(DRAFT) - Taxonomy
                         Species turtle, Atlantic green sea
                                 Species Id M030072
                                   Date 26 AUG 96



TAXONOMY

NAME - turtle, Atlantic green sea OTHER COMMON NAMES - ELEMENT CODE - 03/10/84 AOU CODE - 04/24/85 09/29/88 CATEGORY - Reptiles PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Reptilia, ORDER AND SUBORDER - Testudinata, FAMILY AND SUBFAMILY - Cheloniidae, GENUS AND SUBGENUS - Chelonia, SPECIES AND SSP - mydas, mydas SCIENTIFIC NAME - Chelonia mydas mydas AUTHORITY - Linnaeus TAXONOMY REFERENCES - 3082 and 1045 COMMENTS ON TAXONOMY - Although there is insufficient taxonomic information to distinguish between stocks, there may be geographically and genetically distinct populations of the green sea turtle. Replacement of extinct populations by transplanting individuals from another population has not succeeded, and even if it were to succeed, the animals would be biologically different. Although the treatment of the species as a single stock may be ill-advised, because of its worldwide distribution, the lack of data indicating discrete stocks, and the difficulties in distinguishing separate stocks, populations and subspecies, the species is necessarily considered a single stock in the Indo-Pacific region and a single stock in the Atlantic Ocean and adjacent seas for purposes of this review.*91* In various parts of its range, the green sea turtle also may be called tortuga verde, greenback turtle, edible turtle, soup turtle, tortue verte, tortuga blanca, tartaruga verde, aruana and krape. It is distinguishable from other sea turtles by its four large plates on each side of its upper shell and one pair of prefrontal scales; shell plates that do not overlap; and paddle shaped limbs that normally have only one claw. The color of the shell in most adult green sea turtles is highly variable, but background color may be light to dark brown, green, buff, black, or olive. The underside is usually white to pale yellow.*91* Taxonomy - 1
                                  (DRAFT) - Status
                         Species turtle, Atlantic green sea
                                 Species Id M030072
                                   Date 26 AUG 96



STATUS

Coded Status E: Federal Endangered Plan approved by Director T: Federal Threatened REFERENCES FOR STATUS - 1045, 828 and 274 COMMENTS ON STATUS - This species is Endangered in Florida and Threatened elsewhere. It is included in the Recovery Plan For Marine Turtles. Causes of declines in stocks include 1) destruction or modification of habitat, 2) overutilization for commercial, scientific or educational purposes, 3) inadequate regulatory mechanisms, 4) disease and or predation, and 5) losses to incidental catch *8850*. Status - 1
                               (DRAFT) - Distribution
                         Species turtle, Atlantic green sea
                                 Species Id M030072
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - 001, 131, 810, CB1 and AO1 References on County Abundance - 001, 131, 810, CB1 and AO1 REFERENCES FOR HYDROLOGIC UNIT CODES - 001, 131, 810, CB1 and AO1 REFERENCES FOR OTHER DISTRIBUTION - H Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - Aquatic REFERENCES FOR HABITAT - 1045 LAND USE - Water Bays and Estuaries Barren Land Beaches REFERENCES FOR LAND USE - 1045 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine, subtidal OW0 Marine, subtidal AB1 Marine, subtidal AB2 Marine, subtidal RF. Marine, intertidal AB2 Marine, intertidal AB1 REFERENCES FOR NWI - 1045 COMMENTS ON HABITAT ASSOCIATIONS - Nesting beaches are distributed widely in tropical and subtropical regions and as far north as Cape Canaveral Florida. They prefer high energy beaches and nesting requires sand deep enough to deposit the eggs below one meter *8850*. ANIMAL/PLANT SPECIES ASSOCIATIONS - Predators of adult turtles include man, killer whales, and sharks. Eggs are eaten by man, raccoons, coatimundi, dogs, pigs, foxes, peccary, lizards, rats, crabs, and birds.*91* Hatchlings are eaten by dogs, hogs, rats, mongoose, cats, lizards, snakes, jackfish, kingfish, snook, barracuda, groupers, rock cod, and sharks.*91* REFERENCES FOR SPECIES ASSOCIATIONS - 91 COMMENTS ON SPECIES ASSOCIATIONS - Predators of adult turtles include man, killer whales, and sharks. Eggs are eaten by man, raccoons, coatimundi, dogs, pigs, foxes, peccary, lizards, rats, crabs, and birds. Hatchlings are eaten by dogs, hogs, rats, mongoose, cats, lizards, snakes, jackfish, kingfish, snook, barracuda, groupers, rock cod, and sharks. The loss of eggs and hatchings to predation is assumed to be very high. Only one to three percent of the hatchlings reach sexual maturity and only about 50 percent of the eggs hatch.*91* POTENTIAL NATURAL VEGETATION - 065 Northern Cordgrass Prairie (Distichlis-Spartina) REFERENCES FOR PNV - 1045 ECOREGION - Southeastern Mixed Forest: Flat Plains Habitat Associations - 1 REFERENCES FOR ECOREGION - 1045 Habitat Associations - 2
                                (DRAFT) - Food Habits
                         Species turtle, Atlantic green sea
                                 Species Id M030072
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - Omnivore REFERENCES FOR TROPHIC LEVEL - 1045 LIFESTAGE FOOD FOOD PART General Plants Not Specified General Chlorophyta Not Specified General Tracheophyta Not Specified General Poaceae Not Specified General Animals Adult stage General Scyphozoans Adult stage General Sponges Adult stage General Molluscs Adult stage General Diplopods Adult stage Larva Animals Adult stage Adult Plants Not Specified Adult Chlorophyta Not Specified Adult Tracheophyta Not Specified Adult Poaceae Not Specified Adult See Comments; Food See Comments REFERENCES FOR GENERAL FOOD - 1045, 1026 and 1027 REFERENCES FOR ADULT FOOD - 1045, 1026 and 1027 REFERENCES FOR LARVAE FOOD - 1026 COMMENTS ON FOOD - Green sea turtles are primarily herbivores that eat sea grasses and algae. Other organisms living on sea grass blades and algae add to the diet.*91* The green turtle has the unique ability among marine turtles to digest plant material. However, hatchlings and yearlings are primarily carn- ivorous and mature specimens eat marine animals, particularly cniderians whenever available *8850*. COMMENTS ON ADULT FOOD - The adults are mainly herbivores, eating mostly algae and marine grasses, including Thalassia, Zostera, Cymodorea, and Halophila *1045,1026,1027*. However, they do occasionally eat jellyfish, mollusks, and crustaceans *1045,1027*. COMMENTS ON JUVENILE FOOD - The hatchlings and yearlings are primarily carnivorous *8850*. Food Habits - 1
                         (DRAFT) - Environment Associations
                         Species turtle, Atlantic green sea
                                 Species Id M030072
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Temperature: Specified in Comments G Bottom Type [Aquatic]: Rooted aquatic vegetation G Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] G Coastal Features: Reefs G Coastal Features: Sand beaches G Coastal Features: Vegetated offshore islands G Density of Aquatic Vegetation: High G Water Depth Preference: 5-10 ft. G Water Depth Preference: 10-25 ft. LIM Gradient: Specified in Comments LIM Water Temperature: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 1045, 1026 and 1027 REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 1045 and 1026 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - This species lives in waters that are above 20 degrees C in the coldest month. A sloping beach is used for nesting *1045*. Environment Associations - 1
                               (DRAFT) - Life History
                         Species turtle, Atlantic green sea
                                 Species Id M030072
                                   Date 26 AUG 96



LIFE HISTORY

Physical description: The shell is broad and heart-shaped and the head small. They have a length from 91-122 cm and weigh 100-200 kg. The smooth, keelless carapace is brown with brown mottling. The plastron is white to yellow and the head is light brown with yellow markings. The flippers are paddle-shaped and each has one claw *1045,1027*. They are characterized by a single pair of prefrontal scales on the head and usually four costal scutes on the smooth carapace *8850*. Reproduction: The breeding season varies with location but probably does not occur in Virginia. The incubation period is 48-70 days, depending on the beach conditions. Nesting occurs at 2, 3, or 4 year intervals. They have 2-7 clutches per season with 75-200 eggs/clutch. The female builds the nest at night on a sloping beach. She digs a large basin with the front flippers, and at the bottom of this digs a smaller egghole with the rear flippers *1045, 1027,8850*. Courtship and nest occur in the vicinity of nesting beaches and possibly enroute to nesting beaches *8850*. Behavior: This species forages in shallow water marine grasses. Nesting is on shallow beaches and copulation occurs in the water near the nesting beach. Basking is on reefs or uninhabited islands *1045*. This species has the unique ability to digest plant material. However, the hatchlings and yearlings are primarily carnivorous and mature specimens eat marine animals, particularly cniderians when they are available. Hatchling dispersal is likely to be a response to wave forces and ocean currents in their subsequent movements rather than migration *8850*. Origin: The origin of this species is native *1045*. Limiting factors: They use uninhabited beaches for nesting *1045,1026,1027* . It is distinguishable from other sea turtles by its four large plates on each side of its upper shell and one pair of prefrontal scales; shell plates that do not overlap; and paddle shaped limbs that normally have only one claw. The color of the shell in most adult green sea turtles is highly variable, but background color may be light to dark brown, green, buff, black, or olive. The underside is usually white to pale yellow.*91* Nesting: The nesting season varies with location. Nesting is reported between May and November in Florida. Most females nesting on a given beach are never seen again. The ones that do return to nest do not return at a universal interval; however, returns to nest have been observed in two, three, or four year cycles. The females deposit between three and seven clutches per season at about 10 to 18 day intervals. Average clutch sizes vary between 81 and 147 eggs that hatch usually within 48-72 days. Hatchlings emerge, mostly at night, travel quickly to the water and swim out to sea in what is called a "swimming frenzy". At this point, they enter their "lost year" period before they begin diving behavior. This period may be spent in areas where currents concentrate debris and floating vegetation such as sargassum. Since the breeding and nesting grounds are often separated by long distances from the foraging area, long distance migrations are required between these sites.*91* LIFE HISTORY CODES - Life History - 1 (DRAFT) - Life History Species turtle, Atlantic green sea Species Id M030072 Date 26 AUG 96 LIFE HISTORY CODES - Breeding/Spawning Season: June Breeding/Spawning Season: July Breeding/Spawning Season: August Breeding/Spawning Season: September Gestation/Incubation Period: 1-2 months Average Number of Offspring/Reproductive Effort: 101- Number of Broods/Litters (Reproductive Efforts) Per Ye Parental Care of Young: No care given young COMMENTS ON LIFE HISTORY - Green turtles are considered the most palatable of all sea turtles and they became a prime source of meat for mariners and settlers of colonial America *8850*. REFERENCES FOR LIFE HISTORY- 1045, 1026, 1027, 8850, 8819, 8818 and 8822 Life History - 2
                           (DRAFT) - Management Practices
                         Species turtle, Atlantic green sea
                                 Species Id M030072
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Regulating harvest of species being described Beneficial Prohibiting harvest of species being described Beneficial Maintaining undisturbed/undeveloped areas Beneficial Maintaining unique or special habitat features [wetlands, caves, Beneficial Establishing/maintaining nesting and escape cover Beneficial Stocking captive-reared wild-strain animals Beneficial Regulating harvest - restricting number of hunters Existing Regulating harvest of species being described Existing Prohibiting harvest of species being described Existing Stocking captive-reared wild-strain animals Adverse Shoreline modification/development Adverse Dredging Adverse Underwater mining Adverse Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 1045 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 91 REFERENCES FOR EXISTING MANAGEMENT PRACTICES - 1045 COMMENTS ON MANAGEMENT PRACTICES - Green sea turtles may be adversely affected by the following activities: 1. Domestic development - including artificial lighting, man-made barriers, rip-rap, jetties, beach cleaning and traffic; 2. Industrial development- thermal discharge, agrobusiness, radioactive waste, insect control, and trace metals; 3. Pollution - including spills of oil and hazardous materials; 4. Dredging and mining; and 5. Predators attracted to human refuse.*91* Other habitat alterations that affect green sea turtles include the introduction of exotic vegetation, by man on nesting beaches and pollution of the turtles' oceanic habitat. Exotic vegetation may inhibit nesting by forming barriers and dense root mats. The currents which accumulate sargassum weed, where some green sea turtles may spend the early part of their cycle, also accumulate pollutants such as oil, styrofoam, and other plastic. Numerous young green sea turtles have been found dead or moribund along Florida beaches with their jaws and throats obstructed by tar.*91* A complete Recovery Plan is found in reference *8850*. Management Practices - 1
                                   (DRAFT) - References
                            Species turtle, Atlantic green sea
                                    Species Id M030072
                                      Date 26 AUG 96



     

References

828 Service, U.S. Fish and Wildlife. 1983. Republication of the lists of endangered and threatened species. Federal Register 17.11-17.12:1-24. 1026 Bustard, R. 1973. Sea Turtles, Natural History and Conservation. Taplinger Publ. New York, N.Y:22. 1027 Carr, A.F. 1952. Handbook of Turtles. Turtles of the United States, Canada, and Baja California. Comstock Publ. Assoc. Cornell Univ. Press, Ithaca, N.Y:542. 1045 Serv., U.S. Fish and Wildl. 1980. Selected vertebrate endangered species of the sea coast of the United States: Green sea turtle. FWS/OBS-80/1.13. U.S. Dep. Inter., Off. Biol. Serv. Washington, D.C. 8800 Tobey, Franklin J. 1985. Virginia's Amphibians and Reptiles: a Distributional Survey. Virginia Herpetelogical Society:114. 8818 Keinath, J.A., Musick, J.A., Byles, R.A. 1987. Aspects of the biology of Virginia's sea turtles: 1979-1986. Virginia J. Science 38 (4):329-336. 8819 Bellmund, S., Musick, J.A., Klinger, R.E., Byles, R.A., Keinath, J.A., Barnard, D.E. 1987. Ecology of sea turtles in Virginia. VIMS Special Scientific Report 119. VA Inst. Marine Sci., Coll. Wm. and Mary. Gloucester Point, VA:48. 8822 Lutcavage, M., Musick, J.A. 1985. Aspects fo the biology of sea turtles in Virginia. Copeia 1985 2:449-456. 8850 Service , U.S. Fish and Wildlife. 1985. A recovery plan for marine turtles. U.S. Fish and Wildlife Service Atlanta, GA:363. 9286 Virginia Dept. Game Inland Fisheries. 1988. Proceedings of symposium on species of special concern in Virginia. In Press. References - 1