(DRAFT) - Taxonomy
                              Species gannet, northern
                                 Species Id M040022
                                   Date 26 AUG 96



TAXONOMY

NAME - gannet, northern OTHER COMMON NAMES - ELEMENT CODE - 04/05/84 AOU CODE - 06/04/85 CATEGORY - Birds PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Aves, ORDER AND SUBORDER - Pelecaniformes, FAMILY AND SUBFAMILY - Sulidae, GENUS AND SUBGENUS - Sula, SPECIES AND SSP - bassanus, SCIENTIFIC NAME - Sula bassanus AUTHORITY - Linnaeus TAXONOMY REFERENCES - 624, 3950, 2484, 4122, 4106 and 606 COMMENTS ON TAXONOMY - Formerly Pelecanus bassanus Linaeus, under the order steganopodes; long history of controversy as to whether gannets and boobies should be in the genus Sula, or if gannets should be placed in the genus Morus; when classed under genus Morus, usually known as Morus bassanus *3950,2484,4122,4106,606,624* Taxonomy - 1
                                  (DRAFT) - Status
                              Species gannet, northern
                                 Species Id M040022
                                   Date 26 AUG 96



STATUS

Coded Status Federal Migratory Nongame-Protected REFERENCES FOR STATUS - 758 and 4100 Status - 1
                               (DRAFT) - Distribution
                              Species gannet, northern
                                 Species Id M040022
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - CB1 and AO1 References on Seasonal Occurrence - Common to abundant transient and uncommon winter resident, rare summer visitor *700* References on County Abundance - CB3 and AO1 REFERENCES FOR HYDROLOGIC UNIT CODES - CB3 and AO1 OTHER DISTRIBUTION REFERENCES FOR OTHER DISTRIBUTION - H Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - Aquatic REFERENCES FOR HABITAT - 700, 4122 and 4106 LAND USE - Water Chesapeake Bay Atlantic Ocean Coastal Waters Barren Land Bare Exposed Rock REFERENCES FOR LAND USE - 606, 700, 4122, 4106 and 3950 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine, subtidal OW0 V REFERENCES FOR NWI - 606, 3950, 2484 and 4108 COMMENTS ON HABITAT ASSOCIATIONS - Only comes to land to breed *3950,606* ANIMAL/PLANT SPECIES ASSOCIATIONS - Cholera Salmonella Newcastle disease flies lice mites ticks REFERENCES FOR SPECIES ASSOCIATIONS - 1901, 1959, 2035, 1913, 1952 and 1992 COMMENTS ON SPECIES ASSOCIATIONS - Diseases and parasites *1959,2035,1913,1901* Habitat Associations - 1
                                (DRAFT) - Food Habits
                              Species gannet, northern
                                 Species Id M040022
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - Carnivore REFERENCES FOR TROPHIC LEVEL - 706, 2484 and 3950 LIFESTAGE FOOD FOOD PART General Gadiformes Not Specified General Perciformes Adult stage General Clupeiformes Adult stage General Salmoniformes Not Specified General Pleuronectiformes Not Specified General Semionotiformes Not Specified General Osteichthyes Adult stage General Squid, Octopus Not Specified General Molluscs Not Specified General Crustaceans Not Specified General Echinoderms Not Specified General Animals Adult stage General See Comments; Food See Comments Important Osteichthyes Not Specified Important Animals Not Specified Juvenile Osteichthyes Not Specified Juvenile Perciformes Adult stage Juvenile Clupeiformes Adult stage Juvenile Animals Not Specified Juvenile See Comments; Food See Comments Adult Gadiformes Not Specified Adult Perciformes Adult stage Adult Clupeiformes Adult stage Adult Salmoniformes Not Specified Adult Semionotiformes Not Specified Adult Osteichthyes Adult stage Adult Squid, Octopus Not Specified Adult Molluscs Not Specified Adult Crustaceans Not Specified Adult Echinoderms Not Specified Adult Animals Adult stage Adult See Comments; Food See Comments REFERENCES FOR GENERAL FOOD - 4108, 606, 3950, 4111 and 2484 REFERENCES FOR IMPORTANT FOOD - 2484 and 3950 REFERENCES FOR ADULT FOOD - 4108, 606, 3950, 4111 and 2484 REFERENCES FOR JUVENILE FOOD - 606 and 4111 COMMENTS ON FOOD - 9999S= scraps from fishing vessels and other ships *4108,2484,606* Food Habits - 1 (DRAFT) - Food Habits Species gannet, northern Species Id M040022 Date 26 AUG 96 COMMENTS ON ADULT FOOD - 99999S = feed from scraps and waste of fishing vessels and other ships *2484,606,4108* COMMENTS ON JUVENILE FOOD - 9999S = nestlings fed incompletely regurgitated food by parents *606,4111* Food Habits - 2
                         (DRAFT) - Environment Associations
                              Species gannet, northern
                                 Species Id M040022
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Terrestrial Features: Cliffs/ledges G Biodegradable Organics: Specified in Comments G Aquatic Habitat Zonation: Open water [pelagic zone] G Coastal Features: Rocky offshore islands G Vegetations Successional Stage: Bare rock G Relation to Substrate: Occurs on substrate [not penetrating] G Substrate: Mud or silt G Substrate: Plants LIM Terrestrial Features: Cliffs/ledges LIM Biodegradable Organics: Specified in Comments LIM Coastal Features: Rocky offshore islands LIM Vegetations Successional Stage: Bare rock E Biodegradable Organics: Specified in Comments FJ Aquatic Habitat Zonation: Open water [pelagic zone] RJ Aquatic Habitat Zonation: Open water [pelagic zone] RJ Coastal Features: Rocky offshore islands RJ Terrestrial Features: Cliffs/ledges FA Aquatic Habitat Zonation: Open water [pelagic zone] RA Aquatic Habitat Zonation: Open water [pelagic zone] RA Coastal Features: Rocky offshore islands RA Terrestrial Features: Cliffs/ledges BA Biodegradable Organics: Specified in Comments BA Coastal Features: Rocky offshore islands BA Terrestrial Features: Cliffs/ledges BA Vegetations Successional Stage: Bare rock BA Substrate: Mud or silt BA Substrate: Plants BA Relation to Substrate: Occurs on substrate [not penetrating] REFERENCES FOR ENVIRONMENTAL ASSOC_ - 4101, 4122, 4103, 4110, 4107, 4102, 606, 4113, 3950, 4111, 2484 and 4106 REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 4103, 4110, 4102, 4113, 606, 4122 and 4106 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 4109, 3950, 606 and 2484 REFERENCES FOR RESTING ADULT ENVIRONMENTAL ASSOC_ - 3950, 606 and 2484 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 4102, 4113, 606, 4122, 4106, 4101, 4107, 3950, 2484 and 4111 Environment Associations - 1 (DRAFT) - Environment Associations Species gannet, northern Species Id M040022 Date 26 AUG 96 REFERENCES FOR FEEDING JUVENILE ENVIRONMENTAL ASSOC_ - 4122, 4106, 3950 and 606 REFERENCES FOR RESTING JUVENILE ENVIRONMENTAL ASSOC_ - 606, 4122, 4106 and 3950 REFERENCES FOR EGG ENVIRONMENTAL ASSOC_ - 4103 and 4110 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - 00080S = insecticides, heavy metals, and their effects are discussed by sources *4103,4110,4102*; 99999S = will also nest on gentle slopes of rocky islands *4113,606,4122,4106* COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - 00080S = effects of pollutants on breeding birds is discussed *4102*; 99999S = will nest on gentle slopes of rocky islands; breeds mainly in Gulf of St. Lawrence and Newfoundland *606,4113,4122,4106,4102* COMMENTS ON EGG ENVIRONMENTAL ASSOC_ - 00080S = effects of insecticides and heavy metals on eggshells are discussed *4103,4110* Environment Associations - 2
                               (DRAFT) - Life History
                              Species gannet, northern
                                 Species Id M040022
                                   Date 26 AUG 96



LIFE HISTORY

PHYSICAL DESCRIPTION: adults mostly white, head and neck coverts black, sexes similar, length 35-40 inches, wingspread 5.5 to 6 feet, weight 6-7 lbs, detailed account of age classes given by first source *4122,606,624*; REPRODUCTION: breeding season = late April and May *606,3950,4111*; incubation period 39-46 days, average 44 days *2484,4122,4106,4111,3950,606*; reproductive periods/year = 1 brood/season, will occasionally lay second or third replacement clutch if the others fail *3950,4111,4122,4106*; age at sexual maturity = probably fourth or fifth year *4122,4106*; breeding behavior = colonial, monogamous, mate for life *4122.4105,3950,2484,606*; minimum breeding age = 5 years, occasionally 4 years *4107,4111,4122,4106*; clutch size = 1 *606,3950,4111,2484,4122,4106*; BEHAVIOR: territoriality = breeding season, over nest site only *606,4111,4122,4106*; daily periodicity = strictly diurnal *3950*; dispersion = from 2nd to fifth year, many juveniles prospect at several colonies for suitable nest site *4104*; migration = partial, first year juveniles have true migration along with a small portion of older birds, majority of birds 2 years and older show dispersive movements outside of breeding season *4100,606,3950*; foraging strategy/site = plunge dive from heights of 30-100 feet or greater, dive from water surface, scavenge from ships, usually swallow food under water, feed in open water within limits of continental shelf *4108,4109,3950,606,4122,4106*; nest site = on isolated islands, nest consists of mud, seaweed and other floating debris *4101,4102,606,4122,4106,3950,4113,2484*; development of young = altricial and nidicolous, fledge at 84-97 days (ave. 90 days) *3950,606,4111,2484,4122,4106*; parental care of young = both parents incubate, feed, and brood chick (brood 2-3 weeks); adults are known to adopt guillemot chicks (Cepphus gyrlle) but attempts to brood were unsuccessful *4101,606,3950,4111,4122,4106*; ORIGIN: native *700,3950,606*; POPULATION PARAMETERS: relative trend = world population has increased 2.9% annually, numbers are not obvious at individual gannetries due to immigration *4122,4106,4104,4102,4113*; mortality rates = 6% of breeding adults, about 80% of young/subadults annually *4111,4122,3950*; survival rates = breeding success 75-80%, hatching success 82%, fledging success 78%; breeding adult life expecancy = 16.2 years, breeding success increases with experience; life expectancy = 20-21 years *4107,3950,4111,4122,4106* REFERENCES FOR LIFE HISTORY- 700, 624, 3950, 606, 4101, 4100, 2484, 4104, 4109, 4107, 4102, 4108, 4113, 4111, 4122 and 4106 Life History - 1
                           (DRAFT) - Management Practices
                              Species gannet, northern
                                 Species Id M040022
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Controlling pollution [thermal, chemical, physical] Adverse Applying pesticides Adverse Applying insecticides REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 4103 and 4110 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 4110 and 4103 Management Practices - 1
                                   (DRAFT) - References
                                 Species gannet, northern
                                    Species Id M040022
                                      Date 26 AUG 96



     

References

606* Palmer, R.S. 1962. Handbook of North American Birds. I. Yale Univ. Press New Haven, Conn:557. 624* Robbins, C.S., Bruun, B., Zim, H.S. 1966. Birds of North America: A Guide to Field Identification. Golden Press New York, N.Y:340. 700* Ornithology, Virginia Society of. 1979. Virginia's Birdlife: An Annotated Check-list. Virginia Avifauna No. 2. Virginia Society of Ornithology Lynchburg, Va:118. 706* Warren, B.H. 1890. Birds of Pennsylvania. Commonwealth of Pennsylvania Penn:434. 758* (ed.)1982. Fish and wildlife. 50 CFR 10 (Code of Federal Regulations). General Serv. Admin. Washington, D.C:7. 1901* Fowler, M.E. 1978. Miscellaneous waterbirds (Gaviiformes, Podicipediformes, Procellariformes, Pelicaniformes, and Charadriiformes). Zoo and Wild Animal Medicine Fowler, M.E. W.B. Saunders Co. Phildelphia:213-217. 1913* Peters, H.S. 1936. A list of external parasites from birds of the eastern part of the United States. Bird-Banding 7:9-27. 1952* MacDonald, J.W. 1965. Mortality in wild birds. Bird Study 12:181-195. 1959* Wilson, J.E., MacDonald, J.W. 1967. Salmonella infections in wild birds. Brit. Vet. J. 123:212-219. 1992* Locke, L.N., Stotts, V., Wolfhard, G. 1970. An outbreak of fowl cholera in waterfowl on the Chesapeake Bay. J. Wildl. Dis. 6:404-407. 2035* Palmer, S.F., Trainer, D.O. 1971. Newcastle Disease. Infectious and Parasitic Diseases of Wild Birds Davis, J.W., Anderson, R.C., Karstad, L., Trainer, D.O. The Iowa State University Ames, Iowa:3-16. 2484* Bent, A.C. 1922. Life histories of North American petrels and pelicans and their allies. Dover Publ., Inc. New York:355. 3950* Cramp, S., Simmons, K.E.L., (eds.). 1977. THE BIRDS OF THE WESTERN PALEARTIC.. 4100* Landsborough, T.A. 1974. The migration of the gannet: A reassessment of British and Irish ringing dates. British Birds 67(3):89-103. 4101* Fairhurst, J. 1976. Gannet brooding guillemot chicks. Bird References - 1 (DRAFT) - References Species gannet, northern Species Id M040022 Date 26 AUG 96 Study 23(4):285-286. 4102* Nettleship, D.N. 1976. Gannets in North America: Present numbers and recent population changes. Wilson Bull. 88(2):300-313. 4103* Parslow, J.L.F., Jefferies, D.J. 1977. Gannets and toxic chemicals. British Birds 70(9):366-372. 4104* Nelson, J.B. 1977. Interchange between gannet colonies. British Birds 70(3):127-128. 4106* Nelson, B. 1978. THE GANNET. T. & A.D. Poyser Berkhamsted:336. 4107* Nelson, J.B. 1978. 120(1):132-133. 4108* Watson, P.S. 1981. Seabird observations from commercial trawlers in the Irish Sea. British Birds 74(2):82-90. 4109* Brown, R.G.B. 1979. Seabirds of the Senegal upwelling and adjacent waters. Ibis 121(3):283-292. 4110* Cooke, A.S. 1979. Egg shell characteristics of gannets Sula bassana, shags Phalacrocorax aristotelis and great black-backed gulls Larus marinus exposed to DDE and other environmental pollutants. Environ. Pollut. 19(1)6:47-66. 4111* Nelson, J.B. 1966. The breeding biology of the gannet Sula bassana on the Bass Rock, Scotland. Ibis 108:584-626. 4113* Fisher, J., Vevers, H.G.. The breeding distribution, history, and population of the north Atlantic gannet. J. Anim. Ecol.:173-213. 4122* Grubb, T.C., Jr. 1974. Olefactory navigation to the nesting burrow in Leach's petrel (Oceanodroma leucorhoa). Anim. Behav. 22(1):192-202. References - 2