(DRAFT) - Taxonomy
                               Species grouper, Nassau
                                 Species Id M010412
                                   Date 27 AUG 96



TAXONOMY

NAME - grouper, Nassau OTHER COMMON NAMES - Nassau grouper, grouper, rockfish and hamlet ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Osteichthyes, ORDER AND SUBORDER - Perciformes, FAMILY AND SUBFAMILY - Serranidae, GENUS AND SUBGENUS - Epinephelus, SPECIES AND SSP - striatus, SCIENTIFIC NAME - Epinephelus striatus AUTHORITY - Bloch, 1792 TAXONOMY REFERENCES - 200 Taxonomy - 1
                                  (DRAFT) - Status
                               Species grouper, Nassau
                                 Species Id M010412
                                   Date 27 AUG 96



STATUS

Coded Status Sport Fish REFERENCES FOR STATUS - 200 COMMENTS ON STATUS - Despite their economic importance, available information on the life history of black and Nassau groupers is grossly inadequate for effective management of the fishery. Few studies have been published on the biology of the groupers and are mainly focused on tagging programs that describe movement patterns.*200* Status - 1
                               (DRAFT) - Distribution
                               Species grouper, Nassau
                                 Species Id M010412
                                   Date 27 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine RF. Marine RB. Estuarine AB3 REFERENCES FOR NWI - 200 COMMENTS ON HABITAT ASSOCIATIONS - Like most species of groupers, the black, red and Nassau groupers are secretive, occupying caves, ledges and crevices on reefs and shipwrecks. The larger fish generally occupy the lower part of the depth range. The Nassau grouper is generally found near high-relief coral reefs and rocky bottoms from the shoreline to a depth of at least 90m. It is reported that large fish occur at depths greater than about 50 m. In Bermuda, there is a marked size segregation; fish less than 40 cm SL were on inshore banks and larger ones were on offshore banks. *200* Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species grouper, Nassau
                                 Species Id M010412
                                   Date 27 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 200 LIFESTAGE FOOD FOOD PART General Crustaceans Not Specified General Squid, Octopus Not Specified General Fish Not Specified Juvenile Plankton Not Applicable REFERENCES FOR GENERAL FOOD - 200 REFERENCES FOR JUVENILE FOOD - 202 COMMENTS ON FOOD - All the groupers are carnivorous, feeding mainly on fishes, larger crustaceans, and occasionally on cephalopods.*202* Nassau groupers may consume queen conchs after other predators have made soft parts available. In Bahamian waters, however, Nassau groupers have been observed forcefully pulling conchs out of their shells. Moray eels have been reported in stomachs of Nassau groupers collected in the Virgin Islands and Puerto Rico and Jamaica.*200* COMMENTS ON ADULT FOOD - Stomach contents of 150 Nassau groupers collected during daylight off the Virgin Islands and Puerto Rico consisted of about 53% fish, 39% crustaceans, 5% cephalopods, 2% gastropods, and 2% pelecypods.*200* COMMENTS ON JUVENILE FOOD - Many studies report some change in proportion of various food items-- eg., consumption of more fish and less crustaceans with increasing age. There is evidence that snappers and groupers eat other snappers and groupers. The prey are usually rather young stages, but predation on adults does occur.*202* Crustaceans dominated the diet of smaller fish, whereas fish predominated in the stomach contents of larger Nassau groupers over 300 mm SL.*200* Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species grouper, Nassau
                                 Species Id M010412
                                   Date 27 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Substrate: Rocks G A Water Depth Preference: 50-100 ft. G Biodegradable Organics: Wide range of tolerance to organic pollution FJ Bottom Type [Aquatic]: Rooted aquatic vegetation REFERENCES FOR ENVIRONMENTAL ASSOC_ - 200 REFERENCES FOR RESTING ADULT ENVIRONMENTAL ASSOC_ - 200 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - The Nassau grouper is generally found near high-relief coral reefs and rocky bottoms from the shoreline to a depth of at least 90 m. Large fish occur at depths greater than about 50 m. In Bermuda, a marked size segregation was noted: fish less than 40 cm SL were on inshore banks. The species may be resistant to changing salinities. In the old New York Aquarium, several Nassau groupers reportedly lived for several years in water that occasionally became almost fresh and was rather polluted.*200* Environment Associations - 1
                               (DRAFT) - Life History
                               Species grouper, Nassau
                                 Species Id M010412
                                   Date 27 AUG 96



LIFE HISTORY

Morphology: This grouper can be distiguished from other species in the genus Epinephelus by the third spine of the dorsal fin which is longer than the second, and in having a slightly indented interspinous membrane. The caudal fin is slightly emarginated. In Nassau groupers the pelvic fins are shorter than the pectorals and are inserted below or behind the ventral end of the pectoral fin base. The Nassau grouper can be distinguished from Mycteroperca spp. by its less elongated body and eight soft anal fin rays. Other serranid species have less robust bodies, soft dorsal and anal fin bases that are not covered by scales or thick skin, and 10 or fewer dorsal fin spines. Coloration: In this species the body ground color can be tawny to pinkish and red, with five dark vertical bars; the third and fourth bars divide above the lateral line and their adjacent branches join to form a w-shaped mark. The Nassau and red groupers are somewhat similar in general appearance, but can be distinguished because the Nassau grouper has a black saddle on top of the caudal peduncle, distincitve black spots below and behind the eyes, and a characteristic "tuning fork" shaped mark on the top of the head, all of which are lacking in the red grouper.*200* Sex reversal: The Nassau grouper changes from female to male at a length between 300 and 800 mm, which presumably indicated that an individual fish can spend more than one spawning season as a functional female. In the Caribbean, ripe fish were collected between February and May, and sexually inactive ones were collected in October and November. Nassau groupers form spawning aggregations of as many as 100,000 fish off Bimini, Bahamas, for 1 week during the full moon in January. Similar aggregations have been found in other Carribean areas.*200* Movement: Nassau groupers have been reported to form enormous spawning aggregations and undergo spawning migrations. Analyses of gonad samples indicates that fish of both sexes are present and ripe during the aggregations. Tagged Nassau groupers in the Florida Keys promptly returned to their original reef, showing strong home-reef specificity. Tagged Nassau groupers in the Virgin Islands moved at most about 820 m from the site of release. From results of tagging studies, it was reported that the Nassau grouper tends to become established in an area.*200* REFERENCES FOR LIFE HISTORY- 200 Life History - 1
                           (DRAFT) - Management Practices
                               Species grouper, Nassau
                                 Species Id M010412
                                   Date 27 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Providing artificial nesting/spawning sites Beneficial Regulating harvest - setting size limits Existing Regulating commercial harvest gear types REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 200 REFERENCES FOR EXISTING MANAGEMENT PRACTICES - 200 COMMENTS ON MANAGEMENT PRACTICES - The minimum size for any species (including species such as scamp and gag) is 18 inches total length. New gear restrictions prohibit use of longline nets and stab nets (also called sink nets) off the Atlantic coast of Florida. Artificial reef programs designed to improve recreational fishing in the ocean have become popular in recent years; most activity in United States waters has been in Florida. Artificial reefs consisting of a wide variety of materials, ranging from abandoned vessels to designed and prefabricated structures and even junk (debris, large appliances, etc.), have been placed on the coasts of Florida (greatest activity has been in Dade County). A controversy exists as to the best depths for artificial reefs. Among the recreational fishermen, there seems to be a consensus favoring water about 73 m deep, where diving and spearfishing are discouraged and reefs tend to attract large fish. The long-term effect of artificial reef placement would seem to be that recreational catches of groupers would improve, perhaps in some proportion to the numbers and kinds of artificial reefs installed. It is not known if artificial reefs increase fish populations or just redistribute them.*200* Size limits that maximize yield per recruit for gonochoristic (bisexual) species would probably allow continued adequate reproduction of that species regardless of fishing mortality if release-related mortality is low. Protection of reproduction occurs because the size limit that maximizes yield per recruit appears mathematically related to the downward inflection of the von Bertalanffy growth curve. The inflection usually occurs at the initial age of sexual maturity when the fish's energy is partially diverted from somatic growth to reproduction. Thus, size limits that maximize yield per recruit usually allow attainment of sexual maturity.*202* Management Practices - 1
                                   (DRAFT) - References
                                 Species grouper, Nassau
                                    Species Id M010412
                                      Date 27 AUG 96



     

References

200 * Jory, Darryl, Iverson, Edwin. 1989. Species Profiles: Life Histories and Environmental Requirements of Coastal Fishes and Invertebrates (South Florida)--Black, Red and Nassau Groupers. Fish and Wildlife Species Profiles 82(11.110) (ed.). Fish and 202 * Polovina, Jeffrey (Ed.) and Stephen Ralston (Ed.). 1987. Tropical Snapper and Groupers: Biology and Fisheries Management. Polovina, J. and Ralston, S. (ed.). Westview Press Boulder, CO and London, England:659. 191 * South Atlantic Fishery Management Council. 1988. Amendment Number 1 and Environmental Assessment and Regulatory Impact Review To The Fishery Management Plan For The Snapper Grouper Fishery of The South Atlantic Region. (ed.):46. References - 1