(DRAFT) - Taxonomy
                              Species hagfish, Atlantic
                                 Species Id M010135
                                   Date 27 AUG 96



TAXONOMY

NAME - hagfish, Atlantic OTHER COMMON NAMES - Atlantic hagfish, northern hagfish, hagfish, slime eel and Fr. myxine du nord ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - Chordata, Vertebrata CLASS AND SUBCLASS - , ORDER AND SUBORDER - Myxiniformes, FAMILY AND SUBFAMILY - Myxinidae, GENUS AND SUBGENUS - Myxine, SPECIES AND SSP - glutinosa, SCIENTIFIC NAME - Myxine glutinosa AUTHORITY - Linnaeus, 1758 TAXONOMY REFERENCES - 258 and 1717 COMMENTS ON TAXONOMY - A small family of ancient, bottom-living jawless marine fishes inhabiting deeper parts of the shore waters of temperate seas of the world; principally on continental shelves and slopes, near or on muddy bottoms, where they feed on dead and dying fishes and invertebrates. They may also attack living prey. Unlike the lampreys, they are hermaphroditic and have no larval stage. Upon hatching from the egg, the young resemble the adults. Five genera with about 32 species; only 1 species occurs in the Canadian Atlantic area.*258* Taxonomy - 1
                                  (DRAFT) - Status
                              Species hagfish, Atlantic
                                 Species Id M010135
                                   Date 27 AUG 96



STATUS

REFERENCES FOR STATUS - 258 COMMENTS ON STATUS - The Atlantic hagfish is of no direct commercial importance, but is does constitute a nuisance to commercial fishing activities since it attacks and eats dead and dying fishes caught on line trawls and in bottom gill nets, destroying such fishes for commercial use. Attacks on commercially caught fishes occur principally when the gear is on the bottom. It is of importance in biomedical reseach and has been studied extensively, mainly because it is the most primitive of living vertebrates. Detailed knowledge of its biology, especially its physiology, may aid in our understanding of the evolution of fishes. It also has three hearts, one without nerves. Studies of these and other organs have been conducted in hopes of providing insight into cardiac function and activity in mammals. Detailed studies of blood chemistry, endocrine system, and immune responses, to name only a few areas of interest, have been made.*258* Status - 1
                               (DRAFT) - Distribution
                              Species hagfish, Atlantic
                                 Species Id M010135
                                   Date 27 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE REFERENCES FOR HABITAT - 258 LAND USE - Bays and Estuaries Water REFERENCES FOR LAND USE - 258 COMMENTS ON HABITAT ASSOCIATIONS - Atlantic hagfish live on soft, muddy bottoms, usually in depths of 30 m or more, at times to a depth of 958 m. They require the high-salinity (about 30 ppt or more) and low-temperature (below 12 degrees C) conditions usually found in deeper waters. Aquarium observations indicate that they burrow in the soft bottom and may assume a position with the head emergent from the bottom. However, they may also remain completely buried in mud for long periods, suggesting that the species is facultatively anaerobic. Certainly they may remain inactive for long periods. They are nocturnal, being more active at night than in daylight hours, although they are blind. They must be quite abundant in certain locations, such as in Passamaquoddy Bay, NB, judging from the number caught in traps. A trap made from a 25 gal. steel drum, baited with flounder and set overnight in 50 m, may catch upwards of 500 or more hagfish in a single night.*258* ANIMAL/PLANT SPECIES ASSOCIATIONS - Predation and Competition: The Atlantic hagfish has few known enemies, although small hagfish have been found in the stomachs of cod and white hake and six hagfish eggs were taken from the stomachs of halibut caught over eastern Sable Bank. It would seem to be risky for a cod, however, to attempt to eat a large adult hagfish because of the great quantities of viscous slime hagfish can produce. Indeed, the production of large quantities of slime may be related to feeding, for a fish predator, upon grasping a hagfish in its mouth, would likely find its mouth and gills suffused with the viscous, suffocating slime, which could cause death, thus providing the intended victim with a meal. As a scavenger, the hagfish must compete with other creatures of similar food habits, such as crabs, that also feed on dead organisms found on the ocean floor.*258* Habitat Associations - 1
                                (DRAFT) - Food Habits
                              Species hagfish, Atlantic
                                 Species Id M010135
                                   Date 27 AUG 96



FOOD HABITS

TROPHIC LEVEL - SCAVENGER REFERENCES FOR TROPHIC LEVEL - 258 LIFESTAGE FOOD FOOD PART General Polychaetes Not Specified General Crustaceans Not Specified General Fish See Comments General Malacostraca Not Specified Important Crustaceans Not Specified REFERENCES FOR GENERAL FOOD - 258 COMMENTS ON FOOD - Food: Atlantic hagfish comsume the internal organs and flesh or dead and dying creatures found on the sea floor. They will attack, burrow into, and eat fishes caught on bottom longlines or bottom gill nets. The fishes are attacked and entry to the body gained by using the sharp lingual or tongue teeth in a rasping manner, or by biting onto the skin or the dead fish and pulling against the bitten skin until a piece is removed; or entry may be via the mouth or anus, the viscera being eaten first and then the muscle, leaving only skin and bones. European studies indicated that many invertebrates, such as polychaete worms and crustaceans are also consumed. Few food studies of wild-caught Atlantic hagfishes have been conducted but a study of gut content over 100 North Sea specimens suggested that the species may be a more active predator than previously thought. It was found that shrimp, Pandalus borealis, was an important food item. The evidence presented suggests that P. borealis was selected and preyed upon by the hagfish examined. Thus the Atlantic hagfish may be an active predator on shrimps and possibly other species.*258* Predation and Competition: The Atlantic hagfish has few known enemies, although small hagfish have been found in the stomachs of cod and white hake and six hagfish eggs were taken from the stomachs of halibut caught over eastern Sable Bank. It would seem to be risky for a cod, however, to attempt to eat a large adult hagfish because of the great quantities of viscous slime hagfish can produce. Indeed, the production of large quantities of slime may be related to feeding, for a fish predator, upon grasping a hagfish in its mouth, would likely find its mouth and gills suffused with the viscous, suffocating slime, which could cause death, thus providing the intended victim with a meal. As a scavenger, the hagfish must compete with other creatures of similar food habits, such as crabs, that also feed on dead organisms found on the ocean floor.*258* Food Habits - 1
                         (DRAFT) - Environment Associations
                              Species hagfish, Atlantic
                                 Species Id M010135
                                   Date 27 AUG 96



ENVIRONMENTAL ASSOCIATIONS

Environment Associations - 1
                               (DRAFT) - Life History
                              Species hagfish, Atlantic
                                 Species Id M010135
                                   Date 27 AUG 96



LIFE HISTORY

Description: Body elongate, eel-shaped, round in cross section anteriorly, compressed posteriorly; a single gill pore on each side just anterior to ventral fin fold origin; line of nucous pores, low on each side of body, from behind gill openings to beyond anus. Mouth lipless, with one pair of smaller barbels directed posteriorly and one pair of smaller barbels directed anteriorly on either side of single nostril at tip of snout; jawless and toothless, orange-colored tongue (or lingual) teeth present which can be everted for burrowing into host; eye blind, covered by skin and layer of muscle. Fins; no paired fins, no fin rays, a fold of skin resembling a fin begins dorsally about two-thirds back from snout and just before anus and continues around tail, continuing forward on ventral surface to about one-third distance behind snout. Size 46-61 cm long, rarely longer. Color uniformly brown, reddish, or gray above; sometimes mottled. Reproduction: Spawning apparently occurs throughout the year. Female hagfish may carry 1-30 horny-shelled large eggs, each up to 25 mm in length. The eggs are whitish, ovoid in shape, with a cluster of filaments at each end. Hagfish eggs are seldom found in the wild but have been taken in the Bay of Fundy, on Georges Bank, and off the south coast of Newfoundland. Atlantic hagfish are not functional hemaphrodites but each sexually mature individual possesses undeveloped gonadal tissue of the opposite sex. Growth: Newly hatched Atlantic hagfish resemble the adults and thus do not have a larval stage. The smallest individuals that have been seen are about 6.0-7.0 cm long. The rate of growth is unknown and there is no known way to age them. Specimens taken in Passamaquoddy Bay average about 35.0 cm TL.*258* Predation and Competition: The Atlantic hagfish has few known enemies, although small hagfish have been found in the stomachs of cod and white hake and six hagfish eggs were taken from the stomachs of halibut caught over eastern Sable Bank. It would seem to be risky for a cod, however, to attempt to eat a large adult hagfish because of the great quantities of viscous slime hagfish can produce. Indeed, the production of large quantities of slime may be related to feeding, for a fish predator, upon grasping a hagfish in its mouth, would likely find its mouth and gills suffused with the viscous, suffocating slime, which could cause death, thus providing the intended victim with a meal. As a scavenger, the hagfish must compete with other creatures of similar food habits, such as crabs, that also feed on dead organisms found on the ocean floor.*258* Life History - 1
                           (DRAFT) - Management Practices
                              Species hagfish, Atlantic
                                 Species Id M010135
                                   Date 27 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                Species hagfish, Atlantic
                                    Species Id M010135
                                      Date 27 AUG 96



     

References

258 * Scott, W.B., M.G. Scott. 1988. Atlantic Fishes of Canada. Canadian Bulletin of Fisheries and Aquatic Sciences (219) (ed.). University of Toronto Press Toronto, Canada:731. References - 1