(DRAFT) - Taxonomy
                              Species flounder, starry
                                 Species Id M010123
                                   Date 27 AUG 96



TAXONOMY

NAME - flounder, starry OTHER COMMON NAMES - ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - , FAMILY AND SUBFAMILY - Pleuronectidae, GENUS AND SUBGENUS - Platichthys, SPECIES AND SSP - stellatus, SCIENTIFIC NAME - Platichthys stellatus AUTHORITY - TAXONOMY REFERENCES - 232 COMMENTS ON TAXONOMY - The starry flounder is a member of the flatfish family, Pleuronectidae. This family is mraine, but a few species, including the starry flounder, enter fresh water *232*. Taxonomy - 1
                                  (DRAFT) - Status
                              Species flounder, starry
                                 Species Id M010123
                                   Date 27 AUG 96



STATUS

Coded Status Migrant See Comments REFERENCES FOR STATUS - 232 COMMENTS ON STATUS - The starry flounders make inshore-offshore migrations with the seasons *232*. The Alaska Department of Fish and Game has managerial authority over starry flounder extending to 3 mi offshore. Because harvesting of starry flounder is minimal, however, no management plan has been formulated *232*. Status - 1
                               (DRAFT) - Distribution
                              Species flounder, starry
                                 Species Id M010123
                                   Date 27 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE AQUATIC REFERENCES FOR HABITAT - 232 LAND USE - Water Streams and Canals Bays and Estuaries REFERENCES FOR LAND USE - 232 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine REFERENCES FOR NWI - 232 COMMENTS ON HABITAT ASSOCIATIONS - The flatfish family is marine, but a few species, including the starry flounder, enter fresh water. Starry flounder is distributed along the coast of the Arctic Region in nearshore areas and brackish water. It is common in brackish water and may ascend rivers even into fresh water *232*. The larvae are pelagic, and after metamorphosis, starry flounders become bottom dwellers *232*. There is evidence that spawning starry flounders use shallow water near river mouths and sloughs *232*. During summer, the fish are inshore, in shallow water and estuaries *232*. Habitat Associations - 1
                                (DRAFT) - Food Habits
                              Species flounder, starry
                                 Species Id M010123
                                   Date 27 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 232 LIFESTAGE FOOD FOOD PART Larva Plankton Not Specified Juvenile Annelids Not Specified Juvenile Polychaetes Not Specified Juvenile Crustaceans Not Specified Juvenile Molluscs Not Specified Juvenile Bivalve Molluscs Not Specified Adult Worms Not Specified Adult Annelids Not Specified Adult Polychaetes Not Specified Adult Molluscs Not Specified Adult Bivalve Molluscs Not Specified Adult Crustaceans Not Specified Adult Malacostraca Not Specified Adult Brittle/Serpent Star Not Specified Adult Plants Not Specified Adult Osteichthyes See Comments Adult Clupeiformes Egg/Fetus stage Larva See Comments; Food See Comments Juvenile See Comments; Food See Comments Adult See Comments; Food See Comments General Molluscs Not Specified General Bivalve Molluscs Not Specified General Crustaceans Not Specified General Malacostraca Not Specified General Osteichthyes See Comments General Clupeiformes Egg/Fetus stage General See Comments; Food See Comments REFERENCES FOR GENERAL FOOD - 232 REFERENCES FOR ADULT FOOD - 232 REFERENCES FOR JUVENILE FOOD - 232 REFERENCES FOR LARVAE FOOD - 232 COMMENTS ON FOOD - Starry flounders are reported to eat razor and surf clams in Kodiak, clams and sand shrimp in Cook Inlet, and herring eggs on kelp, clam siphons, brittle stars, and crabs in the northeast Gulf of Alaska. The primary foods in the northeast Bering Sea and Southeast Chukchi Sea were brittle stars and clams. Starry flounders near Kotzebue were found with herring eggs in their stomachs *232*. Food Habits - 1 (DRAFT) - Food Habits Species flounder, starry Species Id M010123 Date 27 AUG 96 Types of Feeding Areas Used: Starry flounders feed on organisms that live on or burrow into silt substrates. Mud and occasionally, bits of bark and pebbles have been found in the stomachs *232*. Factors Limiting Availability of Food: Light and temperature may limit the production of plankton, which is the food of pelagic larvae of starry flounder. Adults are benthic feeders and forage on many species. Any disruption of the bottom sediment, however, could destroy the food source of adults *232*. Feeding Behavior: The food habits of the starry flounder vary with increases in the size of the fish. Observations of food in the stomachs indicated that starry flounders cracked the shells of large clams, swallowed smaller clams, brittle stars, and pea crabs whole, and swallowed whole or severed at any level nemerteans, priapulids, and polychaetes worms. Starry flounders in Washington apparently began feeding about sunrise and fed through the day until sunset *232*. COMMENTS ON ADULT FOOD - Adults fed mainly on nemertean and polychaete wroms, small clams and crabs, and brittle stars in Washington. Adults from the outer MacKenzie delta in Canada consumed isopods and plant material *232*. COMMENTS ON JUVENILE FOOD - Juvenile starry flounders in Washington consumed polychaete worms, mysids, amphipods, and clams *232*. COMMENTS ON LARVAE FOOD - The pelagic larvae of starry flounder feed on plankton *232*. Food Habits - 2
                         (DRAFT) - Environment Associations
                              Species flounder, starry
                                 Species Id M010123
                                   Date 27 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS J J J L Water Temperature: Specified in Comments L LIM G G Water Depth Preference: Specified in Comments G Water Depth Preference: 500-1000 ft. BA Water Temperature: Below 15 degrees C BA Water Temperature: Specified in Comments E Water Temperature: Below 15 degrees C E Water Temperature: Specified in Comments G Substrate: Sand G Substrate: Rocks G Substrate: Specified in Comments G Bottom Type [Aquatic]: Gravel G Bottom Type [Aquatic]: Specified in Comments FA Substrate: Mud or silt FA Substrate: Specified in Comments BA Water Depth Preference: Specified in Comments BA Water Depth Preference: 50-100 ft. REFERENCES FOR ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR JUVENILE ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR LARVAE ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR EGG ENVIRONMENTAL ASSOC_ - 232 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Water Quality: No information is available on the pH, dissolved oxygen, or Environment Associations - 1 (DRAFT) - Environment Associations Species flounder, starry Species Id M010123 Date 27 AUG 96 turbidity tolerances of starry flounder. Starry flounder tolerates a wide range of salinities. It is common in brackish water and may ascend rivers even into fresh water. Salinity was a limiting factor in the distribution of starry flounder in the southeastern Chukchi Sea. Starry flounders are more common in low-salinity water and are replaced by other forms in high-salinity water *232*. Water Depth: Starry flounders are normally found in shallow water, but catches have been reported from water deeper than 275 m. Depth distribution for starry flounder may be seasonal. They summer in shallow nearshore water and winter in water up to 300 m *232*. Substrate: Starry flounder prefers a soft sand or mud substrate. When distrubed, it may use dorsal, anal, and caudal fins to cover itself with sand or mud. Starry flounder is taken commercially on gravel, clear shifting sand, hard stable sand, and mud, appearing to avoid only rock. Starry flounder can alter its colors and color patterns to match the substrate *232*. COMMENTS ON LIMITING ENVIRONMENTAL ASSOC_ - Salinity was a limiting factor in the distribution of starry flounder in the southeastern Chukchi Sea. Starry flounders are more common in low-salinity water and are replaced by other forms in high-salinity water *232*. COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - Adult starry flounders in Washington stopped feeding in winter, when the bottom temperatures were lowest (8-9 oC) *232*. Starry flounders feed on organisms that live on or burrow into silt substrates. Mud and, occasionally, bits of bark and pebbles were found in the stomachs *232*. COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - Starry flounders are reported to spawn when water temperatures are near 11 oC. There is evidence that spawning starry flounders usu shallow water near river mouths and sloughs. In California, starry flounders in spawning condition were commonly caught in depths of 30 m or less *232*. Spawning occurs from February to April in Puget Sound and British Columbia, and later farther north, with the height of the spawning season corresponding with water temperatures near 11 oC *232*. COMMENTS ON JUVENILE ENVIRONMENTAL ASSOC_ - Juvenile flatfish in California were found in salinities of 6 to 10 ppt *232*. COMMENTS ON LARVAE ENVIRONMENTAL ASSOC_ - Larval starry flounder in the Canadian Northwest Territories were found in colder, more saline waters below the surface waters *232*. COMMENTS ON EGG ENVIRONMENTAL ASSOC_ - Temperatures for hatching have not been reported in the field; however, in Environment Associations - 2 (DRAFT) - Environment Associations Species flounder, starry Species Id M010123 Date 27 AUG 96 the lab, eggs hatched in 68 hours at 12.5 oC and in 110 hours at 10.5 oC *232*. Environment Associations - 3
                               (DRAFT) - Life History
                              Species flounder, starry
                                 Species Id M010123
                                   Date 27 AUG 96



LIFE HISTORY

Feeding Behavior: Starry flounders in Washington apparently began feeding about sunrise and fed through the day until sunset *232*. Reproductive Habitat: There is evidence that spawning starry flounders use shallow water near river mouths and sloughs. In California, starry flounders in spawning condition were commonly caught in depths of 30 m or less *232*. Reproductive Seasonality: In California, ripe, spawning, and spent fish occurred in November, became more numerous in the middle part of January, and gradually disappeared in February. Spawning occurs from February to April in Puget Sound and British Columbia, and later farther north, with the height of the spawning season corresponding with water temperatures near 11 oC *232*. Age at Sexual Maturity: Sexual maturity for starry flounders in California and Washington is reached at age 2 for males and age 3 for females. Females grow faster and reach a larger size than males. The growth of Washington starry flounders slows after sexual maturation, especially in males *232*. Frequency of Breeding: Spawning in California occurs no more often than once a year. The arctic flounder (Liopsetta glacialis), which ranges from Bristol Bay northward, apparently spawns only every second year *232*. Fecundity: The fecundity of a female of 565 mm standard length was about 11,000,000 eggs. The eggs have diameters ranging from .89 to 1.01 mm, are slightly lighter than seawater, and are nonadhesive *232*. Incubation Period/Emergence: The times from fertilization to hatching were found to be 110 hours at 10.5 oC and 68 hours at 12.5 oC. Metamorphosis from the planktonic to benthic life stage occurs when the fish is about 7 mm long at an age ranging from 27 to 104 days. Although size and age both influence the onset of metamorphosis, the influence of size is stronger. The time to metamorphosis is shorter than that for most flatfish, which would limit the dispersal of starry flounder *232*. Movements Associated with Life Functions: The starry flounders make inshore-offshore migrations with the seasons. During summer, the fish are inshore, in shallow water and estuaries. In the winter, they move into deeper water. Tagging has shown that starry flounders seldom move more than 8 km. Juveniles and adults, however, have been recorded moving up rivers as far as 200 km *232*. Natural Factors Affecting Populations: Starry flounders are pelagic in their larval stage, and growth and survival are strongly influenced by physical and biological factors, such as water temperature and planktonic food sources. Cold years or years with low food supply could lead to high mortality and low dispersal, which reduce local recruitment of stocks. In a study conducted in the southeastern Chukchi Sea, an extremely low density of starry flounders, with smaller sizes than reported in more southern waters, was found. It suggested that the physical climate of the Arctic Ocean area Life History - 1 (DRAFT) - Life History Species flounder, starry Species Id M010123 Date 27 AUG 96 may limit the population size of flatfish and depress normal growth patterns *232*. Human-related Factors Affecting Populations: Starry flounder eggs are buoyant, and the larvae are planktonic. These characteristics make them vulnerable to any toxicants that may be released into the water. Juveniles and adults are benthic and rely on benthic food sources, so that any contaminiation or disruption of the sediment could reduce the population. A summary of possible impacts from human-related activities includes: changes in biological oxygen demand or nutrient loading; changes in the chemical composition of water; changes in dissolved oxygen, temperature, pH, or salinity; entrapment; changes in sedimentation rates, turbidity, or suspended solids; changes in substrate composition and location; and shock waves, blasting *232*. LIFE HISTORY CODES - Periodicity: Active at night Average Number of Offspring/Reproductive Effort: Grea Number of Broods/Litters (Reproductive Efforts) Per Ye Gestation/Incubation Period: 3-4 days Gestation/Incubation Period: 5-7 days Breeding/Spawning Season: January Breeding/Spawning Season: February Breeding/Spawning Season: March Breeding Spawning Season: April Breeding/Spawning Season: November Breeding/Spawning Season: December REFERENCES FOR LIFE HISTORY- 232 Life History - 2
                           (DRAFT) - Management Practices
                              Species flounder, starry
                                 Species Id M010123
                                   Date 27 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Controlling sedimentation Beneficial Developing/maintaining suitable pH Beneficial Developing/maintaining suitable salinity Adverse Salinity alteration Beneficial Other management practices [specified in comments] Adverse Other management practices [specified in comments] Adverse Underwater explosions Adverse Applying other toxicants REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 232 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 232 COMMENTS ON MANAGEMENT PRACTICES - Starry flounder eggs are buoyant, and the larvae are planktonic. These characteristics make them vulnerable to any toxicants that may be released into the water. Juveniles and adults are benthic and rely on benthic food sources, so that any contaminiation or disruption of the sediment could reduce the population. A summary of possible impacts from human-related activities includes: changes in biological oxygen demand or nutrient loading; changes in the chemical composition of water; changes in dissolved oxygen, temperature, pH, or salinity; entrapment; changes in sedimentation rates, turbidity, or suspended solids; changes in substrate composition and location; and shock waves, blasting *232*. Management Practices - 1
                                   (DRAFT) - References
                                 Species flounder, starry
                                    Species Id M010123
                                      Date 27 AUG 96



     

References

232 * State of Alaska Department of Fish, and Game. 1986. Alaska Habitat Management Guide: Life Histories and Habitat Requirements of Fish and Wildlife. (ed.). Alaska Department of Fish and Game Juneau, Alaska:763. References - 1