(DRAFT) - Taxonomy
                               Species cod, Greenland
                                 Species Id M010592
                                   Date 27 AUG 96



TAXONOMY

NAME - cod, Greenland OTHER COMMON NAMES - Greenland cod, rock cod, ogac, uvac, pilot and Fr. ogac ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - Gadiformes, FAMILY AND SUBFAMILY - Gadidae, GENUS AND SUBGENUS - Gadus, SPECIES AND SSP - ogac, SCIENTIFIC NAME - Gadus ogac AUTHORITY - Richardson, 1836 TAXONOMY REFERENCES - 258 COMMENTS ON TAXONOMY - In general, subspecies of Atlantic cod are not acknowledged. Although Atlantic cod vary greatly in many morphological and meristic characters, such as growth and vertebral numbers, these differences are, in part at least, a reflection of environmental conditions.*258* Considered a subspecies, Gadus morhua ogac, by Svetovidov (1948), an opinion since abandoned. Validity of Gadus ogac as full species distinct from G. morhua is well established; among other characteristics G. ogac has a larger eye, narrower caudal peduncle and dark peritoneum.*258* In the family of codfish, medium to large-sized bottom fishes, living in cool seas, except for one freshwater holarctic genus, Lota. Found mainly in the northern hemisphere. Most species are bottom fishes inhabiting inshore waters and continental shelves, some to depths of 1300 m. Cods are generally heavy-bodied, elongate, usually with a stout caudal peduncle. Head large: mouth large; teeth usually well developed on upper (premaxillary) and lower jaws and on head of vomer; eye moderate. Gill rakers usually short and stubby. Branchiostegals 6-8. Fins are well developed. Scales cycloid, small. Lateral line well developed. Swim bladder without pneumatic duct (physoclistous). Vertebrae 42-82. Color generally somber, ranging from tan to brown to silvery. Predaceous fishes feeding heavily on other fishes and invertebrates. The cods are of great economic importance and support extensive commercial fisheries.*258* Taxonomy - 1
                                  (DRAFT) - Status
                               Species cod, Greenland
                                 Species Id M010592
                                   Date 27 AUG 96



STATUS

Coded Status Commercial/consumption REFERENCES FOR STATUS - 258 COMMENTS ON STATUS - Of minor commercial importance. From time to time attempts have been made to establish a market for Greenland cod. In 1979-81 a fishery was operated from Newfoundland with only limited success. The wholesome flesh is whitish and flaky but firmer and tougher and less desirable than that of Atlantic cod. It was noted that those caught in the Miramichi estuary were not esteemed as food.*258* Status - 1
                               (DRAFT) - Distribution
                               Species cod, Greenland
                                 Species Id M010592
                                   Date 27 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

COMMENTS ON SPECIES ASSOCIATIONS - Predation and competition: Its interaction with other species is not well documented. It is probably eaten by seals, but in stomach contents is not usually distinguished from Atlantic cod.*258* Parasites and disease: The Greenland cod from Atlantic waters is hist to the sealworm Phocanema decipiens in flesh and the copepod Lernaeocera branchialis on gills. The cestode Pyramiocephalus phocarum was recorded from the intestine of Greenland cod from Hudson Bay.*258* Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species cod, Greenland
                                 Species Id M010592
                                   Date 27 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 258 LIFESTAGE FOOD FOOD PART General Osteichthyes Not Specified General Salmoniformes Not Specified General Gadiformes Not Specified General Pleuronectiformes Not Specified General Crustaceans Not Specified General Malacostraca Not Specified General Molluscs Not Specified General Polychaetes Not Specified REFERENCES FOR GENERAL FOOD - 258 COMMENTS ON FOOD - Little attention has been given to the food of Greenland cod in Canadian waters. In Greenland waters it eats mainly fishes, especially capelin, polar cod, smaller Greenland cod, and Greenland halibut; among the invertebrates, amphipods, shrimps, crabs, molluscs, and polychate works are commonly eaten.*258* Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species cod, Greenland
                                 Species Id M010592
                                   Date 27 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Temperature: Specified in Comments G Human Association: Specified in Comments G REFERENCES FOR ENVIRONMENTAL ASSOC_ - 258 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Greenland cod inhabit cold temperate to arctic waters, usually in inshore regions in the northern part of its range. Along the Labrador coast, it is the common codfish in harbors and fjords, occurring less commonly in offshore waters. The occurrence of this species in the Bras d'Or Lake further emphasizes its tolerance of reduced salinity.*258* COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - It has been suggested that spawning occurs in brackish waters along the shores of Greenland.*258* Environment Associations - 1
                               (DRAFT) - Life History
                               Species cod, Greenland
                                 Species Id M010592
                                   Date 27 AUG 96



LIFE HISTORY

Description: Body heavy, elongate, compressed posteriorly. Head large; snout blunt; mouth large, terminal but lower jaw slightly overhung by snout; teeth small, numerous on jaws and vomers; distinct and stout barbel near tip of lower jaw; eye moderate. Gill rakers 43. Branchiostegals 7. Fins; dorsals, 3, first of 13-17 rays, second 14-20, third 15-20, all distincly separate, total dorsal rays 45-53; caudal square to only slightly concave, total caudal rays 47-51; anals, 2, first of 18-23 rays, second 15-20; pelvics located in front of pectorals, anterior rays slightly extended and filamentous; pectorals high on sides, fan-like. Scales small, cycloid. Lateral line distinct, arched over pectoral fin, complete, not distinctly colored. Peritoneum uniformly brownish black to black. Vertebrae 53-57. Size averages about 1.3-3.6 kg. Color variable, generally dark brownish black above with yellowish to bronze marbling on sides; white to gray below; unlike Atlantic cod, lateral line not light in color.*258* Growth: The Greenland cod is considered not to live as long as the Atlantic cod, and certainly not to grow as large. Little work on its growth in Canadian waters has been published. In Greenland waters it attains sexual maturity at 3 or 4 yr, a length of 70 cm, and a maximum age of 11 yr, although more commonly it attains an age of 8-10 yr.*258* Reproduction: Spawning is said to occur in February or March. It has been suggested that spawning occurs in brackish waters. It was said to spawn along the shores of Greenland. Details on the number of eggs per female are lacking but probably 1-2 million are produced. The egg is relatively large, and without an oil globule.*258* Predation and competition: Its interaction with other species is not well documented. It is probably eaten by seals, but in stomach contents is not usually distinguished from Atlantic cod.*258* Parasites and disease: The Greenland cod from Atlantic waters is hist to the sealworm Phocanema decipiens in flesh and the copepod Lernaeocera branchialis on gills. The cestode Pyramiocephalus phocarum was recorded from the intestine of Greenland cod from Hudson Bay.*258* LIFE HISTORY CODES - Breeding/Spawning Season: February Breeding/Spawning Season: March Average Number of Offspring/Reproductive Effort: Grea REFERENCES FOR LIFE HISTORY- 258 Life History - 1
                           (DRAFT) - Management Practices
                               Species cod, Greenland
                                 Species Id M010592
                                   Date 27 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                  Species cod, Greenland
                                    Species Id M010592
                                      Date 27 AUG 96



     

References

References - 1