(DRAFT) - Taxonomy
                                  Species porbeagle
                                 Species Id M010730
                                   Date 26 AUG 96



TAXONOMY

NAME - porbeagle OTHER COMMON NAMES - Mackerel shark and Fr: maraiche ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - Chondrichthyes, ORDER AND SUBORDER - Squaliformes, FAMILY AND SUBFAMILY - Lamnidae, GENUS AND SUBGENUS - Lamna, SPECIES AND SSP - nasus, SCIENTIFIC NAME - Lamna nasus AUTHORITY - Bonnaterre, 1788 TAXONOMY REFERENCES - 258 Taxonomy - 1
                                  (DRAFT) - Status
                                  Species porbeagle
                                 Species Id M010730
                                   Date 26 AUG 96



STATUS

Coded Status Commercial/consumption Commercial/industrial See Comments REFERENCES FOR STATUS - 258 COMMENTS ON STATUS - There is no directed fishery in Canadian waters at the present time although there was a longline fishery for porbeagle before the discovery of high mercury levels in shark muscles in 1970 and the cessation of the fishery. Elsewhere within their range, porbeagle are caught by longlines and sold fresh and salted for food, the liver for oil, and the carcass for fish meal. Porbeagle caught incidentally in Canadian waters in weirs or gill nets or on longlines are sometimes shipped to United States markets, where they are used for human food and sold in specialty restaurants as shark, mackerel shark, or mako. The flesh is dense and not unlike that of swordfish in texture *258*. Status - 1
                               (DRAFT) - Distribution
                                  Species porbeagle
                                 Species Id M010730
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE REFERENCES FOR HABITAT - 258 LAND USE - Water REFERENCES FOR LAND USE - 258 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine OW0 REFERENCES FOR NWI - 258 COMMENTS ON HABITAT ASSOCIATIONS - A pelagic, epipelagic, or littoral shark usually more common on continental shelves but occurring sometimes well offshore; also known to frequent inshore areas, including harbors. More common in the Canadian region during the spring, summer, and fall but occasionally caught on fishing banks during winter months. It seems to prefer cool rather than cold waters, is usually found in temperatures below 16 oC, and has been caught in temperatures as low as 6 oC in European waters *258*. ANIMAL/PLANT SPECIES ASSOCIATIONS - Phyllobothrium dagnallium Dinobothrium sp. REFERENCES FOR SPECIES ASSOCIATIONS - 258 COMMENTS ON SPECIES ASSOCIATIONS - Predation and competition: The porbeagle appears to have few enemies other than man *258*. Parasites and disease: There are few records of parasites from porbeagle caught in the western North Atlantic. A study found Phyllobothrium dagnallium in stomach, intestine, and spiral valve, and Dinobothrium sp. from specimens caught around the Magdalen Islands in the Gulf of St. Lawrence *258* Habitat Associations - 1
                                (DRAFT) - Food Habits
                                  Species porbeagle
                                 Species Id M010730
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 258 LIFESTAGE FOOD FOOD PART General Clupeiformes Not Specified General Perciformes Not Specified General Gadiformes Not Specified General Squid, Octopus Not Specified General Osteichthyes Not Specified REFERENCES FOR GENERAL FOOD - 258 COMMENTS ON FOOD - The porbeagle is said to feed mainly on small pelagic schooling species such as herring, gaspereau, and mackerels but cod, white hake, red hake, haddock, cusk, and squid are also eaten. Detailed food studies have not been conducted in Canadian Atlantic waters but examination of stomachs of a few individuals in the Bay of Fundy - Gulf of Maine showed that herring, gaspereau, mackerel, redfish, and squid were eaten *258*. Food Habits - 1
                         (DRAFT) - Environment Associations
                                  Species porbeagle
                                 Species Id M010730
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Aquatic Habitat Zonation: Open water [pelagic zone] G Aquatic Habitat Zonation: Specified in Comments G Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] G Coastal Features: Specified in Comments G Human Association: Specified in Comments G Water Temperature: Between 15-21 degrees C G Water Temperature: Below 15 degrees C G Water Temperature: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 258 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - A pelagic, epipelagic, or littoral shark usually more common on continental shelves but occurring sometimes well offshore; also known to frequent inshore areas, including harbors. More common in the Canadian region during the spring, summer, and fall but occasionally caught on fishing banks during winter months. It seems to prefer cool rather than cold waters, is usually found in temperatures below 16 oC, and has been caught in temperatures as low as 6 oC in European waters *258*. Environment Associations - 1
                               (DRAFT) - Life History
                                  Species porbeagle
                                 Species Id M010730
                                   Date 26 AUG 96



LIFE HISTORY

Reproduction: The porbeagle is ovoviviparous, the developing young being retained within the brood chamber (oviduct) of the female. The young cannibalize fertilized eggs in the oviduct and grow large with noticeable expanded abdomens and branchial regions. The young, 60-75 cm long, are born alive, 1-5 pups per female, usually in late summer in northwest Atlantic waters although females with young may be found throughout the year *258*. Growth: There is little information on rate of growth. Porbeagles are thought to reach sexual maturity at about 5 yr or older and to have a life expectancy of about 20-30 yr. Age determinations are considered possible by counting growth rings on vertebral centra. Of 59 specimens caught off Georges and Browns banks and Trinity Ledges, NS, males averaged about 1.5 m TL and females 1.63 m TL. Round weights of 41 specimens ranged from 17 to 156 kg and averaged 75 kg. Dressed weights of 17 individuals averaged 68 percent of round weights. Maximum size to about 3.05 m but usually 1.83-2.44 m. The all-tackle world record by rod and line was of one weighing 210.92 kg caught off Cornwall, England *258*. Description: Body spindle-shaped, rather heavy, greatest depth at dorsal fin; caudal peduncle slender, flattened, with distinct lateral keels and short secondary keels at caudal base; precaudal pits present. Head stout; snout pointed; mouth moderate to large; teeth similar in upper and lower jaws with large central cusp and two lateral cusps, lateral cusps often not evident on specimens less than 1.2 m long, 24-32 tooth rows in upper jaw, 20-26 in lower jaw; eye large; pectoral origin behind fifth gill slit; gill slits large. Fins: dorsals, 2, first relatively large, second quite small, located over anal; caudal nearly symmetrical, ventral lobe well developed and only slightly smaller than dorsal lobe; anal small, same size as second dorsal; pelvics small; pectorals large, twice as long as broad. Porbeagles have heat-regulating mechanisms (heat exchangers) that raise body temperatures 2.7-8.3 oC above ambient temperatures, allowing them to operate more efficiently in cool water. Size possibly to 3.05 m long but usually 1.83-2.44 m. Color dark bluish gray to bluish black above, white tip on lower trailing edge of first dorsal, lower surfaces of pectoral dark on outer half, mottled white and black toward bases; belly white, unblotched *258*. Predation and competition: The porbeagle appears to have few enemies other than man *258*. Parasites and disease: There are few records of parasites from porbeagle caught in the western North Atlantic. A study found Phyllobothrium dagnallium in stomach, intestine, and spiral valve, and Dinobothrium sp. from specimens caught around the Magdalen Islands in the Gulf of St. Lawrence *258* LIFE HISTORY CODES - Average Number of Offspring/Reproductive Effort: 1 Average Number of Offspring/Reproductive Effort: 2 Average Number of Offspring/Reproductive Effort: 3-4 Average Number of Offspring/Reproductive Effort: 5-7 Life History - 1 (DRAFT) - Life History Species porbeagle Species Id M010730 Date 26 AUG 96 LIFE HISTORY CODES - Breeding/Spawning Season: July Breeding/Spawning Season: August Breeding/Spawning Season: September REFERENCES FOR LIFE HISTORY- 258 Life History - 2
                           (DRAFT) - Management Practices
                                  Species porbeagle
                                 Species Id M010730
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                    Species porbeagle
                                    Species Id M010730
                                      Date 26 AUG 96



     

References

258 * Scott, W.B., M.G. Scott. 1988. Atlantic Fishes of Canada. Canadian Bulletin of Fisheries and Aquatic Sciences (219) (ed.). University of Toronto Press Toronto, Canada:731. References - 1