(DRAFT) - Taxonomy
                                 Species eider, king
                                 Species Id M040072
                                   Date 26 AUG 96



TAXONOMY

NAME - eider, king OTHER COMMON NAMES - ELEMENT CODE - 04/12/84 AOU CODE - 06/04/85 CATEGORY - Birds PHYLUM AND SUBPHYLUM - Chordata, Vertebrata CLASS AND SUBCLASS - Aves, ORDER AND SUBORDER - Anseriformes, FAMILY AND SUBFAMILY - Anatidae, Anthyinae GENUS AND SUBGENUS - Somateria, SPECIES AND SSP - spectabilis, SCIENTIFIC NAME - Somateria spectabilis AUTHORITY - Linnaeus TAXONOMY REFERENCES - 539 Taxonomy - 1
                                  (DRAFT) - Status
                                 Species eider, king
                                 Species Id M040072
                                   Date 26 AUG 96



STATUS

Coded Status Federal Migratory Game (Consumptive Recreational) Accidental REFERENCES FOR STATUS - 424 and 758 Status - 1
                               (DRAFT) - Distribution
                                 Species eider, king
                                 Species Id M040072
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - 001, 095, 115, 131, 199, 550, 650, 700, 800, 810 and 830 References on Seasonal Occurrence - Accidental south to Florida *609*; rare in Chesapeake Bay; a few seen most winters in the lower Bay; viewed from Chesapeake Bay Bridge-Tunnel from Norfolk to Cape Charles, Virginia; seen across mouth of Bay *582* OTHER DISTRIBUTION REFERENCES FOR OTHER DISTRIBUTION - A Comments on Distribution - Accidental south to Florida *609*; rare in Chesapeake Bay; a few seen most winters in the lower Bay; viewed from Chesapeake Bay Bridge-Tunnel from Norfolk to Cape Charles, Virginia; seen across mouth of BAy *582* Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - Aquatic REFERENCES FOR HABITAT - 424 LAND USE - Water Streams and Canals Lakes Bays and Estuaries Wetland Nonforested Wetland Barren Land Sandy Areas other than Beaches Bare Exposed Rock Mixed Barren Land REFERENCES FOR LAND USE - 539, 538, 2825 and 424 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine Riverine Estuarine Lacustrine REFERENCES FOR NWI - 539, 538, 2825 and 424 ANIMAL/PLANT SPECIES ASSOCIATIONS - Diseases and Parasites: Helminths: Nematodes *2051* Protozoan: Blood parasites *1991* REFERENCES FOR SPECIES ASSOCIATIONS - 1897, 1942, 2051 and 1991 COMMENTS ON SPECIES ASSOCIATIONS - Diseases and Parasites: General reference *538,2825*; Habitat Associations - 1
                                (DRAFT) - Food Habits
                                 Species eider, king
                                 Species Id M040072
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - OMNIVORE REFERENCES FOR TROPHIC LEVEL - 424 LIFESTAGE FOOD FOOD PART General Animals Larva stage General Animals Adult stage General Molluscs Adult stage General Bivalve Molluscs Adult stage General Plants Leaves/Needles General Plants Fruit/Seeds General Plants Bole/Stem General Crustaceans Adult stage General Malacostraca Adult stage General Echinoderms Adult stage General Sea/Heart Urchins Adult stage General Trichoptera Larva stage General Anthozoans Adult stage General Monocotyledoneae Leaves/Needles General Hydrocharitaceae Leaves/Needles General Chlorophyta Leaves/Needles General Coleoptera Larva stage General Angiospermae Buds General Angiospermae Leaves/Needles General Angiospermae Fruit/Seeds General Salicaceae Buds General Salicaceae Leaves/Needles General Ranunculaceae Leaves/Needles General Bryophyta Bole/Stem General Tracheophyta Bole/Stem General Bryophyta Leaves/Needles General Tracheophyta Leaves/Needles General Insects Larva stage General Diptera Larva stage General Dicotyledoneae Buds General Dicotyledoneae Leaves/Needles General Dicotyledoneae Fruit/Seeds General Salicaceae Fruit/Seeds REFERENCES FOR GENERAL FOOD - 539, 538, 2825, 554 and 424 Food Habits - 1
                         (DRAFT) - Environment Associations
                                 Species eider, king
                                 Species Id M040072
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Level: Permanently flooded areas G Water Level: Semi-permanently flooded areas G Water Depth Preference: 1-5 ft. G Water Depth Preference: 5-10 ft. G Water Depth Preference: 10-25 ft. G Water Depth Preference: 25-50 ft. G Water Depth Preference: 50-100 ft. G Water Depth Preference: 100-200 ft. G Soil Needs: Rocky G Soil Needs: Sand G Soil Drainage: Well drained G Soil Moisture: Dry G Soil Compaction: Easily penetrated G Terrestrial Features: Rock outcrops G Terrestrial Features: Depressions G Terrestrial Features: Bare ground G Coastal Features: Reefs G Coastal Features: Sand beaches G Coastal Features: Rocky offshore islands G Coastal Features: Sandy offshore islands G Coastal Features: Rocky beaches G Coastal Features: Vegetated offshore islands G Aquatic Features: Pool areas G Inland Wetlands: Inland shallow fresh marshes G Inland Wetlands: Inland open fresh water G Inland Wetlands: Inland saline marshes G Coastal Wetlands: Coastal shallow fresh marsh G Coastal Wetlands: Coastal deep fresh marshes G Coastal Wetlands: Coastal open fresh water G Coastal Wetlands: Sounds and bays G Aquatic/Terrestrial Ecotones: Grassland/water G Hardwood Forest: Willow G Vegetations Successional Stage: Bare rock G Vegetations Successional Stage: Climax grassland G Human Association: Wildlife refuges/sanctuaries REFERENCES FOR ENVIRONMENTAL ASSOC_ - 539, 538, 2825 and 424 Environment Associations - 1
                               (DRAFT) - Life History
                                 Species eider, king
                                 Species Id M040072
                                   Date 26 AUG 96



LIFE HISTORY

PHYSICAL DESCRIPTION:length: 530-600 mm *609*; weight female=1530-2010 g (1830 g average), male=1500-1870 g *539*; adult male (breeding): crown and back of head pale bluish gray forming a smooth crest, separated from greenish cheeks by a whitish line; black spot below the eye; black 'V' mark on the sides of the throat; black feathers bordering the enlarged base of the bill; neck, throat, foreback white; breast creamy white; rest of body and tail brownish black except for a large white patch on each side of the rump; middle and lesser wing coverts white; marginal coverts dusky; greater coverts and secondaries; black; primaries and coverts brownish black; bill bright orange to red with a whitish nail and seasonally enlarged orange to reddish forehead knob; eye dark brown; legs and feet yellowish to dull orange with darker webs *539*; adult female cinnamon buff head and neck finely streaked with black; darker on crown and lighter on throat; rudimentary crest on hind neck; neck brownish black; feathers edged with tawny and buff; rump, tail coverts, breast, sides, flanks cinnamon buff with darker 'U' shaped marks especially on the flanks; tail dark brown; primaries and secondaries blackish brown; secondaries and coverts tipped white; eye brown; bill gray; legs and feet greenish gray to dull yellow with darker webs *539*; eclipse male dark brownish black; lacks crescent markings of the female and retains upper wing coverts and some white feathers on breast or foreback *539*; subadult male in second year resembles adults but median wing coverts are margined or shaded with dusky coloration *539*; juveniles both sexes quite brownish; during late winter male gradually acquires darker back, scapulars, and flanks; develops varying amount of white on breast and rump *539*; voice female=distinctive hollow sounding notes; male=tremulous cooing *539*; REPRODUCTION: courtship period from May to June; nesting begins last half of June and continues 2-3 weeks after arrival on breeding grounds *539,424*; incubation period 22-24 days *539*; clutch size 2-6 eggs (usually 5), average 5.04 eggs *539,538,424*; no renesting due to short breeding season *539*; sexual maturity in third year *538*; pair bond renewed yearly during social courtship during winter and spring *538*; pair bonds formed during second winter or spring of life; displays among spring migrants *539*; male displays modifications of comfort movements; head rolling, bathing, body shaking, wing flapping to expose black abdomen and throat markings, upward stretch, cooing displays include reaching and pushing; head turning conspicuous *539,538*; female displays inciting; male responds by swimming rapidly ahead of her and performing head-turning *539*; copulatory behavior extend selves prone on water; male performs all the displays used during pair formation (comfort movement displays); male treads; performs single display, swims away rapidly while performing head-turning *539*; BEHAVIOR: not social nesters; 6-10/square mile *538*; home range widely dispersed *424*; active at all hours during the summer *538*; will fly over land; flocks of 75-350; follow shoreline but usually keep at least one mile from land; fly abreast in long line; fly 30-40 feet above water *554*; corridors; Southhampton Island on the south, Boothia Peninsula in the center, Ellsmore Island on the north to coastal west Canada, and southwest Greenland; pass around the north and south ends of Baffin Island or head overland over its middle; smallest flocks less than 10, Life History - 1 (DRAFT) - Life History Species eider, king Species Id M040072 Date 26 AUG 96 largest 1100 *424*; first to arrive at coastlines to breed are males; flocks of nearly equal sex ration arrive later; leave late April from winter areas, arrive mid-June on breeding grounds *424*; immatures do not go far north in summer and do not visit breeding grounds *554*; spring migration during the day; midday pause; peaks during mid-morning and mid-afternoon *538*; fall: males migrate early July daily through August; females migrate mid-August to September; males congregate temporarily at sea before leaving; most in east migrate east to Greenland; yearlings summering near the molting grounds join molting males *424*; immatures may remain on molting ground throughout the year; fall migration more continuous from daybreak till sunset *538*; 100000 molt off coast of western Greenland *539*; males complete molt and then continue migration in September and October *424*; unsuccessful females and those that do not attend young form flock on breeding ground and undergo molt migration after broods merge *539*; forage in open sea or coastlines with sources of food at depths shallow enough to permit easy diving; forage in deeper waters than the common eider *538*; dive to 150-180 feet; partially opens wings when diving; probably uses wings and feet to swim *554*; food includes 95% animal matter and 5% plant matter; molluscs make up 46%, 20% is blue mussels; crustaceans make up 19%, 50% is king crab; echinoderms make up 17%, 8% is sand dollars and 6% is sea urchin *2825,424*; 24% of total stomach volume is gravel *554*; also eats caddisfly larvae, sea anemones, eelgrass, wigeon grass, algae, larvae of aquatic insects and beetles, bivalves, crabs, brittle stars, sea cucumbers, midges *539,538,554,538,424*; summer foods young and females eat midge larvae and aquatics, twigs, buds, leaves, catkins of willows, seeds of water buttercup, stems and leaves of moss, fly larvae, grasses, sedges, broad-leaved herbs *539,424*; winter food molluscs, sea urchins, sand dollars, crustaceans, algae, eelgrass (Zostera), wigeon grass (Ruppia) *539*; high Arctic nesting grounds, females arrive mid-June, spread out over tundra habitats, concentrate on river islands as protection from arctic fox predation; nest well away from water; on dry and rocky slopes up to 1/4 mile from nearest water; along fresh water ponds, lakes, and streams on arctic tundra, usually near coastlines with occassional nesting just above the high-tide line of seacoasts *539*; penetrates inland along watercourses; breeds further north than any other duck; likes areas that first become free of snow; dry, well vegetated areas on ground with moss and lichen growth; bare stretch of gravel or sand, amongst loose stones *2825*; off Canadian coast, not as common off Alaska as the common eider; nest beside lakes, on small islands in lakes, low marshy country, almost bare hillsides, low mossy tundra, dry grassy tundra; usually well scattered nests but when fox predation is high, concentrate nests on river islands; often on dry rocky slopes; appear more frequently on interior lakes *538*; closest nests 200 yards apart *538*; isolated or grouped nests; on barrier islands *424*; incubation by female alone; as soon as clutch completed males begin to migrate to molt (molt less than 1000 miles from breeding areas) *539*; determinate nesters, little significant renesting *424*; nest built entirely of down *607*; lays within one week of arrival on territory *2825*; one egg laid per day; eggs bright olive; 64 X 43 mm ; 73 g *539*; male closely attends female only during egg laying; follows her to nest site; female spends little time off nest while incubating *538*; brood brought to ponds and eventually get to the Life History - 2 (DRAFT) - Life History Species eider, king Species Id M040072 Date 26 AUG 96 coast sometimes via streams; very fast growth rate; fly in slightly over 1 month *424*; broods merge with over 100 ducklings and several females *539*; attending females may remain in breeding area till September *538*; non-breeding habitat found at sea; rest on drift ice; feed off coastlines in deep waters; large flocks during non-breeding season; 15000 birds winter off St. Lawrence Island in February; pair formation begins in these blocks *539*; winter on North Atlantic coast and sometimes on Great lakes; frequently found south of natural habitat *554*; found in ocean and along coasts *609*; winter as far north as open water permits; 40-90 degrees north along Atlantic coast from Nova Scotia to Virginia (Audubon Christmas counts); found singly or in duos *424*; POPULATION PARAMETERS: North American population 1-1.5 million *539*; Audubon Christmas counts (1954-1962) 16.5 at 53 stations *538*; 100000 in eastern Arctic and Greenland; sex ratio 1 male per 3.4 females during migration in August; 60% males on breeding ground *424*; mortality factors: starvation caused by late breakup of sea ice; 10% hunting mortality in Greenland; foxes take large numbers of eggs after a crash in the lemming population *424*; gulls take eggs also *538*; AQUATIC/TERRESTRIAL ASSOCIATIONS: predators: arctic foxes, gulls *2825,424* REFERENCES FOR LIFE HISTORY- 539, 538, 2825, 582, 607, 554, 609 and 424 Life History - 3
                           (DRAFT) - Management Practices
                                 Species eider, king
                                 Species Id M040072
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                   Species eider, king
                                    Species Id M040072
                                      Date 26 AUG 96



     

References

424* Bellrose, R.C. 1978. Ducks, Geese and Swans of North America. 2nd ed.. Stackpole Books Harrisburg, Penn:540. 538* Johnsgard, P.A. 1975. Waterfowl of North America. Indiana Univ. Press Bloomington:575. 539* Johnsgard, P.A. 1978. Ducks, Geese, and Swans of the World. Univ. Nebraska Press Lincoln:404. 554* Kortright, F.H. 1967. The Ducks, Geese, and Swans of North America. Stackpole Books Harrisburg, Penn:476. 582* Meanley, B. 1982. Waterfowl of the Chesapeake Bay Country. Tidewater Publ. Centreville, Md:210. 607* Pearson, G.T. (ed.)1936. Birds of America. Garden City Publ. Co. Garden City, N.Y:260. 609* Peterson, R.T. 1980. Birds of eastern and central North America. 4th Ed.. Houghton Mifflin Co. Boston, Mass:384. 758* (ed.)1982. Fish and wildlife. 50 CFR 10 (Code of Federal Regulations). General Serv. Admin. Washington, D.C:7. 1897* Humphreys, P. 1978. Ducks, geese, swans (Anseriformes). Zoo and Wild Animal Medicine Fowler, M.E. W.B. Saunders Co. Philidelphia:183-209. 1942* Wobeser, G.A. 1981. DISEASES OF WILD WATERFOWL.:300. 1991* Bennett, G.F., Inger, J.G. 1972. Blood parasites of gamebirds from insular Newfoundland. Can. J. Zool. 50:705-706. 2051* Wehr, E.E. 1971. Nematodes. Infectious and Parasitic Diseases of Wild Birds Davis, J.W., Anderson, R.C., Karstad, L., Trainer, D.O. The Iowa State University Ames, Iowa:185-233. 2825* Soothill, E., Whitehead, P. 1978. WILDLIFE OF THE WORLD. Blandford Press:297. References - 1