(DRAFT) - Taxonomy
                               Species bass, black sea
                                 Species Id M010169
                                   Date 27 AUG 96



TAXONOMY

NAME - bass, black sea OTHER COMMON NAMES - black sea bass, blackfish, black bass and black will ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Osteichthyes, ORDER AND SUBORDER - Perciformes, FAMILY AND SUBFAMILY - Serranidae, GENUS AND SUBGENUS - Centropristis, SPECIES AND SSP - striata, SCIENTIFIC NAME - Centropristis striata AUTHORITY - Linnaeus TAXONOMY REFERENCES - 201 COMMENTS ON TAXONOMY - A subspecies, C.s. melana, occurs along the eastern and northern coastal areas of the Gulf of Mexico. Taxonomy - 1
                                  (DRAFT) - Status
                               Species bass, black sea
                                 Species Id M010169
                                   Date 27 AUG 96



STATUS

Coded Status Sport Fish Commercial Migrant Existing, FMP exists Commercial/consumption REFERENCES FOR STATUS - 184, 191 and 201 COMMENTS ON STATUS - Management regulations for black sea bass caught in the Fishery Conservation Zone (the area in which the United States asserts exclusive fishery management authority and which extends seaward from 3 to 200 nm) were established in the Fishery Management Plan for the snapper-grouper fishery of the South Atlantic Region (Prepared by the South Atlantic Fishery Management Council, March, 1983).*201* For both the commercial and recreational sectors, all species in the management unit are sought as food fish.*184* Black sea bass are migratory in the northern part of their range, whereas south of Cape Hatteras, North Carolina, they are apparently residents in given areas year round. In the Middle Atlantic Bight black sea bass move inshore and northward in spring and offshore and south in fall, probably due to changes in temperature.*201* Status - 1
                               (DRAFT) - Distribution
                               Species bass, black sea
                                 Species Id M010169
                                   Date 27 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC REFERENCES FOR HABITAT - 201 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Estuarine Marine REFERENCES FOR NWI - 201 COMMENTS ON HABITAT ASSOCIATIONS - Black sea bass eggs are pelagic. Little is known of the early life history of black sea bass. The pelagic existence of this fish is short. Larvae longer than 13 mm SL were not collected, presumably because they become demersel or estuarine near that size. It is not known what part of the population remains at sea, nor are the routes and mechanisms of larval transport known. Juvenile black sea bass occur in saline areas of estuaries along the coast from Florida to Massachusetts and in offshore areas.*201* Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species bass, black sea
                                 Species Id M010169
                                   Date 27 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 201 LIFESTAGE FOOD FOOD PART General Echinoderms Not Specified General Molluscs Not Specified General Crustaceans Not Specified General Fish Not Specified General See Comments; Food Not Applicable Adult Crustaceans Not Specified Adult Fish Not Specified Adult See Comments; Food Not Applicable Juvenile Crustaceans Not Specified Juvenile See Comments; Food Not Applicable Important Crustaceans Not Specified Important Fish Not Specified Important See Comments; Food Not Applicable REFERENCES FOR GENERAL FOOD - 201 REFERENCES FOR IMPORTANT FOOD - 201 REFERENCES FOR ADULT FOOD - 201 REFERENCES FOR JUVENILE FOOD - 201 COMMENTS ON FOOD - Black sea bass are carnivorous bottom-feeders that eat crustaceans (primarily crabs and shrimp), fish, mollusks, and echinoderns.*201* COMMENTS ON ADULT FOOD - Adults eat mainly crabs and fish. *201* COMMENTS ON JUVENILE FOOD - The young eat shrimp, isopods, and amphipods.*201* Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species bass, black sea
                                 Species Id M010169
                                   Date 27 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Temperature: Greater than 27 degrees C G Water Temperature: Between 21-27 degrees C G Water Temperature: Between 15-21 degrees C G Water Temperature: Below 15 degrees C G Water Temperature: Specified in Comments G Bottom Type [Aquatic]: Specified in Comments E Water Temperature: Between 15-21 degrees C E Water Temperature: Specified in Comments L Water Depth Preference: Less than 1 ft. L Water Depth Preference: 1-5 ft. L Water Depth Preference: 5-10 ft. L Water Depth Preference: 10-25 ft. L Water Depth Preference: 25-50 ft. L Water Depth Preference: 50-100 ft. L Water Depth Preference: 100-200 ft. L Water Depth Preference: Specified in Comments J Water Temperature: Greater than 27 degrees C J Water Temperature: Between 21-27 degrees C J Water Temperature: Between 15-21 degrees C J Water Temperature: Below 15 degrees C J Water Temperature: Specified in Comments J Bottom Type [Aquatic]: Specified in Comments J J J J J J J J J Estuarine habitat zone: specified in comments A Water Temperature: Specified in Comments A Water Depth Preference: 25-50 ft. A Water Depth Preference: 50-100 ft. A Water Depth Preference: 100-200 ft. A Water Depth Preference: 200-500 ft. A Water Depth Preference: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 201 Environment Associations - 1 (DRAFT) - Environment Associations Species bass, black sea Species Id M010169 Date 27 AUG 96 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 201 REFERENCES FOR FEEDING JUVENILE ENVIRONMENTAL ASSOC_ - 201 REFERENCES FOR EGG ENVIRONMENTAL ASSOC_ - 201 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - In North Carolina estuaries, black sea bass were collected at 6 to 29 degrees C.*201* The black sea bass is a dominant species associated with the hard-or live-bottom sponge-coral habitat in the South Atlantic Bight. This habitat, which is located on the open shelf from depths of 18 to 55 m, consists of low rock ridges, outcroppings, coral patches, and shipwrecks. The black sea bass is also one of the most abundant species on artificial reefs along the South Atlantic coast.*201* COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - Adult black sea bass are found in offshore areas in depths of 10 to 120 m; however, most are at 20 to 60 m.*201* Off South Carolina the lowest bottom temperatures recorded in the depth range inhabited by adult sea bass are 10 degrees C or higher during all seasons. Although no seasonal movements were observed in South Carolina waters, it has been noted that larger specimens were found in deep water, which suggest gradual offshore movements with increasing age; however, reduced fishing pressure offshore away from ports may be a factor. It has been reported that average age increased with depth from 2.4 years at 18-26 m to 4.8 years at 37-46 m.*201* COMMENTS ON FEEDING JUVENILE ENVIRONMENTAL ASSOC_ - Juvenile black sea bass occur in saline areas of estuaries along the coast from Florida to Massachusetts and in offshore areas. Within estuaries, juveniles are found around jetties, piers, wrecks, and shell bottom such as oyster reefs.*201* Black sea bass have been collected year round in North Carolina estuaries at salinities of 1 to 36 ppt and temperatures of 6 to 29 degrees C. In South Carolina estuaries juveniles (20-140 mm SL) were found at salinities of 8.8 to 37.8 ppt and temperatures of 5.6 to 30.4 degrees C, mainly from July to November. Young-of-the-year black sea bass have been collected throughout the year in the lower reaches of a Georgia estuary, mostly over shell bottom. COMMENTS ON FEEDING LARVAE ENVIRONMENTAL ASSOC_ - Larvae were collected in tows from the surface to a depth of 33m, 4 to 82 km from shore where water depths ranged from 15 to 51 m.*201* COMMENTS ON EGG ENVIRONMENTAL ASSOC_ - Black sea bass eggs are pelagic, are 0.9-1.0 mm in diameter and hatch in about 75 h at 16 degrees C. Eggs from artificially spawned gulf black sea bass hatch in about 38 h after fertilization at 23 degrees C under an 11 h photoperiod.*201* Environment Associations - 2
                               (DRAFT) - Life History
                               Species bass, black sea
                                 Species Id M010169
                                   Date 27 AUG 96



LIFE HISTORY

Morphology: Dorsal X, 11; anal III, 7; total gill rakers plus tubercles usually 22-28 (20-29), lower limb usually 14 to 18 (14-19); scales large, lateral line scales 47 (46-49). Body robust, the back somewhat elevated anteriorly; head large; snout moderately pointed; mouth large, oblique. Spinous and soft dorsal fins continuous, males having the noticeabley higher fins; caudal fin rounded in juveniles, trilobed and with one upper ray extended in adults. Breeding males develop an adipose hump on the nape.*201* Color in life: smokey gray, dusky brown, or blue-black above, slightly paler below; centers of scales pale blue or white, forming longitudinal stripes along back and sides; sides sometimes mottled or with dark and light vertical crossbars. Breeding males with vivid hues of fluorescent blue and green around eyes and nape; females are lighter and brownish or gray-blue instead of blue-black. Four juvenile color phases have been described: an overall light grayish phase peppered with small dark spots; a dark phase with dull white spots; a striped phase with a single horizontal dark stripe; and a barred phase with six vertical bars. The Atlantic subspecies of black sea bass differs from Gulf of Mexico subspecies in the following details: gill rakers and tubercles usually 25 to 26, instead of 22-24; pectoral rays 18 or 19, instead of 17 or 18; jaw usually 16% or 17% of standard length, instead of 18% to 19%; black spot at base of last dorsal spines distinct in juveniles, rather than indistinct to almost absent; dark blotches on jaws usually absent in young. Other species of Centropristis differ in being very pale to white, with seven diffuse to distinct dark vertical bars on side, fins pale, soft dorsal and caudal with dark to inky spots.*201* Spawning: Black sea bass are protogynous hermaphrodites. Most individuals function first as females, undergo sexual succession, and become functional males. Spawning of black sea bass extends from January to June offshore in the South Atlantic region, peaking from March to May. Minor spawning occurs in September and October. Spawning occurs later in the northern part of the range. The frequency of occurence of transitional individuals increases after both spawning periods, indicating that sexual succession is primarily a post-spawning process.*201* Black sea bass mature between age I and IV. Sex ratios of black sea bass differ significantly from 1:1 in favor of females at the smaller sizes and younger ages. An increased frequency of males occurs between 200 and 240 mm SL and ages IV and V. Individuals undergoing sexual succession ranged in length from 118 to 327 mm SL and from ages I to VIII; modes were at 200 mm SL and age III.*201* Fecundity and eggs: Estimates of fecundity have ranged from 17,000 in a fish of 108 mm SL (age II) to 1,050,000 in a fish of 438 mm SL of undetermined age. Fecundity was significantly related to length, weight, and age.*201* Black sea bass eggs are pelagic, are 0.9-1.0 mm in diameter and hatch in about 75 h at 16 degrees C.*201* Larvae: Life History - 1 (DRAFT) - Life History Species bass, black sea Species Id M010169 Date 27 AUG 96 The pelagic existence of this fish is short. Larvae longer than 12 mm SL were not collected, presumably because they become demersal or estuarine near that size. It is not known what part of the population remains at sea, nor are the routes and mechanisms of larval transport known.*201* Juveniles: Juvenile black sea bass occur in saline areas of estuaries along the coast from Florida to Massachusetts and in offshore areas. Within estuaries, juveniles are found around jetties, piers, wrecks, and shell bottom such as oyster reefs.*201* Black sea bass have been collected year round in North Carolina estuaries at salinities of 1 to 36 ppt and temperatures of 6 to 29 degrees C. In South Carolina estuaries juveniles (20-140 mm SL) were found at salinities of 8.8 to 37.8 ppt and temperatures of 5.6 to 30.4 degrees C, mainly from July to November. Young-of-the-year black sea bass have been collected throughout the year in the lower reaches of a Georgia estuary, mostly over shell bottom.*201* Migration: Black sea bass are migratory in the northern part of their range, whereas south of Cape Hatteras, North Carolina, they are apparently residents in given areas year round. In the Middle Atlantic Bight black sea bass move inshore and northward in spring and offshore and south in fall, probably due to changes in temperature. Off South Carolina the lowest bottom temperatures recorded in the depth range inhabited by adult sea bass are 10 degrees C or higher during all seasons. Although no seasonal movements were observed in South Carolina waters, it has been noted that larger specimens were found in deep water, which suggest gradual offshore movements with increasing age; however, reduced fishing pressure offshore away from ports may be a factor. It has been reported that average age increased with depth from 2.4 years at 18-26 m to 4.8 years at 37-46 m.*201* LIFE HISTORY CODES - Breeding/Spawning Season: January Breeding/Spawning Season: February Breeding/Spawning Season: March Breeding Spawning Season: April Breeding/Spawning Season: May Breeding/Spawning Season: June Gestation/Incubation Period: 3-4 days Average Number of Offspring/Reproductive Effort: Grea REFERENCES FOR LIFE HISTORY- 201 Life History - 2
                           (DRAFT) - Management Practices
                               Species bass, black sea
                                 Species Id M010169
                                   Date 27 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Existing Regulating harvest of species being described Existing Regulating harvest - setting size limits Existing Regulating commercial harvest gear types Existing Regulating commercial harvest levels REFERENCES FOR EXISTING MANAGEMENT PRACTICES - 201 COMMENTS ON MANAGEMENT PRACTICES - A minimum size limit of 203 mm TL (8 inches) was adopted for black sea bass on the basis of a yield-per-recruit analysis that indicated the harvest is less than the maximum possible.*201* Restrictions on fish trap dimensions and trap fishing areas were adopted for black sea bass caught in the South Atlantic Fishery Conservaton Zone. Minimum mesh size of traps is 1 by 2 inches (rectangular) or 1 1/2 inches (hexagonal).*201* Commercial harvest: In the South Atlantic region black sea bass are mainly harvested by a trap fishery; small amounts are caught with trawls and hand lines. In the early 1900's the fishery was primarily a handline fishery. The trap fishery, which developed in 1960, operates year-round along the 18 m contour. Wire-mesh crab pots are used, baited with menhaden, herring, squid, mullet, spot, or croaker. Highest landings occur from early winter to spring due to increased participation in the fishery by shrimp and sport fishermen. Fifteen to 40 traps are set singly over patch reefs. Fishing time ranges from 20 to 45 minutes for each trap. This fishery differs considerably from the trap fishery north of Cape Hatteras, North Carolina, which operates from May to December, and in which 400 to 1,5000 unbaited wooden slat traps are set out for the entire season in strings of 10 or 20 traps per line, with about 200 traps fished daily. Trawl catches of black sea bass are primarily landed north of Cape Hatteras, North Carolina. Black sea bass are also caught in the handline fishery for snappers and groupers. Black sea bass landings increased sharply in the South Atlantic region after 1960 due to the development of the trap fishery.*201* The South Atlantic Fishery Management Council concluded from YPR analysis that black sea bass are probably in the range of growth overfishing, i.e.; excessive harvesting of small fish. Sport fishing: The recreational catch of black sea bass has increased, partly due to the development of the headboat fishery in the mid-1960's. The black sea bass is the most abundant species in the North Carolina charter boat bottom fishing catches of reef fishes, accounting for 75% of the catch by number and 64% of the catch by weight.*201* Management Practices - 1
                                   (DRAFT) - References
                                 Species bass, black sea
                                    Species Id M010169
                                      Date 27 AUG 96



     

References

184 * South Atlantic Fishery Management Council. 1983. Fishery Management Plan, Regulatory Impact Review, and Final Environmental Impact Statement for the Snapper-Grouper Fishery of the South Atlantic Region. South Atlantic Fishery Management Council (ed.). National Marine Fishery Service:273. 191 * South Atlantic Fishery Management Council. 1988. Amendment Number 1 and Environmental Assessment and Regulatory Impact Review To The Fishery Management Plan For The Snapper Grouper Fishery of The South Atlantic Region. (ed.):46. 201 * Mercer, Linda. 1989. Species Profiles: Life Histories and Environmental Requirements of Coastal Fishes and Invertebrates (South Atlantic) --Black Sea Bass. Species Profile Series from the Fish and Wildlife Service 82(11.99) (ed.):16. References - 1