(DRAFT) - Taxonomy
                                 Species sanderling
                                 Species Id M040145
                                   Date 26 AUG 96



TAXONOMY

NAME - sanderling OTHER COMMON NAMES - ruddy plover, beach-bird, surf snipe, white snipe, bull peep and whitey ELEMENT CODE - 07/19/84 AOU CODE - 08/29/86 CATEGORY - Birds PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Aves, ORDER AND SUBORDER - Charadriiformes, FAMILY AND SUBFAMILY - Scolopacidae, Calidridinae GENUS AND SUBGENUS - Calidris, SPECIES AND SSP - alba, SCIENTIFIC NAME - Calidris alba AUTHORITY - Pallas TAXONOMY REFERENCES - 3607 and 541 COMMENTS ON TAXONOMY - AKA: ruddy plover, beach-bird, surf snipe, white snipe, beach plover, whitey, bull peep *607* Taxonomy - 1
                                  (DRAFT) - Status
                                 Species sanderling
                                 Species Id M040145
                                   Date 26 AUG 96



STATUS

Coded Status Federal Migratory Nongame-Protected REFERENCES FOR STATUS - 3609 and 758 Status - 1
                               (DRAFT) - Distribution
                                 Species sanderling
                                 Species Id M040145
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - 001, 131, 199, 550, 650, 700, 710, 740, 800, 810 and CB1 References on County Abundance - 001, 131, 199, 550, 650, 700, 710, 740, 800, 810, CB1 and 999 REFERENCES FOR HYDROLOGIC UNIT CODES - 001, 131, 199, 550, 650, 700, 710, 740, 800, 810 and CB1 OTHER DISTRIBUTION REFERENCES FOR OTHER DISTRIBUTION - H Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - Terrestrial REFERENCES FOR HABITAT - 541 REFERENCES FOR FSIZE - 3609 and 541 LAND USE - Streams and Canals Reservoirs Water Lakes Bays and Estuaries Wetland Nonforested Wetland Barren Land Beaches Sandy Areas other than Beaches Mixed Barren Land REFERENCES FOR LAND USE - 6069, 4182, 4652, 609, 619, 3606, 3607, 3609, 541 and 1082 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine, intertidal FL2 Estuarine, intertidal FL2 Palustrine Riverine Lacustrine REFERENCES FOR NWI - 4182, 4652 and 541 ANIMAL/PLANT SPECIES ASSOCIATIONS - 541 POTENTIAL NATURAL VEGETATION - 065 Northern Cordgrass Prairie (Distichlis-Spartina) 101 Oak - Hickory - Pine Forest (Quercus-Carya-Pinus) 103 Southern Floodplain Forest (Quercus-Nyssa-Taxodium) REFERENCES FOR PNV - 609 ECOREGION - Southeastern Mixed Forest: Flat Plains REFERENCES FOR ECOREGION - 609 Habitat Associations - 1
                                (DRAFT) - Food Habits
                                 Species sanderling
                                 Species Id M040145
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - Carnivore REFERENCES FOR TROPHIC LEVEL - 541 LIFESTAGE FOOD FOOD PART General Malacostraca Juvenile stage General Copepods Adult stage General Crustaceans Adult stage General Crustaceans Juvenile stage General Molluscs Adult stage General Insects Larva stage General Insects Pupa stage General Insects Adult stage General Branchiopods Adult stage General Malacostraca Adult stage General Annelids Adult stage General Diptera Larva stage General Diptera Adult stage General Other Plant Species Fruit/Seeds General Saxifragaceae Buds General Lycopsida Leaves/Needles General Chlorophyta Bole/Stem General Chlorophyta Leaves/Needles General Chlorophyta Rhizome General Ostracods Adult stage General Polychaetes Adult stage General Plants Bole/Stem General Plants Leaves/Needles General Plants Rhizome General Plants Buds General Plants Fruit/Seeds General Tracheophyta Bole/Stem General Tracheophyta Leaves/Needles General Tracheophyta Rhizome General Tracheophyta Buds General Tracheophyta Fruit/Seeds General Angiospermae Buds General Dicotyledoneae Buds General Animals Larva stage General Animals Pupa stage General Animals Juvenile stage General Animals Adult stage General Spermopsida Buds Juvenile Animals Larva stage Juvenile Animals Pupa stage Juvenile Insects Larva stage Juvenile Insects Pupa stage Adult Bryophyta Not Specified Adult Plants Fruit/Seeds Adult Polychaetes Not Specified Food Habits - 1 (DRAFT) - Food Habits Species sanderling Species Id M040145 Date 26 AUG 96 LIFESTAGE FOOD FOOD PART Adult Crustaceans Not Specified Adult Insects Not Specified Adult Diptera Adult stage Adult Diptera Larva stage Adult Molluscs Not Specified Adult Ostracods Not Specified Adult Siphonaptera Not Specified REFERENCES FOR GENERAL FOOD - 3609, 3606, 3607 and 541 REFERENCES FOR ADULT FOOD - 541 and 1169 REFERENCES FOR JUVENILE FOOD - 3609 Food Habits - 2
                         (DRAFT) - Environment Associations
                                 Species sanderling
                                 Species Id M040145
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Substrate: Sand G Relation to Substrate: Occurs on substrate [not penetrating] G Bottom Type [Aquatic]: Sand G Stability of Bottom: Unstable G Density of Aquatic Vegetation: Low G Aquatic Habitat Zonation: Shallows with emergent vegetation [littoral zone] G Water Level: Permanently flooded areas G Elevation: Below sea level G Elevation: 0-100 ft. G Elevation: 100-300 ft. G Soil Needs: Sand G Soil Needs: Clay G Soil Drainage: Well drained G Terrestrial Features: Ridges G Terrestrial Features: Bare ground G Coastal Features: Sand beaches G Coastal Features: Sand bars G Coastal Features: Mudflats G Coastal Features: Dunes G Coastal Features: Hind-dune G Coastal Features: Sandy offshore islands G Aquatic Features: Pool areas G Aquatic Features: Backwaters G Inland Wetlands: Inland shallow fresh marshes G Inland Wetlands: Inland open fresh water G Coastal Wetlands: Coastal shallow fresh marsh G Coastal Wetlands: Coastal open fresh water G Coastal Wetlands: Sounds and bays G Percent Canopy Closure of Preferred Trees: Specified in Comments G Vegetations Successional Stage: Sand dune G Human Association: Wildlife refuges/sanctuaries REFERENCES FOR ENVIRONMENTAL ASSOC_ - 609, 619, 3606, 3607, 3609, 541 and 1082 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - 00740S=0% *541* Environment Associations - 1
                               (DRAFT) - Life History
                                 Species sanderling
                                 Species Id M040145
                                   Date 26 AUG 96



LIFE HISTORY

PHYSICAL DESCRIPTION: breeding adults-crown and hindneck=cinnamon rufous to rusty streaked with dusky and grayish white; sides of head and neck=similar but paler; foreneck and chest=same but barred; chin, throat, underparts=mostly pure white with some spotting on the sides of the breast; scapulars=black centrally with pale whitish or brownish margins; rump and median tail coverts=brownish gray with white borders, the white increasing laterally; tail feathers=mostly gray with white margins, the outermost pair being largely white; wing coverts=mostly dusky with whitish margins but the greater coverts are brownish gray with broad white tips; secondaries=white on basal half and edged with white; primaries=dusky with the inner ones having considerable white on the basal parts of the outer webs; eye=dark brown; bill=black; legs and toes=black; hind toe absent *541*; winter adult-crown, back, upper wing coverts=gray with dusky shaft streaks; anterior lesser coverts=dusky; underparts=white superciliary stripe *541*; juveniles-crown and back=blackish spotted with whitish buff; wing coverts=have wide buffy fringes; underparts=white with buffy wash on breast *541*; sexes alike *1082*; length=18-20 cm *609*; average length=20.5 cm *1082*; weight, female=52-62 g, male=50-62 g *541*; wing, male=113-125 mm, female=113-137 mm *541*; culmen, male=23-26 mm, female=24-28 mm *541*; often stand or hop on one leg; flock with themselves or in groups with knots, small plovers, and other species; flight is swift, direct and low over water *3609*; REPRODUCTION: breeding season=mid-June to early July *1082*; arrive on breeding grounds near end of May and in late June *541*; incubation period=23-24 days *3607*; both birds may incubate *3609*; incubation begins when clutch complete *541*; male may leave female, help with incubation or raise brood alone while female initiates a second clutch *541*; reproductive periods/year=double clutch *541*; breeding behavior=during courtship the male rises with rapid wingbeats and then begins a short downward flight, uttering a song as he descends *3607*; monogamous; old pair bonds may be reestablished; pairs form soon after migrant flocks break up; flight display is a mate advertisement device that ceases with formation of pair bond; bird ascends to a height of 30-60 feet, then levels off and begins to fly with rapidly vibrating wings alternated with brief glides with the head drawn back into body, song uttered in bursts; bird descends within 200 yards of point of takeoff; ground display of male=following female with hunched head, drooping wings, erected feathers, tail feathers spread and depressed *541*; clutch size=4 eggs *1082*; sometimes 3 eggs *541*; egg=light olive brown speckled and spotted with different shades of brown, chiefly at large end *607*; 35.7 X 24.7 mm *3609*; 11.2 g; egg=20.5% of female's body weight; eggs laid at 26-29 hour intervals *541*; BEHAVIOR: territoriality=portion of level or sloping ground above which flight displays are performed and includes an area about 400 yards in diameter; territorial behavior has been observed but not all individuals establish territories *541*; breeding density=low; 3-4 pairs/square mile *541*; migration=found on sandy shores during migration *1082*; flies over ocean, rests on water *607*; during migration found on beaches and about large bodies of water in the interior *619*; spring migration begins March to late April; fall migration begins in August *3609*; food and Life History - 1 (DRAFT) - Life History Species sanderling Species Id M040145 Date 26 AUG 96 foraging=carnivore *541*; feeds in backwash of waves; whole flock moves in to ebbing water to feed and retreats before next wave *3606*; may also search and probe on beach *3606*; probe in wet sand for tiny animals and snatch those exposed by the washing of the waves *617*; feed in wet sand by running quickly and producing a series of closely spaced shallow probes; on wintering areas they typically forage in small flocks on sandy areas of tidal flats, usually in groups of up to 10 birds *541*; juvenile foods=insect larvae and nymphs *3609*; foods include young stages of burrowing sand crabs, amphipods *3606*; small crustaceans, molluscs, insects and larvae *3607*; sandfleas, shrimp, marine worms, flies and larvae, seeds, buds of saxifrage, moss, algae *3609*; midge, mosquito, and cranefly larvae, sand flies, ostracods, polychaete worms *541*; nesting and nest site=Arctic, circumpolar *609*; eastern Greenland, parts of northern Siberia and its offshore islands, also parts of arctic Canada, Alaska, northwest Spitsbergen *1082*; high Arctic *541*; stony tundra *609*; stony, sparsely vegetated regions of the Arctic tundra, often near water *3607*; gravel ridges; near freshwater, usually a lake or pond *3609*; rock desert and barren tundra with scant vegetation; not far from freshwater but some distance from coast; stony well-drained ridges, gently slopes or level alluvial plains; less than 200 feet above sea level *541*; stony or clay soils in moor and marsh areas *3609*; nest=deep scrape close to thick vegetation and sparsely lined with leaves *1082*; nest cover=Arctic willow *3609*; cover=Salix arctica, Dryas octopetala, Saxifraga oppositifolia; nest placed between bare clay lumps on moss *3609*; nest placed in open situation; fully exposed to sky *541*; care and development of young=young leave the nest a few hours after hatching and are looked after by both parents for a few weeks *3607*; female leads intruders away from nest and also protects hatched brood; male leaves female after eggs hatch; female guards young for 12-14 days; leads young to freshwater beaches; female broods young in cold or wet weather; young are precocial *3609*; fledging period=17 days *541*; non-breeding habitat and behavior=winters from United States, Britain, China to Southern Hemisphere *609*; winters along both United States' coasts *3606*; outer beaches, tideflats, lakeshores *609*; sandy shores *1082*; large bays; sandy substrate *3606*; flat sandy beaches, mudbanks, estuaries; rare inland *3607*; gregarious, mix with other flocking species present; under rough conditions they mass together on the high beach or gather in small groups in the lee of nearby dunes *3606*; rest on crest of beach or dunes behind it *3609*; when approached, generally runs rather than flying away; lives in groups that are more or less continously active *3607*; ORIGIN: native *609* REFERENCES FOR LIFE HISTORY- 609, 1082, 607, 619, 3606, 617, 3607, 3609 and 541 Life History - 2
                           (DRAFT) - Management Practices
                                 Species sanderling
                                 Species Id M040145
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Developing/maintaining mudflats Beneficial Controlling water levels Adverse Draining wetlands, marshes, ponds, lakes REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 4684 and 4685 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 4469 Management Practices - 1
                                   (DRAFT) - References
                                    Species sanderling
                                    Species Id M040145
                                      Date 26 AUG 96



     

References

541* Johnsgard, P.A. 1981. The Plovers, Sandpipers, and Snipes of the World. Univ. Nebrasks Press Lincoln:493. 607* Pearson, G.T. (ed.)1936. Birds of America. Garden City Publ. Co. Garden City, N.Y:260. 609* Peterson, R.T. 1980. Birds of eastern and central North America. 4th Ed.. Houghton Mifflin Co. Boston, Mass:384. 617* Potter, E.F., Parnell, J.F., Teulings, R.P. 1980. Birds of the Carolinas. Univ. N.C. Press Chapel Hill, N.C:408. 619* Reed, C.A. 1965. North American Bird Eggs. Dover Publ., Inc. New York, N.Y:372. 758* (ed.)1982. Fish and wildlife. 50 CFR 10 (Code of Federal Regulations). General Serv. Admin. Washington, D.C:7. 1082* Soothill, E., Soothill, R. 1982. Wading Birds of the World. Blandford Press Puole and Dorset, England:232-233. 1169* Bent, A.C. 1927. Life histories of North American shore birds. Part I. U.S. Natl. Mus. Bull. No. 142:420. 3606* Cogswell, H.L. 1977. WATER BIRDS OF CALIFORNIA. University of California Press Berkeley, CA:334. 3607* Gooders, J. 1979. BIRDS OF MARSH AND SHORE. Orbis Publishing London, England:264. 3609* Bent, A.C. 1962. LIFE HISTORIES OF NORTH AMERICAN SHORE BIRDS. 13. Dover Publications, Inc. New York:420. 4182* Recher, H.F. 1966. Some aspects of the ecology of migrating shorebirds. Ecology 47:393-407. 4469* Fredrickson, L.H.. Personal communication.. 4652* Smith, J... 4684* Rundle, W.D. 1980. Managment, habitat selection, and feeding of migrant rails and shorebirds.:228. 4685* Rundle, W.D., Fredrickson, L.H. 1981. Managing seasonally flooded impoundments for migrant rails and shorebirds. Wildlife Society Bulletin 9(2):80-87. 6069* Silliman, J., Mills, G.S., Alden, S. 1977. Effects of flock size on foraging activity in wintering sanderlings. Wilson Bull. 89:434-438. References - 1