(DRAFT) - Taxonomy
                             Species pipefish, northern
                                 Species Id M010118
                                   Date 26 AUG 96



TAXONOMY

NAME - pipefish, northern OTHER COMMON NAMES - Common pipefish, Pipefish and Fr: Syngnathe brun ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - Osteichthyes, ORDER AND SUBORDER - Gasterosteiformes, FAMILY AND SUBFAMILY - Syngnathidae, GENUS AND SUBGENUS - Syngnathus, SPECIES AND SSP - fuscus, SCIENTIFIC NAME - Syngnathus fuscus AUTHORITY - Storer, 1839 TAXONOMY REFERENCES - 258 COMMENTS ON TAXONOMY - Syngnathus Peckianus Storer, 1839 and Siphostoma fuscum Jordan and Gilber, 1882 are synonyms *258*. Dawson and Vari (1982) show a south-north cline in the frequency of dorsal fin rays, and in the frequency of subdorsal trunk rings in S. fuscus; counts higher northward *258*. Taxonomy - 1
                                  (DRAFT) - Status
                             Species pipefish, northern
                                 Species Id M010118
                                   Date 26 AUG 96



STATUS

Coded Status Migrant See Comments REFERENCES FOR STATUS - 258 COMMENTS ON STATUS - Migrations to deep water in autumn and return to shallow water in the spring have been noted in the Chesapeake Bay area *258*. Of no known importance, except that the family is biologically interesting and unusual in that the parental roles are reversed, the male incubating the eggs on his body *258*. Status - 1
                               (DRAFT) - Distribution
                             Species pipefish, northern
                                 Species Id M010118
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE REFERENCES FOR HABITAT - 258 LAND USE - Water Bays and Estuaries REFERENCES FOR LAND USE - 258 COMMENTS ON HABITAT ASSOCIATIONS - Northern pipefish live in beds of seaweed and eelgrass in warmwater coastal areas, salt marshes, and estuaries, sometimes entering brackish water areas. They are carried along the coasts under the masses of floating seaweed, and travels to open sea are rare. A study noted that the report of four specimens taken south of No Mans Land, MA, 5 February 1930, in 34.7 m of water, is of interest because the occurrence was unusual. Migrations to deep water in autumn and return to shallow water in the spring have been noted in the Chesapeake Bay area *258*. ANIMAL/PLANT SPECIES ASSOCIATIONS - Portuguese man-of-war (Physalia pelagica) Cod (Gadus morhua) Cestode (Rhynchobothrium heterospine) Trematode (Cymbephallus vitellosus) Trematode (Opecoeloides manteri) REFERENCES FOR SPECIES ASSOCIATIONS - 258 COMMENTS ON SPECIES ASSOCIATIONS - Predation and competition: There are no published reports of enemies of this species. It may become entangled in tentacles of Physalia pelagica, the Portuguese man-of-war. A study reported that there is a specimen about 15.0 cm long in the United States National Museum (USNM 154775) which was "removed from the stomach of cod" (Gadus morhus) captured on the Grand Bank *258*. Parasites and disease: There is no information on parasites of this species in the Canadian Atlantic region, and infestation in other areas appears light. A study gave details of the parasites reported in the northern pipefish from United States waters. A cestode, Rhynchobothrium heterospine, and two species of digenetic trematodes, Cymbephallus vitellosus and Opecoeloides manteri, are listed *258*. Habitat Associations - 1
                                (DRAFT) - Food Habits
                             Species pipefish, northern
                                 Species Id M010118
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 258 LIFESTAGE FOOD FOOD PART General Crustaceans Not Specified General Osteichthyes Egg/Fetus stage General Osteichthyes Larva stage REFERENCES FOR GENERAL FOOD - 258 COMMENTS ON FOOD - Food of northern pipefish consists of minute organisms, copepods, amphipods, fish eggs, and larvae. A study, referring to descriptions of feeding behavior given by other authors, noted that after water has been expelled from the tubular mouth, prey is taken in with the return rush of water into the mouth *258*. Food Habits - 1
                         (DRAFT) - Environment Associations
                             Species pipefish, northern
                                 Species Id M010118
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Aquatic Vegetation [specified type]: Specified in comments G Estuarine habitat zone: specified in comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 258 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Northern pipefish live in beds of seaweed and eelgrass in warmwater coastal areas, salt marshes, and estuaries, sometimes entering brackish water areas. They are carried along the coasts under the masses of floating seaweed, and travels to open sea are rare. A study noted that the report of four specimens taken south of No Mans Land, MA, 5 February 1930, in 34.7 m of water, is of interest because the occurrence was unusual. Migrations to deep water in autumn and return to shallow water in the spring have been noted in the Chesapeake Bay area *258*. Environment Associations - 1
                               (DRAFT) - Life History
                             Species pipefish, northern
                                 Species Id M010118
                                   Date 26 AUG 96



LIFE HISTORY

Reproduction: There is no definite information on times of spawning in the Canadian Atlantic area. Reports of males carrying eyed-eggs and larvae, 19 mm long, found in August in Cobequid Bay, NS, would support the view that spawning occurs in June and July as reported for Long Island Sound populations. A study noted that spawning in the Gulf of Maine area takes place between March and August. A female in the Chesapeake Bay region, 19.0 cm long, was reported to contain 860 nearly ripe eggs. Males may be in brooding condition at 83 mm, although the brood pouch is seldom developed in males smaller than 90-100 mm. Specimens from Nova Scotia-Massachusetts are reported to have 17-20 pouch rings. The eggs are deposited by the female, a few at a time, into the brood pouch of the male through a small opening (slit) in the pouch, and are supposedly fertilized at this time. Eggs are 0.75-1.0 mm in diameter. Incubation within the pouch occurs in about 10 d and the young are 8 or 9 mm long when they disperse. They live an independent life from time of emergence and do not return to the brood pouch *258*. Growth: Details of growth of wild northern pipefish have not been documented. Observations indicated aquarium specimens were 70 mm long at 2 mo and length-frequency data showed an average length increment of about 60-120 mm TL during the June-October period in the Delaware River estuary. Northern pipefish were mature at 1 yr; however a study estimated an age of 2 yr or more at maturity *258*. Maximum size for Canadian Atlantic pipefish is 30.5 cm long, but they rarely grow to a length greater than 20.3 cm *258*. Description: Body slender, small, elongate, hexagonal in cross section in front of vent, changing to 4-sided behind dorsal fin; 18-21 trunk rings, tail rings 34-39; dorsal keels present, those of body and tail not continuous, lateral keels from tip of pectorals to beginning of dorsal. Head short, slender, tapering; snout tubular, blunt; mouth small, terminal, toothless; eye moderately large, high on head, eyes can rotate independent of each other. Branchiostegals 2 or 3. Fins: soft rayed; dorsal 37-49 (Canadian specimens), base long; caudal rounded; anal 3, minute, absent in males; pelvics absent; pectorals about 12-14. Lateral line absent. Vertebrae 55-60. Size usually about 20 cm long. Color variably dark greenish, olive, brownish above, crossbarred and darkly mottled; lower parts of sides sprinkled with white dots; ventral surface from head to vent golden yellow; brood pouch flaps of males flesh color; dorsal and pectoral fins pale, caudal fin brown. Color may change to suit surroundings *258*. Predation and competition: There are no published reports of enemies of this species. It may become entangled in tentacles of Physalia pelagica, the Portuguese man-of-war. A study reported that there is a specimen about 15.0 cm long in the United States National Museum (USNM 154775) which was "removed from the stomach of cod" (Gadus morhus?) captured on the Grand Bank *258*. Parasites and disease: There is no information on parasites of this species in the Canadian Atlantic region, and infestation in other areas appears Life History - 1 (DRAFT) - Life History Species pipefish, northern Species Id M010118 Date 26 AUG 96 light. A study gave details of the parasites reported in the northern pipefish from United States waters. A cestode, Rhynchobothrium heterospine, and two species of digenetic trematodes, Cymbephallus vitellosus and Opecoeloides manteri, are listed *258*. LIFE HISTORY CODES - Parental Care of Young: Male Breeding/Spawning Season: March Breeding Spawning Season: April Breeding/Spawning Season: May Breeding/Spawning Season: June Breeding/Spawning Season: July Breeding/Spawning Season: August Gestation/Incubation Period: 1-2 weeks REFERENCES FOR LIFE HISTORY- 258 Life History - 2
                           (DRAFT) - Management Practices
                             Species pipefish, northern
                                 Species Id M010118
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                Species pipefish, northern
                                    Species Id M010118
                                      Date 26 AUG 96



     

References

258 * Scott, W.B., M.G. Scott. 1988. Atlantic Fishes of Canada. Canadian Bulletin of Fisheries and Aquatic Sciences (219) (ed.). University of Toronto Press Toronto, Canada:731. References - 1