(DRAFT) - Taxonomy
                            Species perch, Pacific Ocean
                                 Species Id M010240
                                   Date 26 AUG 96



TAXONOMY

NAME - perch, Pacific Ocean OTHER COMMON NAMES - ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - , FAMILY AND SUBFAMILY - , GENUS AND SUBGENUS - Sebastes, SPECIES AND SSP - alutus, SCIENTIFIC NAME - Sebastes alutus AUTHORITY - TAXONOMY REFERENCES - 232 Taxonomy - 1
                                  (DRAFT) - Status
                            Species perch, Pacific Ocean
                                 Species Id M010240
                                   Date 26 AUG 96



STATUS

Coded Status Migrant Depleted Existing, FMP exists See Comments REFERENCES FOR STATUS - 232 COMMENTS ON STATUS - Adult Pacific ocean perch do not migrate long distances. Seasonal movements of perch are largely between deep and shallow bottoms within a limited area *232*. Before heavy commercial exploitation began in the early 1960's, Pacific ocean perch were a dominant groundfish in the Gulf of Alaska. But perch stocks now have been severely reduced *232*. Pacific ocean perch within the 200-mi limit are managed by the North Pacific Fishery Management Council in their groundfish fishery management plans *232*. Status - 1
                               (DRAFT) - Distribution
                            Species perch, Pacific Ocean
                                 Species Id M010240
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE REFERENCES FOR HABITAT - 232 LAND USE - Water Bays and Estuaries REFERENCES FOR LAND USE - 232 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine REFERENCES FOR NWI - 232 COMMENTS ON HABITAT ASSOCIATIONS - Pacific ocean perch were found in each region but were generally restricted to outer shelf and upper slope depth zones. Adult Pacific ocean perch are generally found on the continental slope *232*. ANIMAL/PLANT SPECIES ASSOCIATIONS - Anemone (Metridium senile) Walleye pollock (Theragra chalcogramma) REFERENCES FOR SPECIES ASSOCIATIONS - 232 COMMENTS ON SPECIES ASSOCIATIONS - Juveniles inhabit areas where cover and protection among sessile invertebrates such as the anemone (Metridium senile). Pacific ocean perch may compete with walleye pollock (Theragra chalcogramma) for food because they feed on largely the same organisms *232*. Habitat Associations - 1
                                (DRAFT) - Food Habits
                            Species perch, Pacific Ocean
                                 Species Id M010240
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 232 LIFESTAGE FOOD FOOD PART Juvenile Copepods Not Specified Juvenile Crustaceans Not Specified General Crustaceans Not Specified General Osteichthyes Not Specified Juvenile See Comments; Food See Comments General See Comments; Food See Comments REFERENCES FOR GENERAL FOOD - 232 REFERENCES FOR JUVENILE FOOD - 232 COMMENTS ON FOOD - Juvenile perch of all sizes in Southeast Alaska feed on copepods and euphausiids. Stocks in the Gulf of Alaska fed almost entirely on euphausiids, whereas those in the Bering Sea consume fishes, euphausiids,and other crustaceans *232*. Types of Feeding Area Used: Schools of feeding perch are found mainly at depths of 150 to 200 m. Dense concentrations of feeding perch were found in the western Gulf of Alaska southeast of Kodiak Island, southwest of Shumagin Islands, and south of Unimak Island *232*. Factors Limiting Availability of Food: Pacific ocean perch may compete with walleye pollock (Theragra chalcogramma) for food. Rapid growth and survival of larval rockfish, as with other ocean fish (such as pollock and Pacific halibut), is probably dependent on ocean currents and weather conditions that result in a concentration of available food *232*. Feeding Behavior: Perch in the Gulf of Alaska feed heavily from May to September and hardly at all through the rest of the year. The feeding rate changes during the day, being most intensive at noon and least intensive in the morning *232*. COMMENTS ON JUVENILE FOOD - Juvenile perch of all sizes in Southeast Alaska feed on copepods and euphausiids *232*. Food Habits - 1
                         (DRAFT) - Environment Associations
                            Species perch, Pacific Ocean
                                 Species Id M010240
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Depth Preference: 500-1000 ft. G Water Depth Preference: Specified in Comments E Bottom Type [Aquatic]: Specified in Comments BA Bottom Type [Aquatic]: Specified in Comments E Coastal Features: Specified in Comments BA Coastal Features: Specified in Comments J Bottom Type [Aquatic]: Specified in Comments A Bottom Type [Aquatic]: Specified in Comments A Water Temperature: Below 15 degrees C A Water Temperature: Specified in Comments J Water Temperature: Below 15 degrees C J Water Temperature: Specified in Comments G Dissolved Oxygen: Specified in Comments FA Water Depth Preference: 200-500 ft. FA Water Depth Preference: 500-1000 ft. FA Water Depth Preference: Specified in Comments BA Water Temperature: Specified in Comments BA Water Depth Preference: 1000-1500 ft. BA Water Depth Preference: Specified in Comments J Water Depth Preference: 200-500 ft. J Water Depth Preference: Specified in Comments A Water Depth Preference: 1000-1500 ft. A Water Depth Preference: Specified in Comments A Water Depth Preference: 500-1000 ft. A Water Depth Preference: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR JUVENILE ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR EGG ENVIRONMENTAL ASSOC_ - 232 Environment Associations - 1 (DRAFT) - Environment Associations Species perch, Pacific Ocean Species Id M010240 Date 26 AUG 96 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Pacific ocean perch were found in each region where sampling occurred but were generally restricted to outer shelf and upper slope depth zones. Adult Pacific ocean perch are generally found on the continental slope at depths of 200 to 300 m. Large concentrations have been found around submarine canyons. Nearshore, rocky-bottom coastal areas exposed to open sea conditions and adjacent bays and straits are probably nursery areas for Pacific ocean perch. Juveniles inhabit areas where cover and protection are afforded by cracks and crevices in and under rocks and ledges and among sessile invertebrates such as the anemone (Metridium senile). Both juveniles and adults were found only over areas with a hard or firm substrate, never over muddy substrate. These areas of clean substrate may be caused by ocean currents and that current, rather than substrate type, was a controlling factor in Pacific ocean perch habitat. Distribution of adults may be determined more by food and hydrographic conditions than by substrate. Water temperature is an important environmental factor controlling distribution of perch. Adult rockfish live within 4.0 to 6.5 oC and the young (14 to 26 cm) at lower temperatures (2.5 to 3.5 oC). The vertical distribution of perch in the Gulf of Alaska has been related to the depth of the layer of oxygen deficiency (where O2 content is less than 1 ml/l). Rockfish concentrations throughout the year remain above this layer, which varies from 350 m deep in summer to 420 m in winter *232*. COMMENTS ON ADULT ENVIRONMENTAL ASSOC_ - Adult Pacific ocean perch are generally found on the continental slope at depths of 200 to 300 m. Large concentrations have been found around submarine canyons. Adults were found only over areas with a hard or firm substrate, never over muddy substrate. These areas of clean substrate may be caused by ocean currents and that current, rather than substrate type, was a controlling factor in Pacific ocean perch habitat. Distribution of adults may be determined more by food and hydrographic conditions than by substrate. Water temperature is an important environmental factor controlling distribution of perch. Adult rockfish live within 4.0 to 6.5 oC *232*. When perch become sexually mature, the perch move into deeper waters (up to 320-370 m or deeper) *232*. COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - Schools of feeding perch are found mainly at depths of 150 to 200 m *232*. COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - The location of spawning relates to water temperature, with females spawning in the warmer areas of the gulf. Spawning in the Bering Sea occurs at depths of 360 to 420 m *232* Nearshore, rocky-bottom coastal areas exposed to open sea conditions and adjacent bays and straits are probably nursery areas *232*. COMMENTS ON JUVENILE ENVIRONMENTAL ASSOC_ - Juveniles inhabit areas where cover and protection are afforded by cracks and crevices in and under rocks and ledges and among sessile invertebrates such as the anemone (Metridium senile). Juveniles were found only over Environment Associations - 2 (DRAFT) - Environment Associations Species perch, Pacific Ocean Species Id M010240 Date 26 AUG 96 areas with a hard or firm substrate, never over muddy substrate. These areas of clean substrate may be caused by ocean currents and that current, rather than substrate type, was a controlling factor in Pacific ocean perch habitat. The young (14 to 26 cm) live at lower temperatures (2.5 to 3.5 oC) *232*. Sometime during their first year of life, the juvenile perch become demersal and are found near the ocean bottom in areas 110 to 140 m deep *232*. COMMENTS ON EGG ENVIRONMENTAL ASSOC_ - Nearshore, rocky-bottom coastal areas exposed to open sea conditions and adjacent bays and straits are probably nursery areas *232*. Environment Associations - 3
                               (DRAFT) - Life History
                            Species perch, Pacific Ocean
                                 Species Id M010240
                                   Date 26 AUG 96



LIFE HISTORY

Reproductive Habitat: Spawning occurs in the northern Gulf of Alaska, with densities of larvae being highest in the Yakutat area. High larval densities are also found in the Kodiak Island area. In the Bering Sea, spawning takes place south and southeast of the Pribilof Islands. A study related the location of spawning to water temperature, with females spawning in the warmer areas of the gulf. Spawning in the Bering Sea occurs at depths of 360 to 420 m *232*. Reproductive Seasonality: Pacific ocean perch are ovoviviparous, meaning they are internally fertilized and give birth to live young. Copulation takes place October through February, and spawning (release of young) takes place from March through June. Significant concentrations of larvae have been reported in the Yakutat region during April and May *232*. Reproductive Behavior: Mating has not been observed; however, it has been suggested that males may copulate several times with different females. A study stated mating and fertilization take place simultaneously; however, another study stated that fertilization may occur two months after mating *232*. Age at Sexual Maturity: Fish from all stocks begin to mature at age 5, and all individuals are mature at age 9. Fifty percent of the stock is mature at age 7 *232*. Frequency of Breeding: Pacific ocean perch breed annually *232*. Fecundity: Fecundity is higher in Bering Sea stocks (75,000 eggs at age 15, 205,000 at age 20) than in the Gulf of Alaska (33,000 at age 15, 48,000 at age 20). It is reported that 10,000 to 270,000 larvae may be released *232*. Incubation Period: Spawning (release of live young) takes place four to five months after copulation *232*. Movements Associated with Life Functions: Larval Pacific ocean perch are planktonic, with their distribution largely controlled by ocean currents. Sometime during their first year of life, the juvenile perch become demersal and are found near the ocean bottom in areas 110 to 140 m deep. When they become sexually mature, the perch move into deeper waters (up to 320-370 m or deeper). Adult Pacific ocean perch do not migrate long distances. Seasonal movements of perch are largely between deep and shallow bottoms within a limited area *232*. Natural Factors Influencing Populations: A study noted strong fluctuations in year class strength. Extreme success of failure of year classes is apparently characteristic of the species. Female perch in the Gulf of Alaska cast their larvae in places where water currents are conducive of high productivity. Weather and current conditions resulting in dispersal of plankton may have a negative effect on larval pollock survival. It seems possible the perch larvae are similarly affected. Before heavy commercial exploitation began in the early 1960's, Pacific ocean perch were a dominant Life History - 1 (DRAFT) - Life History Species perch, Pacific Ocean Species Id M010240 Date 26 AUG 96 groundfish in the Gulf of Alaska. But perch stocks now have been severely reduced, and possibly as a result of release from competition with perch, pollock have become much more abundant. Pollock and perch feed on largely the same organisms, and it is possible that competition with pollock will prevent perch stocks from recovering even if fishing pressure is relieved *232*. Human-related Factors Influencing Populations: A summary of possible impacts from human-related activities includes: alteration of preferred water temperature, pH, dissolved oxygen, and chemical composition; introduction of water-soluble substances; increase in suspended organic or mineral material; alteration of preferred substrate; reduction of food supply; seismic shockwaves; and human harvest *232*. LIFE HISTORY CODES - Dispersion: Clumped Nest Materials: Gravel Periodicity: Most active in early summer Periodicity: Most active in late summer Breeding/Spawning Season: January Breeding/Spawning Season: February Breeding/Spawning Season: October Breeding/Spawning Season: November Breeding/Spawning Season: December Breeding/Spawning Season: March Breeding Spawning Season: April Breeding/Spawning Season: May Breeding/Spawning Season: June Mating System (Single breeding season): Polygamy (mal Number of Broods/Litters (Reproductive Efforts) Per Ye Average Number of Offspring/Reproductive Effort: 1,00 Average Number of Offspring/Reproductive Effort: Grea Gestation/Incubation Period: 3-4 months Gestation/Incubation Period: 5-6 months REFERENCES FOR LIFE HISTORY- 232 Life History - 2
                           (DRAFT) - Management Practices
                            Species perch, Pacific Ocean
                                 Species Id M010240
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Developing/maintaining suitable salinity Beneficial Developing/maintaining suitable pH Adverse Salinity alteration Beneficial Prohibiting harvest of species being described Adverse Other management practices [specified in comments] Beneficial Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 232 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 232 COMMENTS ON MANAGEMENT PRACTICES - Human-related Factors Influencing Populations: A summary of possible impacts from human-related activities includes: alteration of preferred water temperature, pH, dissolved oxygen, and chemical composition; introduction of water-soluble substances; increase in suspended organic or mineral material; alteration of preferred substrate; reduction of food supply; seismic shockwaves; and human harvest *232*. Pacific ocean perch within the 200-mi limit are managed by the North Pacific Fishery Management Council in their groundfish fishery managements plans *232*. Management Practices - 1
                                   (DRAFT) - References
                               Species perch, Pacific Ocean
                                    Species Id M010240
                                      Date 26 AUG 96



     

References

232 * State of Alaska Department of Fish, and Game. 1986. Alaska Habitat Management Guide: Life Histories and Habitat Requirements of Fish and Wildlife. (ed.). Alaska Department of Fish and Game Juneau, Alaska:763. References - 1