(DRAFT) - Taxonomy
                           Species dowitcher, short-billed
                                 Species Id M040142
                                   Date 26 AUG 96



TAXONOMY

NAME - dowitcher, short-billed OTHER COMMON NAMES - ELEMENT CODE - 04/06/84 AOU CODE - 10/15/85 CATEGORY - Birds PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Aves, ORDER AND SUBORDER - Charadriiformes, FAMILY AND SUBFAMILY - Scolopacidae, GENUS AND SUBGENUS - Limnodromus, SPECIES AND SSP - griseus, SCIENTIFIC NAME - Limnodromus griseus AUTHORITY - Gmelin TAXONOMY REFERENCES - 624 Taxonomy - 1
                                  (DRAFT) - Status
                           Species dowitcher, short-billed
                                 Species Id M040142
                                   Date 26 AUG 96



STATUS

Coded Status Federal Migratory Nongame-Protected REFERENCES FOR STATUS - 4179, 758 and 733 COMMENTS ON STATUS - Protected - cannot be legally hunted or sold *733* Status - 1
                               (DRAFT) - Distribution
                           Species dowitcher, short-billed
                                 Species Id M040142
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - 001, 003, 005, 007, 009, 011, 013, 015, 017, 019, 021, 023, 025, 027, 029, 031, 033, 035, 036, 037, 041, 043, 045, 047, 049, 051, 053, 057, 059, 061, 063, 065, 067, 069, 071, 073, 075, 077, 079, 081, 083, 085, 087, 089, 091, 093, 095, 097, 099, 101, 103, 105, 107, 109, 111, 113, 115, 117, 119, 121, 125, 127, 131, 133, 135, 137, 139, 141, 143, 145, 147, 149, 153, 155, 157, 159, 161, 163, 165, 167, 169, 171, 173, 175, 177, 179, 181, 183, 185, 187, 191, 193, 195, 197, 199, 510, 515, 520, 530, 540, 550, 560, 570, 580, 590, 595, 600, 610, 620, 630, 640, 650, 660, 670, 678, 680, 683, 685, 690, 700, 710, 720, 730, 735, 740, 750, 760, 770, 775, 780, 790, 800, 810, 820, 830, 840, CB1, CB2, CB3 and AO1 References on Seasonal Occurrence - Abundant transient; rare to uncommon visitor along coast; rare fall and spring transient in mountain regions *700,609,624* References on County Abundance - 007, 037, 089, 141, 025, 075, 107, 125, 143, 177, 011, 061, 109, 145, 029, 111, 147, 031, 047, 065, 081, 113, 049, 067, 083, 135, 003, 019, 117, 137, 053, 157, 023, 057, 073, 105, 121, 159, 175, 193, 540, 610, 680, 750, 820, 009, 041, 059, 091, 161, 195, 620, 690, 760, 830, 027, 043, 077, 127, 163, 179, 197, 560, 630, 770, 840, 013, 045, 063, 079, 095, 165, 181, 570, 640, 775, CB1, 097, 133, 149, 167, 183, 510, 580, 720, 780, CB2, 033, 099, 115, 153, 169, 185, 515, 590, 660, 730, 790, CB3, 035, 051, 069, 085, 101, 155, 171, 187, 520, 595, 670, 735, 005, 021, 036, 071, 087, 103, 119, 139, 173, 191, 530, 600, 678, 740, 550, 093, 700, 131, 199, 710, 650, 001, 800, AO1 and 810 REFERENCES FOR HYDROLOGIC UNIT CODES - 550, 093, 700, 131, 199, 710, 650, 001, 800, AO1 and 810 OTHER DISTRIBUTION REFERENCES FOR OTHER DISTRIBUTION - F Comments on Distribution - Resident only in some coastal counties, transient over entire state *609,700,624* Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - Aquatic REFERENCES FOR HABITAT - 4188, 4189, 4179 and 4181 LAND USE - Cropland and Pasture Water Streams and Canals Lakes Reservoirs Bays and Estuaries Chesapeake Bay Atlantic Ocean Coastal Waters Wetland Nonforested Wetland Barren Land Beaches REFERENCES FOR LAND USE - 4188, 4184, 4189, 4182, 4187 and 4181 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Lacustrine, littoral FL. N 2 Lacustrine, littoral BB. N 3 Riverine, lower perennial BB. N Riverine, lower perennial FL. Palustrine FL. Marine, intertidal BB2 Estuarine, intertidal EM. Estuarine, intertidal FL2 Estuarine, intertidal FL3 REFERENCES FOR NWI - 4181, 1169, 528, 4182, 4187 and 4189 COMMENTS ON HABITAT ASSOCIATIONS - Occurs along edges of muddy ponds, bays, marshy lakes and ponds, on mud flats and sand flats *1169,528* POTENTIAL NATURAL VEGETATION - 065 Northern Cordgrass Prairie (Distichlis-Spartina) 101 Oak - Hickory - Pine Forest (Quercus-Carya-Pinus) 103 Southern Floodplain Forest (Quercus-Nyssa-Taxodium) REFERENCES FOR PNV - 609 ECOREGION - Southeastern Mixed Forest: Flat Plains REFERENCES FOR ECOREGION - 609 Habitat Associations - 1
                                (DRAFT) - Food Habits
                           Species dowitcher, short-billed
                                 Species Id M040142
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - Carnivore REFERENCES FOR TROPHIC LEVEL - 4186, 4182, 4184 and 4166 LIFESTAGE FOOD FOOD PART General Plants Fruit/Seeds General Polychaetes Not Specified General Oligochaetes Not Specified General Annelids Not Specified General Molluscs Not Specified General Malacostraca Not Specified General Ostracods Not Specified General Crustaceans Egg/Fetus stage General Animals Egg/Fetus stage Important Polychaetes Not Specified Important Annelids Not Specified Important Molluscs Not Specified Important Malacostraca Egg/Fetus stage Important Crustaceans Egg/Fetus stage Important Animals Egg/Fetus stage Adult Plants Fruit/Seeds Adult Polychaetes Not Specified Adult Oligochaetes Not Specified Adult Annelids Not Specified Adult Molluscs Not Specified Adult Malacostraca Not Specified Adult Ostracods Not Specified Adult Crustaceans Egg/Fetus stage Adult Animals Egg/Fetus stage Adult See Comments; Food See Comments REFERENCES FOR GENERAL FOOD - 4166, 4184, 4182 and 4186 REFERENCES FOR IMPORTANT FOOD - 4184 REFERENCES FOR ADULT FOOD - 4166, 4184, 4182 and 4186 COMMENTS ON ADULT FOOD - 9999S=feeds on grasshoppers, beetles, flies, maggots, marine worms, leeches, fish eggs, small mollusks, and occasionally seeds of aquatic plants *528,1169* Food Habits - 1
                         (DRAFT) - Environment Associations
                           Species dowitcher, short-billed
                                 Species Id M040142
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Inland Wetlands: Seasonally flooded lands with emergent vegetation G Aquatic Features: Silt bottom streams G Coastal Features: Mudflats G Coastal Features: Sand beaches G Coastal Features: Sand bars G Soil Needs: Sand G Coastal Wetlands: Coastal salt meadows G Coastal Wetlands: Regularly flooded salt marshes G Coastal Wetlands: Sounds and bays G Coastal Wetlands: Mangrove swamps G Inland Wetlands: Shrub swamps G Water Depth Preference: Less than 1 ft. G Aquatic/Terrestrial Ecotones: Woodland/water G Aquatic/Terrestrial Ecotones: Woodland/grassland G Inland Wetlands: Inland open fresh water G Coastal Features: Specified in Comments G Human Association: Abandoned buildings G Soil Compaction: Easily penetrated G Soil Moisture: Wet G Soil Moisture: Moist G Density of Aquatic Vegetation: Low G Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] G Coastal Wetlands: Coastal salt flats G Water Level: Permanently flooded areas LIM Coastal Features: Specified in Comments LIM Aquatic/Terrestrial Ecotones: Woodland/water LIM Inland Wetlands: Shrub swamps LIM Inland Wetlands: Seasonally flooded lands with emergent vegetation LIM Inland Wetlands: Inland deep fresh marshes LIM Coastal Features: Mudflats LIM Coastal Wetlands: Regularly flooded salt marshes LIM Soil Moisture: Wet LIM Soil Moisture: Moist LIM Soil Compaction: Easily penetrated LIM Soil Needs: Sand LIM Water Depth Preference: Less than 1 ft. LIM Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] FA Coastal Features: Mudflats FA Coastal Features: Sand beaches FA Soil Compaction: Easily penetrated Environment Associations - 1 (DRAFT) - Environment Associations Species dowitcher, short-billed Species Id M040142 Date 26 AUG 96 LIFESTAGE ENVIRONMENTAL ASSOCIATIONS FA Soil Moisture: Wet FA Soil Moisture: Moist FA Soil Needs: Sand FA Coastal Wetlands: Regularly flooded salt marshes FA Coastal Wetlands: Sounds and bays FA Coastal Wetlands: Mangrove swamps FA Water Depth Preference: Less than 1 ft. FA Inland Wetlands: Inland fresh meadows [fen] FA Coastal Features: Specified in Comments FA Density of Aquatic Vegetation: Low FA Coastal Wetlands: Coastal salt flats FA Coastal Wetlands: Coastal salt meadows FA Aquatic Features: Silt bottom streams RA Inland Wetlands: Shrub swamps RA Human Association: Abandoned buildings RA Aquatic Features: Silt bottom streams BA Inland Wetlands: Shrub swamps BA Inland Wetlands: Wooded swamps BA Aquatic/Terrestrial Ecotones: Woodland/water BA Aquatic/Terrestrial Ecotones: Woodland/grassland BA Aquatic/Terrestrial Ecotones: Grassland/water BA Inland Wetlands: Inland shallow fresh marshes BA Coastal Features: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 4188, 4184, 4189, 4179, 4186, 4187, 4181, 4182, 1169 and 528 REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 4188, 4184, 4189, 4179, 4186, 4187, 4181 and 4182 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 4188, 4184, 4189, 4179, 4186, 4187, 4181, 4182, 1169 and 528 REFERENCES FOR RESTING ADULT ENVIRONMENTAL ASSOC_ - 4187, 1169 and 528 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 4189, 624, 4179 and 4187 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - 99999S=breeds in arctic regions of Canada and U.S., in marshy habitats *4189,4179,4187*; feeds almost exclusively in intertidal mud and sand flats *4186,4189,4184,4188,4182,4181*; 00420S=forages and breeds on islands *4181,4187* COMMENTS ON RESTING ADULT ENVIRONMENTAL ASSOC_ - rests in marshes and meadows hidden on ground in vegetation *1169* COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - 00420S=feeds on coastal islands *4181* COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - 99999S=breeds in artic regions of Canada & U.S. *624,4189,4179,4187*; 00420S=sometimes breeds on islands *4181* Environment Associations - 2
                               (DRAFT) - Life History
                           Species dowitcher, short-billed
                                 Species Id M040142
                                   Date 26 AUG 96



LIFE HISTORY

ORIGIN: Native *700,609*; BEHAVIOR: Territoriality=breeding *4187*; Daily Periodicity=diurnal *1169*; feeding habitat use is related to tide, peak use at midpoints of rising and ebbtides *4181*; Migration=breeding/wintering *758,4179,624,700*; Foraging Strategy/Sites=feed in cohesive flocks by probing soft substrate with bill, feed primarily in intertidal regions of mudflats and sandflats in water less than 6 cm deep *4188,4184,4189,4179,4186,4182,4181*; Nest Site=on tundra and taiga borders near or in muskeg, sedge, wet or grass meadows, on coast and islands *4189,4179,4187*; Development of Young=precocial *4179*; Parental Care of Young=adults known to move eggs to new nest site after human disturbance *4185*; AQUATIC/TERRESTRIAL ASSOCIATIONS: Calidris pusillus, Calidris minutilla, Charadrius semipalmatus, Arenaria ineterpres *1169,528 * REFERENCES FOR LIFE HISTORY- 700, 609, 4187, 1169, 758, 4179, 624, 4188, 4184, 4189, 4186, 4182, 4181 and 4185 Life History - 1
                           (DRAFT) - Management Practices
                           Species dowitcher, short-billed
                                 Species Id M040142
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Maintaining undisturbed/undeveloped areas Beneficial Maintaining early stages of ecological succession Beneficial Restricting/regulating human disturbance of populations Beneficial Maintaining unique or special habitat features [wetlands, caves, Beneficial Maintaining/protecting riparian habitats Beneficial Developing/maintaining brackish marsh Beneficial Developing/maintaining saline marsh Beneficial Developing/maintaining freshwater marsh Beneficial Developing/maintaining mudflats Beneficial Developing/maintaining/protecting wetlands Beneficial Creating impoundments Beneficial Maintaining bogs Beneficial Maintaining wilderness environment Beneficial Developing/maintaining water holes, ponds, potholes, etc. Adverse Draining wetlands, marshes, ponds, lakes Adverse Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 4185, 4188, 4184, 4189, 4181 and 4182 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 4184, 4189, 4181 and 4182 COMMENTS ON MANAGEMENT PRACTICES - 999(A)=Adversely affected by loss of habitat through developement, draining, and filling of marshlands ; industrial pollution, clean farming, intensive agricultural practices, intensive recreational development, urban and agricultural development, creation of suburban residential areas, channelization, application of of herbicides, pesticides, and insecticides *1169* Management Practices - 1
                                   (DRAFT) - References
                             Species dowitcher, short-billed
                                    Species Id M040142
                                      Date 26 AUG 96



     

References

528* Imhof, T.A. 1976. Alabama Birds, 2nd Ed.. Univ. Alabama Press:445. 609* Peterson, R.T. 1980. Birds of eastern and central North America. 4th Ed.. Houghton Mifflin Co. Boston, Mass:384. 624* Robbins, C.S., Bruun, B., Zim, H.S. 1966. Birds of North America: A Guide to Field Identification. Golden Press New York, N.Y:340. 700* Ornithology, Virginia Society of. 1979. Virginia's Birdlife: An Annotated Check-list. Virginia Avifauna No. 2. Virginia Society of Ornithology Lynchburg, Va:118. 733* Commission, Pennsylvania Game. 1977. The Commonwealth of Pennsylvania Game Laws. Penn. Game Comm. Harrisburg, Penn:137. 758* (ed.)1982. Fish and wildlife. 50 CFR 10 (Code of Federal Regulations). General Serv. Admin. Washington, D.C:7. 1169* Bent, A.C. 1927. Life histories of North American shore birds. Part I. U.S. Natl. Mus. Bull. No. 142:420. 4166* Gross, A.O. 1950. The herring gull-cormorant control project. Proc. 10th Int. Congr. Ornithol.:532-536. 4179* Baker, M.C., Miller, A.E. 1973. Niche relationships among six species of shorebirds on the wintering and breeding ranges. Ecol. Monogr. 43(2):193-212. 4181* Burger, J., Howe, M.A., Caldwell, D.C., Chase, J. 1977. Effects of tide cycles on habitat selection and habitat partitioning by migrating shorebirds. Auk 94:743-758. 4182* Recher, H.F. 1966. Some aspects of the ecology of migrating shorebirds. Ecology 47:393-407. 4184* Mallory, E.P., Schneider, D.C. 1979. Agonistic behavior in short-billed dowitchers (Limnodromus griseus) feeding on a patchy resource. Wilson Bull. 91(2):271-278. 4185* Jehl, J.R. 1979. Nest shifting by a dowitcher. Bird-Banding 50(4):367-368. 4186* Quammen, M.L. 1982. Influence of subtle substrate differences on feeding by shorebirds on intertidal mudflats. Mar. Biol. 71(3):339-343. 4187* Gill, R.E., Jr., Petersen, M.R., Jorgensen, J.D. 1981. Birds of the northcentral Alaska Peninsula, USA, 1976-1980. Arctic 34(4):286-306. References - 1 (DRAFT) - References Species dowitcher, short-billed Species Id M040142 Date 26 AUG 96 4188* Quammen, M.L. 1981. Use of exclosures in studies of predation by shorebirds on intertidal mudflats. Auk 98(4):812-817. 4189* Baker, M.C. 1979. Morphological correlates of habitat selection in a community of shorebirds (Charadriiformes). Oikos 33(1):121-126. References - 2