(DRAFT) - Taxonomy
                                  Species monkfish
                                 Species Id M010508
                                   Date 26 AUG 96



TAXONOMY

NAME - monkfish OTHER COMMON NAMES - monkfish, goosefish, American angler, American goosefish, all-mouth, fishing frog, baudroie, diable de mer, poisson-pecheur and Fr. baudroie d'Amerique ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - Lophiiformes, FAMILY AND SUBFAMILY - Lophiidae, GENUS AND SUBGENUS - Lophius, SPECIES AND SSP - americanus, SCIENTIFIC NAME - Lophius americanus AUTHORITY - Valenciennes, 1837 TAXONOMY REFERENCES - 258 COMMENTS ON TAXONOMY - Lophius americanus, considered at one time synonymous with Lophius picatorius of the eastern Atlantic, was shown to be a distinct species by Berrill (1929).*258* Family Lophiidae or goosefishes, is a small family of marine fishes occurring, generally worldwide, in Arctic, Atlantic, Pacific, and Indian oceans, some inhabiting warm regions; mainly deepwater fishes, benthic and epipelagic; on muddy bottoms of the continental slope, usually beyond 200 m. Body much depressed and broad anteriorly, with a slender caudal peduncle. Head large and depressed with numerous conical tubercles and ridges on dorsal and lateral surfaces; snout with prominent spines; mouth wide, with numerous sharp, depressible teeth in both jaws; eye relatively small. Gill opening usually at or behind, rarely extending in front of, pectoral fin base; no gill rakers.*258* Taxonomy - 1
                                  (DRAFT) - Status
                                  Species monkfish
                                 Species Id M010508
                                   Date 26 AUG 96



STATUS

Coded Status See Comments REFERENCES FOR STATUS - 258 COMMENTS ON STATUS - There is no directed fishery for monkfish in Canadian Atlantic waters. When captured it is usually taken on longline or by trawl. Catches from NAFO subareas 3 and 4 totalled 266 t in 1982; total catch from all NAFO subareas for that year was 3204 t. Most of the Canadian catch is used in the manufacture of fish meal. When marketed fresh or frozen, the flesh of the caudal section or tail is said to have a good flavor, and is considered a delicacy by some. The species is used extensively in physiological and biochemical studies.*258* Status - 1
                               (DRAFT) - Distribution
                                  Species monkfish
                                 Species Id M010508
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE REFERENCES FOR HABITAT - 258 LAND USE - Water REFERENCES FOR LAND USE - 258 COMMENTS ON HABITAT ASSOCIATIONS - The monkfish is a bottom-dwelling sluggish fish, living over a wide variety of substrates from tideline down to 668 m. Monkfish were found to be associated with fine gravel and clay sediment-type bottom on the Scotian Shelf. It tolerates a wide range of temperature, 0-21 deg C. Although widely distributed over the Scotian Shelf at all depths and along the Laurentian Channel, it appeared to avoid cold surface waters of the Browns-LaHave area and north of Banquereau. Preferred depth ranges were given as 73-90 and 128-144 m, preferred temperatures 6-10 deg, and salinities 33-24 ppt. Studies of seasonal distribution indicated that monkfish invade shallow waters of the banks in summer and migrate to deeper waters in winter.*258* COMMENTS ON SPECIES ASSOCIATIONS - Predation and competition: There is little information on predation. Monkfish have been eaten by swordfish. The young are probably eaten by predaceous fishes, and in the laboratory the larvae have been observed to have been eaten by crustaceans.*258* Parasites and disease: Species of parasites includes one species of protozoan, six species of trematodes, and the larvae of the nematode Phocanema sp. Immature monkfish were infected by the protozoan Haemogregarina sp. in the Gulf of Maine.*258* Habitat Associations - 1
                                (DRAFT) - Food Habits
                                  Species monkfish
                                 Species Id M010508
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 258 LIFESTAGE FOOD FOOD PART General Fish Not Specified General Crustaceans Not Specified General Molluscs Not Specified General Squid, Octopus Not Specified General Worms Not Specified General Osteichthyes Not Specified General Clupeiformes Not Specified General Perciformes Not Specified General Pleuronectiformes Not Specified General Chondrichthyes Not Specified General See Comments; Food Adult See Comments; Food REFERENCES FOR GENERAL FOOD - 258 REFERENCES FOR ADULT FOOD - 258 COMMENTS ON FOOD - A voracious predator, the monkfish consumes a wide variety of foods and has been reported to take any available prey, occasionally items that have proved indigestible. Stomach contents examined included such fishes as herring, sand lance, alewives, menhaden, smelt, cod, haddock, mackerel, cunner, striped bass, sculpins, sea ravens, flounders, and skates. Many invertebrates including crabs, squid, other molluscs, starfish, and marine worms are eaten, and even seabirds have been found in stomachs.*258* COMMENTS ON ADULT FOOD - A voracious predator, the monkfish consumes a wide variety of foods and has been reported to take any available prey, occasionally items that have proved indigestible. Stomach contents examined included such fishes as herring, sand lance, alewives, menhaden, smelt, cod, haddock, mackerel, cunner, striped bass, sculpins, sea ravens, flounders, and skates. Many invertebrates including crabs, squid, other molluscs, starfish, and marine worms are eaten, and even seabirds have been found in stomachs. The adult monkfish feeds mainly on bottom organisms and has been observed to partially bury itself in the bottom substrate awaiting prey. The fishing 'lure' on the head is used as an aid in the attraction and ultimate capture of food items.*258* Food Habits - 1
                         (DRAFT) - Environment Associations
                                  Species monkfish
                                 Species Id M010508
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Bottom Type [Aquatic]: Specified in Comments G Water Depth Preference: Specified in Comments G Water Temperature: Below 15 degrees C G REFERENCES FOR ENVIRONMENTAL ASSOC_ - 258 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - The monkfish is a bottom-dwelling sluggish fish, living over a wide variety of substrates from tideline down to 668 m. Monkfish were found to be associated with fine gravel and clay sediment-type bottom on the Scotian Shelf. It tolerates a wide range of temperature, 0-21 deg C. Although widely distributed over the Scotian Shelf at all depths and along the Laurentian Channel, it appeared to avoid cold surface waters of the Browns-LaHave area and north of Banquereau. Preferred depth ranges were given as 73-90 and 128-144 m, preferred temperatures 6-10 deg, and salinities 33-24 ppt. Studies of seasonal distribution indicated that monkfish invade shallow waters of the banks in summer and migrate to deeper waters in winter.*258* Environment Associations - 1
                               (DRAFT) - Life History
                                  Species monkfish
                                 Species Id M010508
                                   Date 26 AUG 96



LIFE HISTORY

Description: Body strongly depressed, greatest width occurring at level of pectoral fin, body tapering to small, cylindrical caudal peduncle. Head very large, much depressed, wider than body, rounded, numerous conical tubercles on top and sides of head, frontal ridges rough; humeral spines long, straight, well developed; mouth very large, terminal, lower jaw projecting considerably beyond upper; a row of large, irregular, canine-like teeth in jaws, mostly depressible, strong teeth on vomer and palatines; eye small, on top of head. Branchiostegals 6. Fins; dorsals, 2, first dorsal of 3 isolated spines on head, the foremost modified into angling apparatus, bearing at its end a fleshy tab, second spines depressible, third spine shorter, a group of short, slender spines behind head, connected by black membrane; second dorsal of 9-12 rays; caudal moderate, thick truncate, broom-shaped; anal 8-10 rays; pelvics thickened, jugular, below head, 1 spine, 5 soft rays, well in front of pectorals; pectorals 19-28 ryas, heavy bases somewhat arm-like. Scales absent, skin smooth, slippery, a series of branched, fleshy fringes or tabs along edge of lower jaw, extending back on sides of head almost to base of pectorals; 2 rows of similar, but smaller tabs from above and behind pectoral fins to end of caudal peduncle; a few tabs on pectoral fins. Vertebrae 26-31. Size to 120 cm long and weight of 22.7-27.2 kg, but average weight 2.3-4.5 kg. Color uniformly dark brown above, light tan below; pectoral fins becoming darker at tips; pelvic fins reddish.*258* Predation and competition: There is little information on predation. Monkfish have been eaten by swordfish. The young are probably eaten by predaceous fishes, and in the laboratory the larvae have been observed to have been eaten by crustaceans.*258* Parasites and disease: Species of parasites includes one species of protozoan, six species of trematodes, and the larvae of the nematode Phocanema sp. Immature monkfish were infected by the protozoan Haemogregarina sp. in the Gulf of Maine.*258* Reproduction: Spawning occurs from June to September in Canadian waters, some presumably in shoal waters off Nova Scotia. Eggs are pelagic, spherical, and 1.61-1.84 mm in diameter. A single oil globule, measuring 0.40-0.61 mm in diameter, is present. Eggs are deposited as a mucous sheet or veil that floats at the surface and may be 9.1-12.2 m long, 30 cm or more in width, and in a single layer. The sheet, pinkish in color, may contain up to 1.32 million eggs, the product of one female. Egg development was noted at temperatures as low as 5 deg C but estimated time to hatching would be about 100 d, whereas at 15 deg hatching occurred in about 6 or 7 d. Upper limit for development was 17-18 deg C and probably higher. Growth: Larvae are 2.5-4.5 mm long on hatching and float on the surface. Soon after hatching curiously elongated dorsal head spines and pelvic fins develop and pigmentation occurs. Postlarvae apparently descend to the bottom, where they seek protection among algae-covered rocks. Monkfish may reach a length of 5.9 cm before their first winter and 11.4 cm when a little more than a year old. Otolith studies indicated that 76.2 cm monkfish were 9 yr. old and 101.6 cm ones 12 yr old. Life History - 1 (DRAFT) - Life History Species monkfish Species Id M010508 Date 26 AUG 96 LIFE HISTORY CODES - Breeding/Spawning Season: June Breeding/Spawning Season: July Breeding/Spawning Season: August Breeding/Spawning Season: September Spawning Site: Standing Water Average Number of Offspring/Reproductive Effort: Grea Dispersion: Other (specify in comments) REFERENCES FOR LIFE HISTORY- 258 Life History - 2
                           (DRAFT) - Management Practices
                                  Species monkfish
                                 Species Id M010508
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                     Species monkfish
                                    Species Id M010508
                                      Date 26 AUG 96



     

References

References - 1