(DRAFT) - Taxonomy
                                  Species sablefish
                                 Species Id M010402
                                   Date 26 AUG 96



TAXONOMY

NAME - sablefish OTHER COMMON NAMES - ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - , FAMILY AND SUBFAMILY - , GENUS AND SUBGENUS - Anoplopoma, SPECIES AND SSP - fimbria, SCIENTIFIC NAME - Anoplopoma fimbria AUTHORITY - Pallas TAXONOMY REFERENCES - 232 Taxonomy - 1
                                  (DRAFT) - Status
                                  Species sablefish
                                 Species Id M010402
                                   Date 26 AUG 96



STATUS

Coded Status Migrant Existing, FMP exists See Comments REFERENCES FOR STATUS - 232 COMMENTS ON STATUS - Years of tagging studies have shown that sablefish conduct extensive migrations *232*. Sablefish withing the 200-mi limit are managed by the North Pacific Fishery Management Council (NPFMC) in their Gulf of Alaska Goundfish Fishery Management Plan. Sablefish within 3 mi of shore are managed by the Alaska Department of Fish and Game (ADF&G) *232*. Status - 1
                               (DRAFT) - Distribution
                                  Species sablefish
                                 Species Id M010402
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE REFERENCES FOR HABITAT - 232 LAND USE - Water REFERENCES FOR LAND USE - 232 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine REFERENCES FOR NWI - 232 COMMENTS ON HABITAT ASSOCIATIONS - Juvenile sablefish occupy shallower, nearshore waters *232*. Sablefish follow a diurnal vertical migration and feed both near the surface and in bottom water layers (down to 1,200 m) *232* Juveniles are pelagic or semipelagic. The move from a pelagic to more demersal existence may take place at around 30 cm *232*. ANIMAL/PLANT SPECIES ASSOCIATIONS - Pacific halibut (Hippoglossus stenolepis) Lingcod (Ophiodon elongatus) Seal Sea lion REFERENCES FOR SPECIES ASSOCIATIONS - 232 COMMENTS ON SPECIES ASSOCIATIONS - The IPHC listed sablefish as a frequent food item of Pacific halibut (Hippoglossus stenolepis). Other large predators such as lingcod (Ophiodon elongatus) probably also consume sablefish, and sablefish eggs, larvae, and juveniles are probably consumed by many more species. Sea lions eat sablefish, and sablefish are sometimes pursued by seals *232*. Habitat Associations - 1
                                (DRAFT) - Food Habits
                                  Species sablefish
                                 Species Id M010402
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 232 LIFESTAGE FOOD FOOD PART Adult Osteichthyes Not Specified Adult Gadiformes Not Specified Adult Pleuronectiformes Not Specified Adult Clupeiformes Not Specified Adult Atheriniformes Not Specified Adult Perciformes Not Specified Adult Invertebrate Cordates Not Specified General Malacostraca Not Specified General Anthozoans Not Specified General Brittle/Serpent Star Not Specified General Crustaceans Not Specified General Osteichthyes Not Specified General Gadiformes Not Specified General Clupeiformes Not Specified General Perciformes Not Specified General Pleuronectiformes Not Specified General Aves Not Specified General Otariidae See Comments General Phocidae See Comments Juvenile Myctophiformes Not Specified Juvenile Atheriniformes Not Specified Juvenile Crustaceans Not Specified Juvenile Salmoniformes Not Specified Adult See Comments; Food See Comments General See Comments; Food See Comments Juvenile See Comments; Food See Comments REFERENCES FOR GENERAL FOOD - 232 REFERENCES FOR ADULT FOOD - 232 REFERENCES FOR JUVENILE FOOD - 232 COMMENTS ON FOOD - Sablefish are opportunistic feeders, consuming a wide variety of organisms. Their diet is dependant upon their life stage, geographic location, the season, and availability of prey. In the Bering Sea, sablefish consume pandalid shrimp (Pandalus spp.), sea anenomes (Actinaria), brittle stars (Ophiuroidea), and small crustaceans (amphipods and euphausids) in addition to several kinds of fish (Saffron cod, Eleginus gracilis; Pacific cod, Gadus macrocephalus; walleye pollock; Pacific herring; sculpins, Cottidae; and small flounders, Pleuronectidae). Occasional presence of bird remnants and seal fur in sablefish stomachs has been noted *232*. Food Habits - 1 (DRAFT) - Food Habits Species sablefish Species Id M010402 Date 26 AUG 96 Types of Feeding Areas Used: Sablefish follow a diurnal vertical migration and feed both near the surface and in bottom water layers (down to 1,200 m) *232*. Factors Limiting Availability of Food: Little specific information is available concerning limitations of food availability for sablefish. A study speculates that sablefish under natural conditions may feed very infrequently and that the absence of stress may reflect an evolutionary adaptation to that feeding strategy *232*. Feeding Behavior: A study noted an annual cycle in the intensity of sablefish feeding. The fullest stomachs were found at the beginning of summer (April-June), with feeding intensity decreasing in autumn (August), and rising again in February-March. Another study observed feeding sablefish and noted that they appeared to be "premediated" feeders --- seeking out selected prey, then leaving and allowing the prey to reconcentrate before feeding again *232*. COMMENTS ON ADULT FOOD - Adult sablefish in the Gulf of Alaska feed on fish, including walleye pollock (Theragra chalcogramma), arrowtooth flounder (Atheresthes stomias), spiny cheek rockfish (Sebastolobus spp.), Pacific herring (Clupea harengus pallasii), Pacific saury (Cololabias saira), and sand lance (Ammodytes hexapterus). They also feed on free-swimming and bottomdwelling invertebrates *232*. COMMENTS ON JUVENILE FOOD - Young sablefish in their pelagic stage off the coast of Oregon and Washington have been reported to feed on blue lanternfish (Tarletonbeania crenularis), saury, and euphausiids. Young sablefish also feed on Pacific saury. Small sablefish (36-38 cm) off southern British Columbia gather in estuaries of rivers, where they feed on young salmon *232*. Food Habits - 2
                         (DRAFT) - Environment Associations
                                  Species sablefish
                                 Species Id M010402
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Depth Preference: 200-500 ft. G Water Depth Preference: 500-1000 ft. G Water Depth Preference: 1000-1500 ft. G Water Depth Preference: Greater than 1500 ft. G Water Depth Preference: Specified in Comments A Water Depth Preference: 200-500 ft. A Water Depth Preference: 500-1000 ft. A Water Depth Preference: 1000-1500 ft. A Water Depth Preference: Greater than 1500 ft. A Water Depth Preference: Specified in Comments J Water Depth Preference: Specified in Comments G Water Temperature: Below 15 degrees C G Water Temperature: Specified in Comments J Water Temperature: Specified in Comments FA Water Depth Preference: Specified in Comments BA Water Depth Preference: 500-1000 ft. BA Water Depth Preference: 1000-1500 ft. BA Water Depth Preference: Greater than 1500 ft. BA Water Depth Preference: Specified in Comments E Water Depth Preference: Specified in Comments J Aquatic Habitat Zonation: Open water [pelagic zone] J Aquatic Habitat Zonation: Specified in Comments J Water Depth Preference: 200-500 ft. J Water Depth Preference: 500-1000 ft. A Water Depth Preference: 200-500 ft. A Water Depth Preference: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR JUVENILE ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR EGG ENVIRONMENTAL ASSOC_ - 232 Environment Associations - 1 (DRAFT) - Environment Associations Species sablefish Species Id M010402 Date 26 AUG 96 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Water Depth: Adult sablefish are found depths from 150 m to 1,500 m. Juvenile sablefish occupy shallower, nearshore waters. Sablefish larger than 340 mm are not found in nearshore commercial catch and index samples, indicating that young sablefish probably do not remain in the nearshore zone beyond their second summer *232*. Water Temperature: Sablefish distribution in the Bering Sea is controlled by temperature, with sablefish found in the relatively warm (3 to 5 oC) continental slope zone. Young sablefish occupy a wider range of temperatures *232*. COMMENTS ON ADULT ENVIRONMENTAL ASSOC_ - Adult sablefish are found depths from 150 m to 1,500 m. As the fish get larger (45-50 cm) they migrate to deeper waters, with adults found in areas with depths greater than 150 m *232*. COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - Sablefish follow a diurnal vertical migration and feed both near the surface and in bottom water layers (down to 1,200 m) *232*. COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - Sablefish breed in deep waters (250 to 750 m) *232*. COMMENTS ON JUVENILE ENVIRONMENTAL ASSOC_ - Juvenile sablefish occupy shallower, nearshore waters. Sablefish larger than 340 mm are not found in nearshore commercial catch and index samples, indicating that young sablefish probably do not remain in the nearshore zone beyond their second summer *232*. Young sablefish occupy a wider range of temperatures *232*. Juveniles are pelagic or semipelagic *232*. Young (45 mm) fish are found in shallow (70 to 200 m), more coastal waters *232*. COMMENTS ON EGG ENVIRONMENTAL ASSOC_ - After spawning, the eggs rise to the surface, where development occurs *232*. Environment Associations - 2
                               (DRAFT) - Life History
                                  Species sablefish
                                 Species Id M010402
                                   Date 26 AUG 96



LIFE HISTORY

Reproductive Habitat: Sablefish breed in deep waters (250 to 750 m). After spawning, the eggs rise to the surface, where development occurs. A large percentage of sablefish in the Gulf of Alaska may spawn in the southeastern gulf. Sablefish in the Bering Sea have been observed to spawn only in the south and southeastern areas, especially in the Bower's Ridge and Aleutian Island regions *232*. Reproductive Seasonality: Spawning in the Bering Sea apparently occurs in February. Sablefish along the entire west coast of Canada spawn from January through April, with peak spawning occurring in February *232*. Age at Sexual Maturity: Males mature sooner than females. The average age at 50% maturity is five years for males and seven years for females *232*. Frequency of Breeding: Sablefish breed annually *232*. Fecundity: Fecundity is related to size, with a small (61 cm) female producing about 82,000 eggs and large (98 cm) female producing 1,277,000 *232*. Incubation Period: No information on the rate of egg development is available, though small (15.9 to 24.8 mm) larvae have been captured in the eastern Bering Sea in July, approximately five months after spawning. Juveniles are pelagic or semipelagic. The move from a pelagic to more demersal existence may take place at around 30 cm *232*. Timing of Movements and Use of Areas: Young (45 mm) fish are found in shallow (70 to 200 m), more coastal waters. As the fish get larger (45-50 cm) they migrate to deeper waters, with adults found in areas with depths greater than 150 m. Sablefish follow a diurnal vertical migration pattern, rising as high as the surface water layer during the day and dropping down to the bottom layers at night *232*. Migration Routes: Years of tagging studies have shown that sablefish conduct extensive migrations. Until recently it was felt that, though some fish did migrate long distances, most migration was localized. New evidence however, indicates that a significant number of fish do migrate long distances (over 185 km). Large fish (over 60 cm) in the Gulf of Alaska tend to migrate eastward, while small (less than 60 mm) fish tend to move westward (possilbe drifting with prevailing ocean currents). The change in direction of movement may be associated with the onset of maturity, with large numbers of adult fish moving to the southeastern gulf to spawn *232*. Natural Factors Influencing Populations: The IPHC listed sablefish as a frequent food item of Pacific halibut (Hippoglossus stenolepis). Other large predators such as lingcod (Ophiodon elongatus) probably also consume sablefish, and sablefish eggs, larvae, and juveniles are probably consumed by many more species. Sea lions eat sablefish, and sablefish are sometimes pursued by seals. As with many other ocean species, the survival of sablefish eggs and larvae is dependant upon beneficial weather patterns and ocean currents that carry them into areas where temperature regimes and food Life History - 1 (DRAFT) - Life History Species sablefish Species Id M010402 Date 26 AUG 96 concentrations are favorable to development *232*. Human-related Factors Influencing Populations: A summary of possible impacts from human-related activities includes: alteration of preferred water temperatures, pH, dissolved oxygen, and chemical composition; introduction of water-soluble substances; increase in suspended organic or mineral material; reduction in food supply; human harvest; and seismic shock waves *232*. LIFE HISTORY CODES - Periodicity: Most active in early spring Periodicity: Most active in late spring Breeding/Spawning Season: January Breeding/Spawning Season: February Breeding/Spawning Season: March Breeding Spawning Season: April Average Number of Offspring/Reproductive Effort: Grea REFERENCES FOR LIFE HISTORY- 232 Life History - 2
                           (DRAFT) - Management Practices
                                  Species sablefish
                                 Species Id M010402
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Developing/maintaining suitable salinity Beneficial Developing/maintaining suitable pH Adverse Salinity alteration Beneficial Regulating harvest of species being described Beneficial Other management practices [specified in comments] Adverse Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 232 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 232 COMMENTS ON MANAGEMENT PRACTICES - Human-related Factors Influencing Populations: A summary of possible impacts from human-related activities includes: alteration of preferred water temperatures, pH, dissolved oxygen, and chemical composition; introduction of water-soluble substances; increase in suspended organic or mineral material; reduction in food supply; human harvest; and seismic shock waves *232*. Management Practices - 1
                                   (DRAFT) - References
                                    Species sablefish
                                    Species Id M010402
                                      Date 26 AUG 96



     

References

232 * State of Alaska Department of Fish, and Game. 1986. Alaska Habitat Management Guide: Life Histories and Habitat Requirements of Fish and Wildlife. (ed.). Alaska Department of Fish and Game Juneau, Alaska:763. References - 1