(DRAFT) - Taxonomy
                             Species minnow, sheepshead
                                 Species Id M010140
                                   Date 26 AUG 96



TAXONOMY

NAME - minnow, sheepshead OTHER COMMON NAMES - ELEMENT CODE - 06/12/84 AOU CODE - 04/24/85 CATEGORY - Fish PHYLUM AND SUBPHYLUM - Chordata, Vertebrata CLASS AND SUBCLASS - Osteichthyes, Actinopterygii ORDER AND SUBORDER - Atheriniformes, Cyprnodontoidei FAMILY AND SUBFAMILY - Cyprinodontidae, GENUS AND SUBGENUS - Cyprinodon, SPECIES AND SSP - variegatus, SCIENTIFIC NAME - Cyprinodon variegatus AUTHORITY - Lacepede TAXONOMY REFERENCES - 2539, 2635 and 3349 COMMENTS ON TAXONOMY - Subspecies 1) C. v. ovinus (Mitchell) MA-VA; 2) C. v. variegatus (Lacepede) NC-TX; 3) C. v. riverendi (Poey) Cuba *2635,3349* Taxonomy - 1
                                  (DRAFT) - Status
                             Species minnow, sheepshead
                                 Species Id M010140
                                   Date 26 AUG 96



STATUS

Status - 1
                               (DRAFT) - Distribution
                             Species minnow, sheepshead
                                 Species Id M010140
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - 001 and 131 References on County Abundance - 001 and 131 REFERENCES FOR HYDROLOGIC UNIT CODES - 001 and 131 REFERENCES FOR OTHER DISTRIBUTION - A Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - Aquatic REFERENCES FOR HABITAT - 835, 1230 and 2624 LAND USE - Water Bays and Estuaries REFERENCES FOR LAND USE - 1230, 2624 and 2199 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine, intertidal FL2 N 3 Marine, intertidal FL3 N 3 Estuarine, intertidal FL2 N 3 Estuarine, intertidal FL3 N 3 Estuarine, intertidal FL4 N 3 REFERENCES FOR NWI - 835, 1230, 2624, 2199, 3348, 3350 and 2775 COMMENTS ON HABITAT ASSOCIATIONS - Occurs in sea-brackish-freshwater; most common in brackish, close to shore *2775* POTENTIAL NATURAL VEGETATION - 065 Northern Cordgrass Prairie (Distichlis-Spartina) 101 Oak - Hickory - Pine Forest (Quercus-Carya-Pinus) REFERENCES FOR PNV - 4205 ECOREGION - Southeastern Mixed Forest: Flat Plains REFERENCES FOR ECOREGION - 4205 Habitat Associations - 1
                                (DRAFT) - Food Habits
                             Species minnow, sheepshead
                                 Species Id M010140
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - Omnivore REFERENCES FOR TROPHIC LEVEL - 835, 2635, 1371 and 1362 LIFESTAGE FOOD FOOD PART General Plants Not Specified General Osteichthyes See Comments General Animals Not Specified Adult Plants Not Specified Adult Osteichthyes See Comments Adult Animals Not Specified REFERENCES FOR GENERAL FOOD - 835, 2635, 1371 and 1362 REFERENCES FOR ADULT FOOD - 835, 2635, 1371 and 1362 COMMENTS ON FOOD - 7300S=dead or disabled fish *835,1362* COMMENTS ON ADULT FOOD - 7300S=dead or disabled fish *835,1362* Food Habits - 1
                         (DRAFT) - Environment Associations
                             Species minnow, sheepshead
                                 Species Id M010140
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Temperature: Greater than 27 degrees C G Water Temperature: Between 21-27 degrees C G Water Temperature: Between 15-21 degrees C G Water Temperature: Below 15 degrees C G Turbidity: Clear water G Substrate: Mud or silt G Substrate: Sand G Relation to Substrate: Unattached - normally free living G Bottom Type [Aquatic]: Mud or silt G Bottom Type [Aquatic]: Sand G Density of Aquatic Vegetation: Low G Gradient: Low G Flow: Less than 5 cfs mean annual flow G Aquatic Habitat Zonation: Specified in Comments E Water Temperature: Between 21-27 degrees C E Water Temperature: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 1230, 2624, 2199, 3347, 3348, 3350 and 2775 REFERENCES FOR EGG ENVIRONMENTAL ASSOC_ - 1230 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - 00190S=shallow with little or no vegetation *3348*; 00220S=preference of <20 ppt however has been recorded surviving up to 140 ppt *3348* COMMENTS ON EGG ENVIRONMENTAL ASSOC_ - 00020S=temperature of 25 degrees C necessary for oocytes to mature *1230* Environment Associations - 1
                               (DRAFT) - Life History
                             Species minnow, sheepshead
                                 Species Id M010140
                                   Date 26 AUG 96



LIFE HISTORY

PHYSICAL DESCRIPTION:size=about 3 inches *2635,2637,1371,3118*; color, male=steel blue-green back with yellow-orange sides and belly *835,1230,2635,3118,3350,1549* female=paler, more brown *3350*; REPRODUCTION:breeding season=April to early fall, 25 C water essential to breed, in areas of warm water breeding may occur year round *1230,3346,2637,2199,3349,3351,1362*; incubation period=5-10 days *1230,2635,2199,3351*; broods/year=more than one considered likely over entire range, dependent upon length of season *3351,1362*; age at maturity=both sexes mature at one year of age *2635*; breeding behavior=1) male constructs nest, 2) male attacks female entering territory, 3) female either flees or remains in territory, 4)if the female remains the two fish swim together to the nest where the egg and sperm are released *835,1230,3350,1549,1362*; BEHAVIOR: territoriality=male defends territory during the spawning season; larger males dominate over the smaller in both defense and the size of territory *835,1230,3349,1549,3350,1362*; larger territories are up to two feet in diameter *3350*; after the spawning season the minnow travels in small schools *1371,3350*; developement of young=eggs hatch after 5-10 days; new larvae = 4mm, large yolk sac, continuous dorsal and ventral fin; after 5 days yolk almost absorbed, larvae >5mm body depth increased; at 9mm=many adult characteristics present, slender body, back not fully elevated; at 13mm=almost all adult characteristics developed, back elevated, caudal fin still more rounded than adult, still lighter in color than adult *2199,3351*; ORIGIN: native *835,2637,3118,3350*; AQUATIC/TERRESTIAL ASSOCIATIONS: in Florida=positively associated with Fundulus confluentes and negatively associated with Menidii beryllina, Eucinostomus argenteus, Anchoa m. diaphana, Floridichthys carpio *3348* REFERENCES FOR LIFE HISTORY- 835, 1230, 3346, 2635, 2637, 1371, 3118, 2199, 3348, 3351, 3350, 1549 and 1362 Life History - 1
                           (DRAFT) - Management Practices
                             Species minnow, sheepshead
                                 Species Id M010140
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                Species minnow, sheepshead
                                    Species Id M010140
                                      Date 26 AUG 96



     

References

835* Cooper, E.L. 1983. The Fishes of Pennsylvania. Penn. State Univ. Press University Park. 1230* Wang, J.C.S., Kernehan, R.J. 1979. FISHES OF THE DELAWARE ESTUARIES. Ecological Analysts, Inc. Towson, Md:410. 1362* Hildebrand, S.F., Schroeder, W.C. 1928. Fishes of Chesapeake Bay. Bull. U.S. Bur. Fish. 43 (1927) Pt. 1:366. 1371* Perlmutter, A. 1961. GUIDE TO MARINE FISHES. New York Univ. Press. 1549* Newman, H.H. 1907. Spawning behavior and sexual dimorphism in Fundulus heteroclitus and allied fish. Biol. Bull. 12(5):314-348. 2199* Lipson, A.J., Moran, R.L. 1974. MANUAL FOR IDENTIFICATION OF EARLY DEVELOPMENTAL STAGES OF FISHES OF THE POTOMAC RIVER ESTUARY. POWER PLANT SITING PROGRAM OF THE MARYLAND DEPARTMENT OF NATURAL RESOURCES. (PPSP-MP-13). Md. Dep. Nat. Resour. Md:282. 2539* Grzimek, B. 1972. GRZIMEK'S ANIMAL LIFE ENCYCLOPEDIA.. 2624* Swift, C., Yerger, R.W., Parrish, P.R. 1977. Distribution and natural history of the fresh and brackish water fishes of the Ochlockonee River, Florida and Georgia. Tall Timber Res. Sta. Tallahassee. 2635* Brender, C. 1948. FIELD BOOK OF MARINE FISHES OF THE ATLANTIC COAST. G.P. Putnam's Sons New York. 2637* Ursin, M.J. 1972. A GUIDE TO FISHES OF THE TEMPERATE ATLANTIC COAST. E.P. Dutton New York. 2775* Bailey, R.M., Winn, H.E., Smith, C.L. 1954. Fishes from the Escambia River, Alabama and Florida, with ecological and taxonomic notes. Proc. Nat. Acad. Sci.(Phila.) 106:109-164. 3118* Sterba, G. 1973. FRESHWATER FISHES OF THE WORLD.. TFH Publications. 3346* DeVlaming, V.L., Kuris, A., Parker, F.R., Jr. 1978. Seasonal variation of reproduction and lipid reserves in some subtropical Cyprinodontids. Trans. Amer. Fish. Soc. 107:464-472. 3347* Usher, R.B., II, Bengtson, D.A. 1981. Survival and growth of sheepshead minnow larvae and juveniles on a diet of Artemia nauplii. Prog. Fish-Cult. 43:102-105. 3348* Kilby, J.D. 1955. The fishes of two Gulf Coastal marsh References - 1 (DRAFT) - References Species minnow, sheepshead Species Id M010140 Date 26 AUG 96 areas in Florida. Tulane Stud. Zoo. 2:175-247. 3349* Simpson, D.G., Gunter, G. 1956. Notes on habitats, systematic characters and life histories of Texas saltwater Cyprinodonts. Tulane Stud. Zoo. 4:115-134. 3350* Raney, E.C., al, et. 1953. Reproductive behavior in Cyprinodon variegatus, Lacepede, in Florida. Zoologica 38:97-104. 3351* Kuntz, A. 1914. Notes on the embryology and larval development of 5 species of Teleostean fishes. Bull. U.S. Bur. Fish. 34:409-429. 4205* Jenkins, R.E. 1984. Fishes of Virginia (tentative)..(publication pending) References - 2