(DRAFT) - Taxonomy
                                 Species tern, royal
                                 Species Id M040187
                                   Date 26 AUG 96



TAXONOMY

NAME - tern, royal OTHER COMMON NAMES - ELEMENT CODE - 03/31/84 AOU CODE - 06/04/85 03/30/88 CATEGORY - Birds PHYLUM AND SUBPHYLUM - Chordata, Vertebrata CLASS AND SUBCLASS - Aves, Neornithes ORDER AND SUBORDER - Charadriiformes, FAMILY AND SUBFAMILY - Laridae, Sterninae GENUS AND SUBGENUS - Thalasseus, SPECIES AND SSP - maximus, maximus SCIENTIFIC NAME - Thalasseus maximus maximus AUTHORITY - Boddaert TAXONOMY REFERENCES - 404, 406 and 575 COMMENTS ON TAXONOMY - Scientific name changed--genus: Sterna to Thalasseus; species: maxima to maximus *406*; superorder: Neognathae *404* Taxonomy - 1
                                  (DRAFT) - Status
                                 Species tern, royal
                                 Species Id M040187
                                   Date 26 AUG 96



STATUS

Coded Status Federal Migratory See Comments State endangered Existing, FMP exists REFERENCES FOR STATUS - 4447 and 402 COMMENTS ON STATUS - Because the Royal tern's nesting habitat constitutes a fragile environment and the breeding colonies are so densely packed, the population is vulnerable to both natural forces such as erosion and flooding from high tides *402,1168,545*, and man-caused disasters such as oil spills *402* Breeds along the Eastern Shore. Common to abundant summer resident on coast, rare visitor farther inland. Status - 1
                               (DRAFT) - Distribution
                                 Species tern, royal
                                 Species Id M040187
                                   Date 26 AUG 96



DISTRIBUTION

References on County Occurrence - 001, 131, 810, CB1, CB2, CB3, 133, 103, 119, 115, 073, 199, 095, 181, 093, 650, 700, 800, 550 and 740 References on Seasonal Occurrence - Common to abundant summer resident on coast, in Chesapeake Bay, and on lower tidal rivers *700*; rare visitor farther inland, usually late summer and fall *700*; uncommon to rare winter visitor, usually in extreme southeast *700* REFERENCES FOR HYDROLOGIC UNIT CODES - 001, 131, 810, 133, 103, 119, 115, 073, 199, 095, 181, 093, 650, 700, 800, 550 and 740 OTHER DISTRIBUTION REFERENCES FOR OTHER DISTRIBUTION - H Comments on Distribution - Common to abundant summer resident on coast. Rare visitor farther inland. Uncommon to rare winter visitor. Only three records away from coast.*8511* Distribution - 1
     

HABITAT ASSOCIATIONS

LAND USE - Barren Land Beaches REFERENCES FOR LAND USE - 1168 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine, intertidal BB2 P REFERENCES FOR NWI - 1168, 450 and 545 COMMENTS ON HABITAT ASSOCIATIONS - Favored nesting habitats are sandy islands; either natural or spoil *1168,450*; barren or sparsely vegetated *402,1168,545*; with the nesting site at an elevation above high tide level of 2-3 feet *1168*; or 1-2 feet *545*; preferred nesting habitat is isolated and free of mammalian predators *402,487,450*; associate on breeding grounds with common terns *487*; black skimmers *487,450*, laughing gulls *450*, and sandwich terns *1168,450* POTENTIAL NATURAL VEGETATION - 065 Northern Cordgrass Prairie (Distichlis-Spartina) 101 Oak - Hickory - Pine Forest (Quercus-Carya-Pinus) 103 Southern Floodplain Forest (Quercus-Nyssa-Taxodium) REFERENCES FOR PNV - 700 ECOREGION - Southeastern Mixed Forest: Flat Plains REFERENCES FOR ECOREGION - 700 Habitat Associations - 1
                                (DRAFT) - Food Habits
                                 Species tern, royal
                                 Species Id M040187
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - Carnivore REFERENCES FOR TROPHIC LEVEL - 1168, 487 and 450 LIFESTAGE FOOD FOOD PART General Malacostraca Not Specified General See Comments; Food See Comments Important See Comments; Food See Comments Juvenile See Comments; Food See Comments Adult Malacostraca Not Specified Adult See Comments; Food See Comments REFERENCES FOR GENERAL FOOD - 1168, 487, 450 and 545 REFERENCES FOR IMPORTANT FOOD - 1168 REFERENCES FOR ADULT FOOD - 1168, 487, 450 and 545 REFERENCES FOR JUVENILE FOOD - 402, 487 and 2579 COMMENTS ON FOOD - Small fish *1168,487,450*; royal terns dive for their food which is primarily small fish less than 4 inches *1168* COMMENTS ON ADULT FOOD - Small fish *1168,487,450*; because of large size, greater diversity in prey base *487* COMMENTS ON JUVENILE FOOD - Parents provide food for young *402,487*, into the following winter *2579* Food Habits - 1
                         (DRAFT) - Environment Associations
                                 Species tern, royal
                                 Species Id M040187
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Coastal Features: Sand bars G Water Level: Seasonal/Alternatiely flooded areas G Coastal Features: Sandy offshore islands G Water Depth Preference: Specified in Comments G Human Association: Specified in Comments LIM Water Level: Specified in Comments LIM Human Association: Specified in Comments E Water Level: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 1168, 487 and 545 REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 1168, 450 and 545 REFERENCES FOR EGG ENVIRONMENTAL ASSOC_ - 1168 and 545 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Feeding primarily in shallow bays and inlets *487,450*; frequently at some distance from colony *487*; because breeding site is near high tide levels, an abnormally high tide can wash away eggs and/or young *1168,545*, but royal terns are known to renest *1168*; for colony site location, isolation is important if not essential *450* COMMENTS ON EGG ENVIRONMENTAL ASSOC_ - An abnormally high tide can wash away the eggs *1168,545* Environment Associations - 1
                               (DRAFT) - Life History
                                 Species tern, royal
                                 Species Id M040187
                                   Date 26 AUG 96



LIFE HISTORY

ORIGIN: native *404*; REPRODUCTION: 1-2 eggs/once a season unless renesting necessary because of nest destruction *1168*; BEHAVIOR: 2-3 days after hatch, young form 'creche' *450*; young fledge at about 30 days *450*; both adults incubate, feed young *487,450*, defend nest *487*; Nest site: shallow depression in the sand, with little or no lining, in a densely packed colony *1168*; fall departure for Virginia in mid-September, linger in North Carolina until end of November *1168*; winter from North and South Carolina to Caribbean area and south to Uruguay and central Argentina *404*; AQUATIC/TERRESTRIAL ASSOCIATIONS: Larus atricilla, Thalasseus sandivicensis *1168,450* COMMENTS ON LIFE HISTORY - Laughing gulls are a major source of royal tern egg mortality *450*; royal terns and sandwich terns nest together in densely packed colonies *1168,450* and their young 'creche' together once hatched *1168,450* REFERENCES FOR LIFE HISTORY- 1168, 487, 450 and 404 Life History - 1
                           (DRAFT) - Management Practices
                                 Species tern, royal
                                 Species Id M040187
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Controlling pollution [thermal, chemical, physical] Beneficial Restricting/regulating human use of habitats Beneficial Restricting/regulating human disturbance of populations Beneficial Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 402 and 450 COMMENTS ON MANAGEMENT PRACTICES - (B)=Preservation of undisturbed nesting habitat is of primary importance *402,450* Management Practices - 1
                                   (DRAFT) - References
                                   Species tern, royal
                                    Species Id M040187
                                      Date 26 AUG 96



     

References

402* Akers, J.W. 1979. Royal tern. Proc. Symp. on Endangered and Threatened Plants and Animals of Virginia Linzey, D.W. Ext. Div., Virginia Polytechnic Institute and State Univ. Blacksburg:454-455. 404* Union, American Ornithologists'. 1957. Check-list of North American Birds. American Ornithologists' Union Baltimore, Md:691. 406* Union, American Ornithologists', Nomencl., Comm. on Classif. and. 1976. Thirty-third supplement to the AOU check-list of North American birds. Auk 93:875-879. 450* Buckley, F.G., Buckley, P.A. 1972. The breeding ecology of royal terns Sterna (Thalasseus) maxima maxima. Ibis 114:344-359. 487* Erwin, R.M. 1977. Foraging and breeding adaptations to different food regimes in three seabirds: the common tern Sterna hirundo, royal tern Sterna maxima, and black skimmer Rynchops niger. Ecology 58:389-397. 545* Kale, H.W., II, Sciple, G.W., Tomkins, I.R. 1965. The royal tern colony of Little Egg Island, Ga. Bird-Banding 12:21-27. 575* Mayr, E. 1969. Principles of Systematic Zoology. McGraw-Hill New York, N.Y:428. 700* Ornithology, Virginia Society of. 1979. Virginia's Birdlife: An Annotated Check-list. Virginia Avifauna No. 2. Virginia Society of Ornithology Lynchburg, Va:118. 1168* Bent, A.C. 1921. Life histories of North American gulls and terns. U.S. Mus. Nat. Hist. Bull. 113. Reprint 1963 by Dover Pubs. New York, N.Y:337. 2579* Ashmole, N.P., Tovar, J. Humberto. 1968. Prolonged parental care in royal terns and other birds. Auk 85:90-100. 4447* Service, U.S. Fish & Wildlife. 1985. 50(66):15. References - 1