be included with the hairy-styled type species in section Grossularia. The lack of style pubescence, tubular petals, and greenish calyx are features found in species of the western group related to Ribes lobbii. There have been some morphological differences observed between Florida and South Carolina plants grown in cultivation at Woodlanders (a native plant dealer). Apparently, the spines of Florida plants occur primarily at the nodes while specimens of South Carolina origin have spines along the internodes as well. In addition, South Carolina plants are flowering slightly earlier and usually have smaller fruit (04). Common names used for this species include Miccosukee gooseberry, Florida gooseberry, and spiney gooseberry. The type specimen (No. 1, 112, 807), collected by Coville on March 2, 1924 (01), was deposited in the United States National Herbarium. Technical descriptions and drawings useful for species identification are available by reference to the published papers of Coville (1924) (01) and Sinnott (1985) (02). Living specimens from both the Florida and South Carolina populations may be simultaneously viewed by appointment at Woodlanders (04). Taxonomy - 2
                                  (DRAFT) - Status
                           Species GOOSEBERRY, MICCOSUKEE
                                Species Id ESIS704018
                                   Date 13 MAR 96



STATUS

Coded Status Florida; Federal Threatened Florida; Officially Listed Florida; Unofficially Listed South Carolina; State Recognized South Carolina; Unofficially Listed T: Federal Threatened Commercial Game (Consumptive Recreational) Ornamental COMMENTS ON STATUS - U.S. STATUSES AND LAWS: The Miccosukee gooseberry (Ribes echinellum) has been designated a Threatened species pursuant to the Endangered Species Act of 1973 (50 CFR 17.12; P.L. 93-205, 87 Stat. 884; 16 U.S.C. 1531-1540), as amended. The species has this status wherever found including the States of FL and SC. Critical Habitat has not been designated. Removal and reduction to possession of any Federally listed plant from an area under Federal jurisdiction is unlawful (50 CFR 17.61 and 17.71). This species is protected by the Lacey Act (P.L. 97-79, as amended; 16 U.S.C. 3371 et seq.) which makes it unlawful to possess any wild plant (including roots, seeds, and other parts) within U.S. territorial or special maritime jurisdiction (as defined in 18 U.S.C. 7); or to import, export, transport, sell, receive, acquire, or purchase in interstate or foreign commerce any wild plant (including roots, seeds, and other parts) taken, possessed, transported, or sold in violation of any State law or regulation. It is also unlawful to import, export, transport, sell, receive, acquire, or purchase any wild plant (including roots, seeds, and other parts) taken or possessed in violation of any U.S. law, treaty, or regulation or in violation of Indian tribal law. RESPONSIBLE FEDERAL AGENCIES: USFWS -Responsible for the management/recovery, listing, and law enforcement/protection of this species. USFS -Responsible for the law enforcement/protection of this species with applicable State and Federal laws on public lands under their control. Also responsible for management/recovery on Forest Service lands. Forest Service Regulations prohibit the removal, destruction, or damaging of any plant classified as threatened, endangered or rare (36 CFR 261). All Federal agencies have responsibility to ensure that any action authorized, funded, or carried out by that agency is not likely Status - 1 (DRAFT) - Status Species GOOSEBERRY, MICCOSUKEE Species Id ESIS704018 Date 13 MAR 96 to jeopardize the continued existence of the species and to utilize their authorities to carry out programs for the conservation of the species. STATE STATUSES AND LAWS: STATE: Florida DESIGNATED STATUS: Endangered ADMINISTRATIVE AGENCY: Florida Department of Agriculture and Consumer Services STATE STATUTE: Florida State Law, Chapter 65-426, Status Section 581.185-187 ("Preservation of Native Flora of Florida Act"). UNOFFICIAL LIST: Endangered; In: Ward, D.B., ed. 1979. Rare and endangered biota of Florida. Vol. 5: Plants. Univ. Presses of Florida. 175 pp. STATE: South Carolina DESIGNATED STATUS: Recognized Threatened ADMINISTRATIVE AGENCY: SC Wildlife and Marine Resources Dept. STATUTE: SC Limited Authorities Cooperative Agreement; Append. A UNOFFICIAL LIST: Endangered; Unofficial list: "Native Vascular Plants Endangered, Threatened, or Otherwise in Jeopardy in South Carolina" INTERNATIONAL STATUSES, TREATIES, AND AGREEMENTS: None. ECONOMIC STATUSES: Gooseberries and currants are cultivated for their edible fruits and for their ornamental habit and bloom. Ribes echinellum (Miccosukee gooseberry) is not in demand for these purposes at present (05). It is cultivated commercially by at least one nursery. 75/07/01:40 FR 27823/ - Smithsonian petition 76/06/16:41 FR 24523/ - Proposal to list 1700 plants 78/04/26:43 FR 17909/ - Comment summary for 76/06/16 publication 79/12/10:44 FR 70796/ - Withdrawal notice 80/12/15:45 FR 82480/ - Review, list as Endangered or Threatened 84/01/20:49 FR 2485/ - 1-year petition finding 84/08/31:49 FR 34535/ - Proposal to list as threatened 84/10/22:49 FR 41266/ - Notice of Public Hearing 85/07/18:50 FR 29338/29341 - Final rule listing species as Threatened Status - 2
     

HABITAT ASSOCIATIONS

HABITAT - TERRESTRIAL TERRESTRIAL LAND USE - Deciduous Forest Land COMMENTS ON HABITAT ASSOCIATIONS - Ribes echinellum (Miccosukee gooseberry) is associated with a deciduous, mixed hardwood forest with an overstory canopy dominated by species of oak and hickory (Quercus and Carya). The species was originally discovered in March of 1924 along the northeast shore of Lake Miccosukee, Jefferson County, Florida near Judge Hopkins Landing (01,02). Four additional segments of the Florida population totaling 5 to 10 acres (in Section 7, Range 4E, Township 2N) are described by Schultz and Hardin (03) one-half to one mile east of the original location. In South Carolina, the species was discovered in 1957 (04) along the western and southern side of Stevens Creek. The population is 1.5 miles northeast of Clark's Hill, McCormick County, South Carolina within the boundaries of the Stevens Creek Natural Area (05). Ribes echinellum also occurs outside the boundaries of the Stevens Creek Natural Area. This disjunct segment was located by Jones in 1981 (06). This colony occurs on Sumter National Forest, Edgefield Ranger District. Harper (10) described the site of the type location in Florida as wooded bluff slopes with an unusual absence of evergreen species. It was estimated that only one-tenth of the vegetation is evergreen. Harper listed Quercus schneckii (Quercus shumardii), Q. michauxii, Liquidamber styraciflua, Tilia sp., Fraxinus americana, Acer floridanum (Acer barbatum), Ulmus fulva (Ulmus rubra), and U. alata as the most abundant tree species. Within the small tree size-class, Prunus caroliniana, Cercis canadensis, and Ilex opaca were listed as the dominants. Bignonia crucigera, Parthenocissus quinquefolia, and Smilax rotundifolia were the most abundant woody vines. Grossularia echinella (=Ribes echinella) was the most abundant shrub associated with Adelia ligustrina and Aesculus pavia. The most abundant herbs were Tovara virginiana, Calycocarpum lyoni, Tillandsia usneoides, Arisaema dracontium, Spigelia marilandica, and Tradescantia sp. Schultz and Hardin (11) surveyed essentially the same area in 1985. According to their estimates, the dominant tree canopy was composed of Carya glabra, Fraxinus americana, Ulmus alata, Celtis laevigata, and Quercus virginiana. Common subdominant trees included Acer barbatum, Prunus serotina, P. caroliniana, Liquidamber styraciflua, Cercis canadensis, Magnolia grandiflora, Quercus nigra, Q. michauxii, and Tilia americana. Ribes echinellum, Aesculus pavia, Aralia spinosa, and Rhus radicans were abundant in the shrub layer. The predominant herb species were Trillium underwoodii, Polygonatum biflorum, and Arisaema dracontium. On the eastern side of the peninsula at Lake Miccosukee the dominant tree canopy included Acer barbatum, Carya glabra, Magnolia grandiflora, and Quercus michauxii. In addition to the species listed as present on the west side on the peninsula, Carpus carolinianus was a common sub-dominant tree and Arundinaria gigantea was abundant in the understory. The estimated number of individuals of Ribes echinellum was 5000 over 100 acres. The largest number of individuals were found to be 200 to 300 feet south of the boat landing on the western side of the peninsula at Lake Miccosukee. The site was characterized as a westerly aspect, with the topographic position ranging from mid-slope to bottom-slope and slope ranging from 0-10 percent to 10-35 percent. Light Habitat Associations - 1 conditions were filtered to shady with a mesic moisture status. Elevation ranged from 80 to 120 feet. Schultz and Hardin (11) also surveyed the Florida population-segment in Section 7, Range 4E, Township 2N. The dominant trees were Fagus grandiflora, Magnolia grandiflora, and Acer barbatum while other associated tree species included Quercus michauxii, Q. nigra, Q. virginiana, Carya glabra, and Fraxinus americana. The shrub layer was composed of Ribes echinellum, Aralia spinosa, Aesculus pavia, Euonymus americanus, and Rhus radicans. Arisaema dracontium, Polygonatum biflorum and Zephyranthes atamasco were dominant herbs. The estimated number of individuals of Ribes echinellum was 3000 to 5000 over five to ten acres. The site was characterized as flat relief approximately five feet above the level of the surrounding swamp. The soil moisture status was defined as mesic and apparently well drained at an elevation of 80 to 90 feet. In South Carolina the forest community was originally described as mixed mesophytic by Radford in 1959 (12). The composition of the tree canopy was approximated as 25 percent Carya cordiformis, 15 percent Acer barbatum, 12 percent Quercus prinus, (probably Q. michauxii misidentified), 12 percent Q. rubra, 10 percent Ulmus rubra, 8 percent Celtis laevigata, 8 percent Carya ovata, 3 percent Juglans nigra, 2 percent Liriodendron tulipifera, 2 percent Liquidamber styraciflua, 2 percent Ulmus alata, and 2 percent Quercus alba. Within the subcanopy were Ostrya virginiana, Acer barbatum, Quercus prinus, Q. alba, and Carya cordiformis. The more abundant shrubs were Lindera benzoin, Asimina triloba, Philadelphus inodorus, Staphylea trifolia, Aesculus sylvatica, and Ribes echinellum. The more common herbs included Isopyrum, Claytonia, Trillium, Tiarella, Dentaria, Geranium, Senecio, Dodecatheon, Desmodium, Agrimonia, Phyrma, Helianthus, and Campanula. Jones (08,13,14) established six 1/10 acre circular plots on the primary bluff where the Ribes echinellum population is best developed. Importance value 200 [(Rel. Den. + Rel. BA.) div. by 2] was calculated by species for the tree and sapling class, relative density for the seedling size class, and relative frequency for herbs and vines. The major tree species and their importance values are as follows: Ulmus rubra 25, Fraxinus americana 11, Quercus alba 10, Carya glabra 10, Acer barbatum 8, Carya cordiformis 8, Quercus michauxii 7. The tree canopy was composed of species of Carya and Quercus in association with occassional individuals of Fraxinus americana. The mid-canopy or intermediate tree size-class was composed of Ulmus rubra and Acer barbatum in association with scattered individuals of the Carya, Quercus, and Fraxinus species mentioned above. Basal area levels of the tree size-class ranged from 10 sq m/ha to 32 sq m/ha. The sapling and seedling size-classes were dominated by Acer barbatum 45, 29, Ostrya virginiana 31,25, Ulmus rubra 12,18, respectively. Ribes echinellum was by far the dominant shrub with a mean density of 54,000 stems/ha and a range of 13,000 stems/ha to 161,000 stems/ha. Associated shrub and woody vines were Philadelphus inodorus, Euonymus americanus, Vitis sp., and Parthenocissus quinquefolia. The herb layer was unusually rich for a Piedmont upland forest with 62 herb species identified on 72 1.0 sq m quadrats. The predominate spring herb species by relative frequency were Senecio obovatus 76, Geranium maculatum 75, Osmorhiza longistylis 50, Erythronium americanum 47, Hexastylis arifolia 47, Stellaria media 47, Claytonia virginica 47, Cardamine angustata 46, and Isopyrum biternatum 43. The predominate fall species by relative frequency were Bromus purgans 50, Bignonia capreolata 39, and Solidago caesia 36. Results of the area sampled do not indicate a dominant tree layer with the composition of mixed-mesophytic forest as defined by Braun (15). Species of Quercus and Carya had Importance Values of 24 and Habitat Associations - 2 33, respectively, accounting for over 50 percent of the composition of the tree size-class. Therefore, the community would be best defined as a mesic variant of the oak-hickory type. The site where the vegetative survey was conducted is a lower slope with a northwest to northeast aspect and 28 deg. to 40 deg. slope. Elevation ranges from 220 to 380 feet. The soil pH of the A-horizon ranged from 6.7 to 7.4 and the texture was a sandy loam. In the A-horizon, the level of exchangable calcium was 1000+ ppm, magnesium 51+ ppm, potassium 50 to 105 ppm, and phosphorus 6 to 25 ppm. The soil less than 20 feet up-slope from the area vegetatively sampled is a Wateree series which is a coarse-loamy, mixed, thermic, Typic Dystrochrepts. The pH ranged from 4.6 to 5.1 and the base levels were lower than found in the sample area. Ribes echinellum was present here only as scattered individuals (08). Rock outcrops of cataclastic granatoid gneiss with potassium feldspars and plagioclase (16) are common along the bluffsite. According to Griffin (16), the high pH and calcium levels are likely due to the presence of marble. This is in agreement with Smith (17) and Hodges (18) who are of the opinion the base levels are too high to be attributed to the presence of high base tolerant plant species. The disjunct population segment in South Carolina located by Jones in 1981 is in a forest community dominated by Liquidamber styraciflua and Ostrya virginiana. The stand was heavily disturbed by logging activity approximately 25 to 30 years ago. The understory is open and sparse. The site is characterized as a lower slope position, 10 to 20 percent slope with an easterly aspect. The soil is a Tatum which is derived from fine-grained phyllite and is a clayey, mixed, thermic, Typic Hapludults (08). Habitat Associations - 3
                                (DRAFT) - Food Habits
                           Species GOOSEBERRY, MICCOSUKEE
                                Species Id ESIS704018
                                   Date 13 MAR 96



FOOD HABITS

TROPHIC LEVEL - AUTOTROPH Food Habits - 1
                         (DRAFT) - Environment Associations
                           Species GOOSEBERRY, MICCOSUKEE
                                Species Id ESIS704018
                                   Date 13 MAR 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Terrestrial Features: Rock outcrops G G Environment Associations - 1
                               (DRAFT) - Life History
                           Species GOOSEBERRY, MICCOSUKEE
                                Species Id ESIS704018
                                   Date 13 MAR 96



LIFE HISTORY

HABIT: Ribes echinellum (Miccosukee gooseberry) is a woody shrub (01,02 03). LIFE CYCLE: Perennial (01,02,03). TYPE OF REPRODUCTION: Sexual and vegetative (04,08). Reproduction from seed has been observed, with the best germination condition probably being bare mineral soil (04,19). Vegetative reproduction occurs commonly by rooting at the stem tips whenever they touch the ground (19). The species is also easily rooted from cuttings. REPRODUCTIVE PHENOLOGY: Germination occurs in late spring, leafing in October, and budding in February. Anthesis dates are from March to April, fruiting dates from April to July, and fruit dispersal is from June to August (08,10). Flowering and fruiting occurs approximately two weeks earlier in Florida than in South Carolina (10,11,19,). SEX OR SPORE STATUS: Monoclinous (01). POLLINATION, SPORE AND SEED DISSEMINATION: Seed dissemination agents are birds. Possible disseminators are mockingbirds, catbirds or thrushes (01). Coville (01) postulated that birds with long bills ( mockingbirds, catbirds or thrushes) would be adapted to open the berries to eat the seed. These species of birds are common and would have a low vulnerability to disturbance, habitat modification, or predation. The daily range of these species would be limited, therefore the potential dispersal distance would be limited. Dissemination by these agents would be internal. SEED BIOLOGY: The production of seed does not appear to be a problem. Seed germination tests have been conducted with many different scarification and stratification treatments with no success. Limited success was achieved by leaving the seed in the fruit and allowing the fruit to rot on a germination medium under greenhouse conditions. When fruit was partially buried in bare mineral soil in July at the South Carolina site, some seed germination was observed the following spring (08). POPULATION BIOLOGY: The major limiting factor may be competition from native and non-native (Lonicera japonica is a potential problem on the South Carolina site) plants. Other factors may include soil water status (soil moisture) and poorly drained soils (19). In Florida, the estimated number of individuals was 5,000 over an area of 100 acres in the Section 12, Range 3E, Township 2N population segment and 3,000 to Life History - 1 (DRAFT) - Life History Species GOOSEBERRY, MICCOSUKEE Species Id ESIS704018 Date 13 MAR 96 5,000 over five to ten acres in the Section 7, Range 4E, Township 2N population segment (11). In South Carolina, the density ranged from 13,000 to 161,000 stems/ha within the area where the species is best developed. Approximately 20 to 25 percent of the plants were less than one-foot tall (08,13). The populations are stable and maintaining themselves (19,20). ECOLOGICAL/EDAPHIC FACTORS: Detailed information concerning the soil texture and moisture (soil moisture status) requirements is not known. The occurrence of the species is so local that Soil Conservation Service soil surveys are not useful. The species appears to require loamy, well drained soils with a mesic moisture status. Wateree and Tatum are soil series that the species occurs on in South Carolina (08), but at the periphery of the population. The series within the area where the species is best developed is unknown. In Florida, the soil series that the species occurs on is also unknown. In South Carolina, the rock is cataclastic granatoid gneiss with potassium feldspars and plagioclase, and it is suspected that marble is present (16). In Florida, the rock is limestone (10). Light conditions are filtered to shady. Slopes range from 0 to 35 percent in Florida and 30 to 45 percent in South Carolina. The microclimate is cool-mesic. Elevation ranges from 80 to 120 feet in Florida and 220 to 380 feet in South Carolina. Aspects are northerly. In South Carolina, soils are neutral to slightly basic (pH 6.7 to 7.4) but peripheral individuals are on acid soils (pH 4.6 to 5.1). Disturbance is minimal (08,11). TROPHIC STATUS: Phototrophic (01). CHARACTERISTIC DOMINANCE: Subdominant (01,08,11). The species often forms dense patches eliminating all other understory plants. COMMUNITY ECOLOGY: The associated plant community is a mesic-variant of the oak-hickory forest. The canopy is dominated by species of oak and hickory (Quercus and Carya) in association with Fraxinus americana. A dense sub-canopy exists due to a large number of individuals of Acer barbatum, Ulmus rubra, and U. alata. In the Section 7, Range 4E, Township 2N population segment in Florida, Fagus grandifolia and Magnolia grandiflora were also important components of the overstory (08,10,11,12). The communities are steady-state (climax) to near steady-state with single tree canopy gaps. It is not known what effect disturbance has on the population biology of this species. SPECIES INTERRELATIONSHIPS: No information available. OTHER LIFE HISTORY DESCRIPTORS: No information available. Life History - 2
                           (DRAFT) - Management Practices
                           Species GOOSEBERRY, MICCOSUKEE
                                Species Id ESIS704018
                                   Date 13 MAR 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Suppressing wildfire Beneficial Controlling/Restricting Off-Road Vehicles Beneficial Restricting/regulating human disturbance of populations Beneficial Maintaining undisturbed/undeveloped areas Beneficial Land Acquisition Beneficial Controlling/Restricting Herbicide Use Beneficial Restricting Timber Harvest Beneficial Controlling/Removing Nonnative Vegetation Beneficial Maintaining Sperm/Seed Banks Beneficial Stocking captive-reared wild-strain animals Beneficial Transplanting wild animals Beneficial Transplanting Wild Eggs/Wild Seeds Beneficial Controlling/Restricting Noncommercial Harvest Beneficial Regulating commercial harvest levels Adverse Exotic/Feral/Introducted Species Existing Exotic/Feral/Introducted Species Adverse Collecting Existing Collecting Adverse Soil compaction by heavy equipment in mine areas Existing Soil compaction by heavy equipment in mine areas Adverse Shoreline modification/development Existing Shoreline modification/development Adverse Low Gene Pool Existing Low Gene Pool Adverse Competition Existing Competition Adverse Fire Existing Fire Adverse Forest Alteration Existing Forest Alteration Adverse Harvesting Existing Harvesting COMMENTS ON MANAGEMENT PRACTICES - The threat of habitat destruction or alteration which may be detrimental to the continued existence of Ribes echinellum (Miccosukee gooseberry) is greatest at the Florida site since the population is on private owned lands and the site has potential for lakeside development (06). Logging of the associated hardwoods and severe fire could pose additional threats to this population. Logging occurred near part of the site with observed detrimental effects (05). Although the South Carolina population occurs on land managed as a nature preserve by the South Carolina Wildlife and Marine Resources Department and on U.S. Forest Service land (Sumter National Forest) that will be set aside and given special consideration (06), little is known about management needs of the species. Although the South Carolina Forestry Commission maintains firebreaks at the Stevens Creek Natural Area, incendiary fires are always a threat. One such incident Management Practices - 1 (DRAFT) - Management Practices Species GOOSEBERRY, MICCOSUKEE Species Id ESIS704018 Date 13 MAR 96 occurred in 1978 (07) which burned one clump of the species. These factors are still considered threatening to the population. The inadequacy of existing regulatory mechanisms was one reason for the current status of the species. Cuttings and fruit have been taken from these plants for cultivation. The Florida Endangered Species Act offers additional protection under Florida State Law, Chapter 65-426, which includes prohibitions concerning taking, transport, and the sale of plants listed under the Florida law. In South Carolina, the species is indirectly protected at the Stevens Creek site under the Natural Area prohibitions against unauthorized plant taking (05). The colony on Sumter National Forest in South Carolina is protected by the Endangered Species Act and Forest Service regulations which prohibit taking of protected plants on Federal lands. This factor is no longer threatening to the population. Other natural or manmade factors affecting the continued existence of the species include the small size and number of both the South Carolina and Florida populations (many of the plants may have reproduced vegetatively not sexually). This factor results in both populations being sensitive to even small-scale perturbations. Lightning and incendiary fires are probably the major threat at this time. Both populations are receiving localized competition from the introduced vine, Japanese honeysuckle (Lonicera japonica) (08,09). This factor continues to be threatening. Agricultural pressure around the Florida sites may lead to future development of these sites for agricultural purposes. Another future threat is the overutilization of this species for commercial purposes. Gooseberries and currants are cultivated for their edible fruits and for their ornamental habit and bloom. The species is not in demand for these purposes at present (05). UNAPPROVED PLAN: A Recovery Plan has not yet been initiated; target date for a draft plan is FY 1987. Ribes echinellum (Miccosukee gooseberry), confined to two populations in South Carolina and Florida, is potentially threatened at the Florida site by development, unrestricted timber harvest, herbicide use, agricultural development, and off-road vehicle use. Although the current landowners of the Florida site appear cooperative, a change in ownership or land use could be detrimental to the plant. The South Carolina site is owned primarily (approximately 95 percent of the colonies) by the South Carolina Wildlife and Marine Resources Department and managed by the state's Heritage Trust Program for the benefit of this species. Uncontrolled exotic plants, particularly Japanese honeysuckle (Lonicera japonica), and wildfire are among the few threats to the integrity of this site. More specific information is needed with regard to the effects of this non-native plant invader, and on the specific environmental conditions which may foster expansion of existing gooseberry populations. Some State restrictions should be applied if collecting of wild material becomes excessive. Acquisition of adjacent colonies which are now unprotected should be pursued. Similarly, restrictions of Management Practices - 2 (DRAFT) - Management Practices Species GOOSEBERRY, MICCOSUKEE Species Id ESIS704018 Date 13 MAR 96 recreational use of sites may be desirable/necessary at some point to reduce the potential for collecting and accidental destructive fires. Other actions recommended for the recovery of this species would be to establish a seed bank with seeds from each population, propagate plants for reintroduction to suitable habitats, or reintroduce wild plants or seeds if feasible. Adjacent to the South Carolina population are commercial timber lands which are periodically sprayed to control hardwoods during reseeding operations. This spraying may or may not have an adverse effect on Ribes echinellum but may need to be monitored or controlled. A very small colony of this species occurs on the Sumter National Forest (where "take" is Federally prohibited). The U.S. Forest Service has agreed to manage the site for the species' benefit. The State of South Carolina is pursuing acquisition of the remaining privately-owned colonies in that state. Florida landowners have been contacted and expressed willingness to cooperate in conservation efforts for the species. Management Practices - 3
                                   (DRAFT) - References
                              Species GOOSEBERRY, MICCOSUKEE
                                  Species Id ESIS704018
                                      Date 13 MAR 96



     

References

***** REFERENCES FOR ALL NARRATIVES EXCEPT N-OCCURRENCE ***** 01 Coville, F.V. 1924. Grossularia echinella, a spiny-fruited gooseberry from Florida. J. Agri. Res. 28:71-76. 02 Sinnott, Q.P. 1985. A revision of Ribes L. Subg. Grossularia (Mill.) Pers. Sect. Grossularia (Mill.) Nutt. (Grossulariaceae) in North America. Rhodora 87:189-281. 03 Rehder, A. 1926. New species, varieties, and combinations from the herbarium and collections of the Arnold Arboretum. Journal of the Arnold Arboretum 7:148. 04 McCartney, R. July 1, 1986. Pers. comm. (telephone). Woodlanders, 1128 Colleton Ave., Aiken, SC 29801. 05 U.S. Fish and Wildlife Service. July 18, 1985. Endangered and threatened wildlife and plants; determination of threatened status for Ribes echinellum (Miccosukee gooseberry). Fed. Reg. 50(138):29338-29341. 06 Gardner, J. and A. Boldog. June 20, 1986. Pers. comm. U.S. For. Service, Sumter National Forest, P.O. Box 30, Edgefield, SC 29824. 07 Barr, W.L. February 28, 1978. [Letter to T. S. Kohlsaat]. South Carolina State Comm. of Forestry, P.O. Box 21707, Columbia, SC. 08 Jones, S.M. 1986. Unpubl. data. Dept. Forestry, Clemson Univ., Clemson, SC. 09 Gholson, A. July 1, 1986. Pers. comm. (telephone). Catahootchie, Florida. 10 Harper, R.M. 1925. A botanically remarkable locality in the Tallahassee red hills of middle Florida. Torreya 25:45-54. 11 Schultz, G.E. and E.D. Hardin. 1985. Site survey summary. Florida Natural Areas Inventory. The Nature Conserv. and the FL Dept. of Nat. Resour. 13 pp. 12 Radford, A.E. 1959. A relict plant community in South Carolina. J. Elisha Mitchell Sci. Soc. 75:33-34. 13 Jones, S.M. and B.A. Dunn. 1978. Compositional and soil-site characteristics of a Ribes echinellum (Saxifragaceae) community. ABS Bulletin 25(2):9. 14 Jones, S.M. and B.A. Dunn. 1983. Endangered and threatened forest plants of South Carolina. Dept. of Forestry, Bulletin No. 37. Clemson Univ., Clemson, SC. 117 pp. 15 Braun, E.L. 1950. Deciduous forests of eastern North America. Hafner Press, N.Y. 596 pp. 16 Griffin, V.S. July 3, 1986. Pers. comm. Dept. of Chemistry and Geology, Clemson Univ., Clemson, SC 29631. 17 Smith, B.R. July 3, 1986. Pers. comm. Dept. of Agronomy and Soils, Clemson Univ., Clemson, SC 29631. 18 Hodges, S.C. July 3, 1986. Pers. comm. Dept. of Forestry, Clemson Univ., Clemson SC 29631. 19 Jones, S.M. 1986. Pers. observations, field work dating from 1977 to 1981. Dept. of Forestry, Clemson Univ., Clemson, SC 29631. 20 Milstead, W.L. 1978. Status report on Ribes echinellum. U.S. Fish and Wildlife Service, Atlanta, Georgia. 19 pp. References - 1 (DRAFT) - References Species GOOSEBERRY, MICCOSUKEE Species Id ESIS704018 Date 13 MAR 96 ***** REFERENCES FOR N-OCCURRENCE NARRATIVE ONLY ***** 01 Milstead, W.L. 1978. Status report on Ribes echinellum. U.S. Fish and Wildlife Service, Atlanta, GA. 19 pp. 02 Coville, F.V. 1924. Grossularia echinella, a spiny-fruited gooseberry from Florida. J. of Agri. Res. 28:71-76. 03 Schultz, G.E. and E.D. Hardin. 1985. Site survey summary. FL Nat. Areas Invent. The Nature Conserv. & FL Dept. of Nat. Resour. 13 pp. 04 Radford, A.E. 1959. A relict plant community in South Carolina. J. Elisha. Mitchell Sci. Soc. 75:33-34. 05 Jones, S.M. 1986. Personal observations. Department of Forestry, Clemson University, Clemson, SC. 06 Jones, S.M. 1986. Unpubl. data. Dept. of For., Clemson Univ., Clemson, SC. 07 Rayner, D.A. 1986. Map to S. M. Jones. South Carolina Wildlife and Marine Resources, P.O. Box 167, Columbia, SC 29202. 08 Sinnott, Q.P. 1985. A revision of Ribes L. Subg. Grossularia (Mill.) Pers. Sect. Grossularia (Mill.) Nutt. (Grossulariaceae) in North America. Rhodora 87:189-281. 09 Sinnott, Q.P. July 1, 1986. Pers. comm. (telephone). Database Mgmt. Unit, Germplasm Resour. Info. Network, Plant Genetics and Germplasm Inst., Bldg. 001, BARC-West, ARS Beltsville, MD 20705. References - 2