(DRAFT) - Taxonomy
                             Species flounder, southern
                                 Species Id M010021
                                   Date 27 AUG 96



TAXONOMY

NAME - flounder, southern OTHER COMMON NAMES - flounder, mud flounder, doormat and halibut ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Osteichthyes, ORDER AND SUBORDER - Pleuronectiformes, FAMILY AND SUBFAMILY - Bothidae, GENUS AND SUBGENUS - Paralichthys, SPECIES AND SSP - lethostigma, SCIENTIFIC NAME - Paralichthys lethostigma AUTHORITY - Jordan and Gilbert TAXONOMY REFERENCES - 77 COMMENTS ON TAXONOMY - Other common names are Flounder, mud flounder, doormat, and halibut.*77* Taxonomy - 1
                                  (DRAFT) - Status
                             Species flounder, southern
                                 Species Id M010021
                                   Date 27 AUG 96



STATUS

Coded Status Sport Fish Commercial Commercial/consumption REFERENCES FOR STATUS - 77 COMMENTS ON STATUS - The southern flounder is a valuable sport and commercial fish along the gulf coast. Sport fishing is done by hook and line and by gigging. Most of the commercial catch is incidental to the catch by shrimp trawlers. The southern flounder is "one of the most sought after and prized fish in the area and is recognized for its fine flavor."*77* Status - 1
                               (DRAFT) - Distribution
                             Species flounder, southern
                                 Species Id M010021
                                   Date 27 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Estuarine Habitat Associations - 1
                                (DRAFT) - Food Habits
                             Species flounder, southern
                                 Species Id M010021
                                   Date 27 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 77 LIFESTAGE FOOD FOOD PART Juvenile Invertebrate Cordates See Comments Adult Osteichthyes See Comments REFERENCES FOR ADULT FOOD - 77 REFERENCES FOR JUVENILE FOOD - 77 COMMENTS ON FOOD - Small southern flounders eat a variety of invertebrates, but become piscivorous when they are about 200 mm long (TL).*77* COMMENTS ON ADULT FOOD - In Louisiana, adult southern flounders eat shrimp and fish. In a more detailed study in Louisiana, Fox and White (1969) reported that striped mullet (Mugil cephalus) was the major food item of southern flounders, followed by fat sleepers (Dormitator maculatus) and anchovies (Anchoa). Larger flounders (150 mm long) ate primarily fish, among which anchovies, menhaden (Brevoortia sp.), sciaenids, and mullet (Mugil sp.) were most common.*77* COMMENTS ON JUVENILE FOOD - In Texas, Stokes (1977) reported that small flounders (10 to 150mm long) ate mostly invertebrates (95%) among which mysids were the most common (32%).*77* Food Habits - 1
                         (DRAFT) - Environment Associations
                             Species flounder, southern
                                 Species Id M010021
                                   Date 27 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS A Water Temperature: Specified in Comments J Water Temperature: Between 15-21 degrees C A J LIM Dissolved Oxygen: Low [less than 5 mg/l] oxygen concentrations REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 77 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 77 REFERENCES FOR FEEDING JUVENILE ENVIRONMENTAL ASSOC_ - 77 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Temperature influences the migration of postlarval and adult southern flounders.*77* COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - In Louisiana coastal waters, adult southern flounders have been collected at temperatures ranging from 5 to 35 degrees C.*77* COMMENTS ON FEEDING JUVENILE ENVIRONMENTAL ASSOC_ - Postlarval migration to estuaries from offshore waters peak when water temperatures are about 16 degrees (Stokes 1977).*77* In a laboratory study, postlarval southern flounders attempted avoidance when dissolved oxygen concentrations fell below 3.7 mg/l. No avoidance differences were noted at temperatures of 6.1 degrees, 14.4 degrees, or 25.3 degrees C.*77* Environment Associations - 1
                               (DRAFT) - Life History
                             Species flounder, southern
                                 Species Id M010021
                                   Date 27 AUG 96



LIFE HISTORY

Morphology and Identification Aids: Dorsal fin rays range from 80 to 95; anal rays from 63 to 74, and pecotoral fin (eye side) rays from 11 to 13. Upper gill rakers on the upper limb of the first fill arch range from 2 to 3 and lower gill rakers on the lower limb from 8 to 11. Scales in the lateral line range from 85 to 100. Body depth is 30% to 47% of standard length (SL). The eyes are on the left side and color is light to dark brown with diffuse nonocellated dark spots and blotches. The blindside is white or dusky.*77* Spawning: Most southern flounders spawn in late fall and early winter, but some spawn in in in early spring. In North Carolina, southern flounders migrate out of estuaries in fall to spawn. In Texas, they migrate from estuaries into the Gulf of Mexico from October through December, apparently to spawn. Males move seaward earlier than females and few remain in the estuaries after November. This migration is usually preceded by a drop in water temperature of 4 to 5 degrees C. Southern flounders are caught in gulf waters as deep as 63 m. Females become sexually mature at 2 years of age in Texas. The youngest mature female southern flounder in northern Florida was 4 years old. Of the mature females collected in August, 8% of the 4-year-olds, 5% of the 5-year-olds, and 18% of the 6-year-olds were developing eggs. Southern flounders in Texas were induced to spawn in the laboratory. About three weeks before spawning took place, males began following gravid females in the tanks. The first spawning was on December 21. Spawning was at midday, when females swam to the surface and released eggs that were immediately fertilized by attending males. Fertilization was 30% to 50% successful, and 6% to 35% of the eggs hatched in 61 to 76 hr.*77* Fecundity: Thirteen southern flounders examined in the laboratory, produced a total of 120,000 eggs (average about 9,230).*77* Larvae: In culture, yolk-sac larvae began metamorphosing to postlarvae at 40 to 46 days (8 to 11 mm long): metamorphosis was complete by 50 to 51 days.*77* Juveniles and Adults: Postlarvae of southern flounder 18 to 34 mm in total length (TL) were captured during February, March, and May at Galveston Island, Texas; fish 25 to 51 mm TL were caught in Mississippi River passes during spring. Southern flounder postlarvae are caught along the Gulf of Mexico coast during winter and early spring. In Aransas Bay, Texas, the peak movement of postlarvae flounders into estuaries is in February, when water temperatures are between 16.0 and 16.2 degrees C. In Texas, Breuer (1962) found postlarvae 35 to 50 mm TL in December. Juveniles are generally collected during spring, summer, and early fall. Juveniles 50 to 100 mm TL were caught on the seaward beaches of islands in Louisiana in April, and fish 34 to 57 mm long were caught in marsh areas of the Mobile Delta during December and from February to April. Near the mouth of the Mississippi River, adults and juveniles were captured during summer Life History - 1 (DRAFT) - Life History Species flounder, southern Species Id M010021 Date 27 AUG 96 in addition to a few adults taken in winter. Near Galveston Island, Texas, a single juvenile was captured in September. Juveniles and adults were collected in the Mobile Delta in water of salinities ranging from 0 to 22.2 parts per thousand (ppt). From April 1974 to February 1975, adult southern flounders migrated in shallow waters from the Gulf of Mexico to Aransas Bay, Texas; the migrations were complete by late June. Adult flounders live in Texas bays from June through November, in water with abundant smooth cordgrass (Spartina alterniflora).*77* Growth Characteristics: The largest southern flounder reported in the literature was 762 mm. The von Bertalanffy growth model predicts a maximum age of 20 years. Growth data on the southern flounder are available only from Florida and Mississippi. Annual growth increments in total length (to the nearest 1 mm) for southern flounder in Florida, based on scale measurements and analysis of 177 fish by age group and length (mm), were as follows: 0-I, 79: I-II, 70: II-III, 49; III-IV, 45; IV-V, 46; V-VI, 40; VI-VII, 37; VII-XIII, 34; and VIII-IX, 41 (Nall 1979). Except for ages VIII-IX, growth rate declined with an increase in ages. In Florida the following total length ranges (to the nearest 1 mm) for each age were reported, I, 79; II, 80-142; III, 84-134; and IV, 170.0-215 (Nall 1979). In Mississippi, southern flounders were larger at the same age increment than in Florida: II, 230mm; III, 340 mm; and IV, 480mm.*77* Behavior: In a Louisiana study of day versus night trawling, 89% of southern flounders were caught at night, apparently because they are more vulnerable to trawling at night than during daylight. A tank study confirmed that flounders are more active at night. A tag-recapture study of southern flounders in Texas revealed that movements between and within estuaries rarely exceeded 18 km. The time between release and recapture ranged from 3 to 212 days.*77* REFERENCES FOR LIFE HISTORY- 77 Life History - 2
                           (DRAFT) - Management Practices
                             Species flounder, southern
                                 Species Id M010021
                                   Date 27 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                Species flounder, southern
                                    Species Id M010021
                                      Date 27 AUG 96



     

References

77* Reagan, R. E., W. M. Wingo. 1985. Species Profiles: Life Histories and Environmental Requirements of Coastal Fishes and Invertebrates (Gulf of Mexico) -- Southern Flounder. U.S. Fish and Wildlife Service Biol. Rep. 82(11.30) pp 9. References - 1