(DRAFT) - Taxonomy
                             Species loon, red-throated
                                 Species Id M040003
                                   Date 26 AUG 96



TAXONOMY

NAME - loon, red-throated OTHER COMMON NAMES - red-throated diver ELEMENT CODE - 08/02/84 AOU CODE - 06/04/85 CATEGORY - Birds PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Aves, ORDER AND SUBORDER - Gaviiformes, FAMILY AND SUBFAMILY - Gaviidae, GENUS AND SUBGENUS - Gavia, SPECIES AND SSP - stellata, SCIENTIFIC NAME - Gavia stellata AUTHORITY - Pontoppidan (1763) TAXONOMY REFERENCES - 624, 3950 and 3944 COMMENTS ON TAXONOMY - AKA red-throated diver *3950,3944* Taxonomy - 1
                                  (DRAFT) - Status
                             Species loon, red-throated
                                 Species Id M040003
                                   Date 26 AUG 96



STATUS

Coded Status Federal Migratory Nongame-Protected Biological Indicator See Comments REFERENCES FOR STATUS - 758, 3944 and 3950 COMMENTS ON STATUS - Considered an important ecological indicator *3944* Status - 1
                               (DRAFT) - Distribution
                             Species loon, red-throated
                                 Species Id M040003
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - 001, 073, 093, 095, 097, 103, 115, 117, 119, 127, 131, 133, 181, 193, 199, 650, 700, 710, 740, 800, 810, 830, CB1, CB2, CB3, AO1 and 999 References on Seasonal Occurrence - Common transient and winter resident along coast *700* References on County Abundance - 001, 073, 103, 115, 119, 131, 133, 193, 199, 650, 700, 710, 810, CB1, CB2, CB3, AO1 and 999 REFERENCES FOR HYDROLOGIC UNIT CODES - 001, 073, 103, 115, 119, 131, 133, 193, 199, 650, 700, 710, 810, CB1, CB2, CB3, AO1 and 999 OTHER DISTRIBUTION REFERENCES FOR OTHER DISTRIBUTION - H Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - Aquatic REFERENCES FOR HABITAT - 624, 1073, 3950 and 3944 LAND USE - Wetland Nonforested Wetland Water Lakes Bays and Estuaries Chesapeake Bay Atlantic Ocean Coastal Waters REFERENCES FOR LAND USE - 700, 3949, 1073 and 3950 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine, subtidal EM1 H Marine, intertidal Estuarine, subtidal Estuarine, intertidal Palustrine Lacustrine, limnetic REFERENCES FOR NWI - 700, 624, 1073, 3944, 3950 and 3948 ANIMAL/PLANT SPECIES ASSOCIATIONS - Diseases and Parasites: General Reference *1901* 1157 1157 1157 REFERENCES FOR SPECIES ASSOCIATIONS - 1901 POTENTIAL NATURAL VEGETATION - 065 Northern Cordgrass Prairie (Distichlis-Spartina) 101 Oak - Hickory - Pine Forest (Quercus-Carya-Pinus) 103 Southern Floodplain Forest (Quercus-Nyssa-Taxodium) REFERENCES FOR PNV - 624 and 700 ECOREGION - Southeastern Mixed Forest: Flat Plains Southeastern Mixed Forest: Irregular Plains REFERENCES FOR ECOREGION - 624 AND 700 Habitat Associations - 1
                                (DRAFT) - Food Habits
                             Species loon, red-throated
                                 Species Id M040003
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - Carnivore REFERENCES FOR TROPHIC LEVEL - 1073 and 3950 LIFESTAGE FOOD FOOD PART General Perciformes Adult stage General Clupeiformes Not Specified General Gasterosteiformes Adult stage General Pleuronectiformes Not Specified General Salmoniformes Not Specified General Osteichthyes Adult stage General Osteichthyes Egg/Fetus stage General Amphibia Adult stage General Malacostraca Not Specified General Crustaceans Not Specified General Molluscs Not Specified General Hirudineans Not Specified General Annelids Not Specified General Insects Not Specified General Animals Adult stage General Animals Egg/Fetus stage Important Osteichthyes Adult stage Important Animals Adult stage Juvenile Insects Not Specified Juvenile Malacostraca Not Specified Juvenile Crustaceans Not Specified Juvenile Perciformes Not Specified Juvenile Clupeiformes Not Specified Juvenile Gasterosteiformes Not Specified Juvenile Pleuronectiformes Not Specified Juvenile Salmoniformes Not Specified Juvenile Osteichthyes Egg/Fetus stage Juvenile Salientia Not Specified Juvenile Amphibia Not Specified Juvenile Molluscs Not Specified Juvenile Hirudineans Not Specified Juvenile Annelids Not Specified Juvenile Animals Egg/Fetus stage Juvenile See Comments; Food See Comments Adult Clupeiformes Adult stage Adult Perciformes Adult stage Adult Pleuronectiformes Not Specified Adult Salmoniformes Not Specified Adult Gasterosteiformes Adult stage Adult Osteichthyes Adult stage Adult Osteichthyes Egg/Fetus stage Adult Amphibia Not Specified Adult Salientia Not Specified Adult Insects Not Specified Adult Malacostraca Not Specified Food Habits - 1 (DRAFT) - Food Habits Species loon, red-throated Species Id M040003 Date 26 AUG 96 LIFESTAGE FOOD FOOD PART Adult Crustaceans Not Specified Adult Molluscs Not Specified Adult Hirudineans Not Specified Adult Annelids Not Specified Adult Animals Adult stage Adult Animals Egg/Fetus stage Adult See Comments; Food See Comments Adult Crustaceans Not Specified Adult Amphibia Not Specified REFERENCES FOR GENERAL FOOD - 3949, 1073, 3950 and 1157 REFERENCES FOR IMPORTANT FOOD - 1157, 1073 and 3950 REFERENCES FOR ADULT FOOD - 3949, 1073, 3950 and 1157 REFERENCES FOR JUVENILE FOOD - 3949, 1073, 3950 and 1157 COMMENTS ON ADULT FOOD - 9999S=feed chiefly on fish *1073,3950* COMMENTS ON JUVENILE FOOD - 9999S=fed crustaceans and aquatic insects 1-4 days after hatching, then small whole fish, after fledging eat diet similar to adult *1073,3950* Food Habits - 2
                         (DRAFT) - Environment Associations
                             Species loon, red-throated
                                 Species Id M040003
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Coastal Wetlands: Coastal open fresh water G Coastal Wetlands: Sounds and bays G Coastal Wetlands: Regularly flooded salt marshes G Inland Wetlands: Bogs G Terrestrial Features: Bare ground G Coastal Wetlands: Coastal shallow fresh marsh G Nutrient [Phosphorus and Nitrogen]: Low nutrient concentrations [oligotroph G Aquatic Features: Lake weed beds G Aquatic/Terrestrial Ecotones: Woodland/water G Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] G Aquatic Habitat Zonation: Dimly-lighted region without emergent vegetation G Terrestrial Features: Specified in Comments LIM Terrestrial Features: Specified in Comments LIM Inland Wetlands: Bogs LIM Coastal Wetlands: Coastal open fresh water LIM Coastal Wetlands: Sounds and bays LIM Coastal Wetlands: Regularly flooded salt marshes LIM Nutrient [Phosphorus and Nitrogen]: Low nutrient concentrations [oligotroph LIM Aquatic Features: Lake weed beds LIM Aquatic/Terrestrial Ecotones: Woodland/water LIM Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] LIM Aquatic Habitat Zonation: Dimly-lighted region without emergent vegetation LIM Aquatic Habitat Zonation: Specified in Comments LIM Coastal Wetlands: Coastal shallow fresh marsh FA Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] FA Aquatic Habitat Zonation: Dimly-lighted region without emergent vegetation FA Nutrient [Phosphorus and Nitrogen]: Low nutrient concentrations [oligotroph FA Coastal Wetlands: Coastal open fresh water BA Terrestrial Features: Specified in Comments BA Inland Wetlands: Bogs BA Coastal Wetlands: Coastal shallow fresh marsh BA Coastal Wetlands: Coastal open fresh water BA Coastal Wetlands: Regularly flooded salt marshes BA Coastal Wetlands: Sounds and bays BA Nutrient [Phosphorus and Nitrogen]: Low nutrient concentrations [oligotroph BA Aquatic/Terrestrial Ecotones: Woodland/water BA Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] BA Aquatic Habitat Zonation: Dimly-lighted region without emergent vegetation BA Aquatic Habitat Zonation: Specified in Comments Environment Associations - 1 (DRAFT) - Environment Associations Species loon, red-throated Species Id M040003 Date 26 AUG 96 REFERENCES FOR ENVIRONMENTAL ASSOC_ - 624, 700, 3948, 3949, 3946, 1157, 1073, 3950 and 3944 REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 1073, 3950, 1157, 3949 and 3946 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 3949, 1157, 3946 and 1073 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 1073, 3950, 1157, 3949 and 3946 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - 00410S=uses islands as breeding sites, as well as lakes, ponds and marshes *1073,3950,1157,3949*; 00190S=nesting density has been linked to the abundance and availability of shallow water along the coastline of the breeding area *3946* COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - 99999S=often feeds in the ocean *1157,1073* COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - 00410S=nesting occurs to some extent on islands *1073*; 00190S=nesting density has been linked to the abundance and availability of shallow water along the coastline of the breeding grounds *3946* Environment Associations - 2
                               (DRAFT) - Life History
                             Species loon, red-throated
                                 Species Id M040003
                                   Date 26 AUG 96



LIFE HISTORY

ORIGIN: native *624,700*; REPRODUCTION: breeding season=mid-May to mid-July depending upon latitude *1073,3944*; incubation period=24-29 days *3950,3944*; reproductive periods/year=1, unless first clutch is lost then there may be a replacement clutch *3944*; breeding behavior=pair bonding and courtship may begin during migration, behavior is detailed by sources *1073,3950,3944*; clutch size=normally 2, range=1-3 *1073,3944,3948*; BEHAVIOR: territoriality=breeding, over nest site only *3946*; migration=breeding/wintering *624,700,758,1073,3950*; foraging strategy/site=dives for prey in aquatic environments, during breeding season prefers to feed in littoral zones, usually in water less than 1 meter deep, also feeds in ocean *1157,3946,1073*; nest site=lakes, ponds, marshes, and bogs, near waters' edge, lake size is most commonly 1-5 ha, important factor in nesting distribution is nest site with at least 2 sq km of shallow water coastline, generally only one nesting pair per water body unless a large body of water *3946,3946,1073,3950,3944*; development of young=precocial and nidifugous, fledge to maturity in about 43 days *1073,3950,3944*; parental care of young - see *3944,1073*; LIMITING FACTORS: for breeding, availability of shallow water along coastline, low water levels, and amount of disturbance by humans and predators *3944,3946,3947*; POPULATIONS PARAMETERS: relative trend=see source *3944* REFERENCES FOR LIFE HISTORY- 624, 3947, 3948, 3946, 3950, 1073, 1157, 700, 758 and 3944 Life History - 1
                           (DRAFT) - Management Practices
                             Species loon, red-throated
                                 Species Id M040003
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Predator control Beneficial Restricting/regulating human use of habitats Beneficial Restricting/regulating human disturbance of populations Beneficial Public education - publicity Beneficial Maintaining undisturbed/undeveloped areas Beneficial Maintaining wilderness environment Beneficial Maintaining unique or special habitat features [wetlands, caves, Beneficial Maintaining/protecting riparian habitats Beneficial Developing/maintaining brackish marsh Beneficial Developing/maintaining saline marsh Beneficial Developing/maintaining freshwater marsh Beneficial Developing/maintaining/protecting wetlands Beneficial Other management practices [specified in comments] Adverse Draining wetlands, marshes, ponds, lakes REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 3947, 3944, 3948 and 3950 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 3944 and 3950 COMMENTS ON MANAGEMENT PRACTICES - 999(B)=breeding success and nest density may be aided by regulation fishing and other recreational use of breeding habitat *3944,3947* Management Practices - 1
                                   (DRAFT) - References
                                Species loon, red-throated
                                    Species Id M040003
                                      Date 26 AUG 96



     

References

624* Robbins, C.S., Bruun, B., Zim, H.S. 1966. Birds of North America: A Guide to Field Identification. Golden Press New York, N.Y:340. 700* Ornithology, Virginia Society of. 1979. Virginia's Birdlife: An Annotated Check-list. Virginia Avifauna No. 2. Virginia Society of Ornithology Lynchburg, Va:118. 758* (ed.)1982. Fish and wildlife. 50 CFR 10 (Code of Federal Regulations). General Serv. Admin. Washington, D.C:7. 1073* Bent, A.C. 1963. Life Histories of North American Diving Birds. Dover Publ., Inc. New York:244. 1157* Bergman, R.D., Derkson, D.V. 1977. Observations on arctic and red-throated loons at Storkerson Point, Alaska. Arctic 30:41-51. 1901* Fowler, M.E. 1978. Miscellaneous waterbirds (Gaviiformes, Podicipediformes, Procellariformes, Pelicaniformes, and Charadriiformes). Zoo and Wild Animal Medicine Fowler, M.E. W.B. Saunders Co. Phildelphia:213-217. 3944* Bundy, G. 1976. Breeding biology of the red-throated diver. Bird Study 23:249-256. 3946* Merrie, T.D.H. 1978. Relationship between spatial distribution of breeding divers and the availability of fishing waters. Bird Study 253:119-122. 3947* Booth, C.J. 1978. Breeding success of red-throated divers. British Birds 71:44. 3948* Sage, B.L. 1976. Breeding success of red-throated divers on Hascosay. British Birds 69:409. 3949* Norberg, R.A., Norberg, U.M. 1976. Size of fish carried by flying red-throated divers Gavia stellata (Pont.) to nearly fledged young in nesting tarn. Ornis Fenn. 53:92-95. 3950* Cramp, S., Simmons, K.E.L., (eds.). 1977. THE BIRDS OF THE WESTERN PALEARTIC.. References - 1