(DRAFT) - Taxonomy
                                     Species elk
                                 Species Id M050035
                                   Date 26 AUG 96



TAXONOMY

NAME - elk OTHER COMMON NAMES - elk, Roosevelt, deer, red and wapiti ELEMENT CODE - CATEGORY - Mammals PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - , FAMILY AND SUBFAMILY - , GENUS AND SUBGENUS - Cervus, SPECIES AND SSP - elaphus, roosevelti SCIENTIFIC NAME - Cervus elaphus roosevelti AUTHORITY - TAXONOMY REFERENCES - 232 Taxonomy - 1
                                  (DRAFT) - Status
                                     Species elk
                                 Species Id M050035
                                   Date 26 AUG 96



STATUS

Coded Status Migrant See Comments REFERENCES FOR STATUS - 232 COMMENTS ON STATUS - The Alaska Department of Fish and Game manages elk, which are migratory *232*. Status - 1
                               (DRAFT) - Distribution
                                     Species elk
                                 Species Id M050035
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - TERRESTRIAL REFERENCES FOR HABITAT - 232 LAND USE - Forest Land Evergreen Forest Land Snow or Ice: All Classes REFERENCES FOR LAND USE - 232 COMMENTS ON HABITAT ASSOCIATIONS - The subspecies Roosevelt elk inhabitats the timbered Pacific coast of Washington, Oregon, northwestern California. Where available, spruce-timbered areas are used by elk most during December through March, apparently because they provide extra cover and because snow depths are less *232*. Feeding Locations: There is considerable overlap in the use of habitat during spring, winter, and fall, depending on snow conditions. Spring: Alder (Alnum spp.) communities. Summer: Alpine habitat. Fall: Alder communities. Winter: Spruce (Picea sitchensis); timbered areas typically on south and southeast-facing slopes adjacent to beach fringes where available. Elk wintering on Tonki and Raspberry islands use heath-type habitat, which grows at or near sea level on windblown capes and bluffs *232*. Reproductive Habitat: Most calving occurs at lower elevation in spruce timber *232*. ANIMAL/PLANT SPECIES ASSOCIATIONS - Brown bear Sitka black-tailed deer REFERENCES FOR SPECIES ASSOCIATIONS - 232 COMMENTS ON SPECIES ASSOCIATIONS - Predators: Brown bears are the only animal on the islands capable of preying on elk. Although preadation of elk by brown bears currently does not appear to be a major mortality factor, the killing of an adult female elk by a brown bear has been documented. A study has observed on two occasions brown bears that were apparently stalking elk herds *232*. Sitka black-tailed deer (Odocoileus hemionus sitkensis) feed on many of the same food plants as elk on Afognak, which could lead to increasing competition as populations increase *232*. POTENTIAL NATURAL VEGETATION - 115 Alder thickets (Alnus) 099 Northern Hardwoods - Spruce Forest (Acer-Betula-Fagus REFERENCES FOR PNV - 232 Habitat Associations - 1
                                (DRAFT) - Food Habits
                                     Species elk
                                 Species Id M050035
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - HERBIVORE/BROWSER HERBIVORE/GRAZER REFERENCES FOR TROPHIC LEVEL - 232 LIFESTAGE FOOD FOOD PART General Plants Not Specified General Onagraceae Not Specified General Leguminosae Not Specified General Rosaceae Not Specified General Umbelliferae Not Specified General Umbelliferae Not Specified General Corn Not Specified General Caryophyllaceae Not Specified General Poaceae Not Specified Important Caprifoliaceae Not Specified General Caprifoliaceae Not Specified Important Salicaceae Not Specified General Salicaceae Not Specified General Cyperaceae Not Specified General Sphenopsida Not Specified General Pteropsida Buds General Araliaceae Buds REFERENCES FOR GENERAL FOOD - 232 REFERENCES FOR IMPORTANT FOOD - 232 COMMENTS ON FOOD - Relatively little quantitative information exists concerning seasonal diets of elk on Afognak Island. Food preferences of these elk have been found to be quite diverse. There is a great abundance of vascular plants on Afognak Island; because of this, the elk's diet includes a large variety of succulent and woody plant species *232*. During summer feeding observations conducted on southwestern Afognak, the following plants were found to be utilized by grazing elk: fireweed (Epilobium angustifolium), lupine (Lupinus nootktensis), burnet (Sanguisorba sitchensis), cow parsnip (Heracleum lanatum), sea lovage (Lipusticum hultenii), hellebore (Veratrum eschscholtzii), and water chickweed (Montia spp.) *232*. Consumption of grasses and forbs begins to decrease with the arrival of fall because then species become dry and apparently less palatable. Gradually the fall diets of elk begin to include a few perrenial forbs, notably fireweed, and an abundance of browse species, including elderberry (Sambucus racemosa) and highbush cranberry (Viburnum edule) *232*. During winter months elk on Afognak feed almost exclusively on browse, with Food Habits - 1 (DRAFT) - Food Habits Species elk Species Id M050035 Date 26 AUG 96 elderberry and willow (Salix spp.) the most important species. During late winter, elk have been observed to heavily utilize the cured tops of hairgrass (Deschampsia caespitosa), which occurs in scattered stands in some of the lowland valleys, but grasses are not generally regarded as a major food item in the winter diet of these elk *232*. During spring, the consumption of grasses and grasslike plants and perennial forbs increases with the development of new growth. Sedges (Carex spp.), horsetail (Equisetum spp.), wild celery (Angelica lucida), bluejoint (Calamagrostis canadensis), and cow parsnip are also heavily utilized, as are the buds of fern (Anthyrum cyclosorum) and devil's club (Oplopanax horridus) *232*. Feeding Locations: There is considerable overlap in the use of habitat during spring, winter, and fall, depending on snow conditions. Spring: Alder (Alnum spp.) communities. Summer: Alpine habitat. Fall: Alder communities. Winter: Spruce (Picea sitchensis); timbered areas typically on south and southeast-facing slopes adjacent to beach fringes where available. Elk wintering on Tonki and Raspberry islands use heath-type habitat, which grows at or near sea level on windblown capes and bluffs *232*. Factors Limiting Availability of Food: Heavy snowfall and overgrazing by elk can sufficiently reduce forage to cause malnutrition, resulting in severe winter mortality *232*. Food Habits - 2
                         (DRAFT) - Environment Associations
                                     Species elk
                                 Species Id M050035
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Coniferous Forest: Specified in Comments FA Hardwood Forest: Alder FA Elevation: Specified in Comments FA Coniferous Forest: Specified in Comments FA Slope: Specified in Comments BA Elevation: Specified in Comments BA Coniferous Forest: Specified in Comments FA Aspect: Southeast FA Aspect: South FA Coastal Features: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 232 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 232 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Aquatic: Because of its widespread availability, water does not appear to be a limiting factor for elk on Afognak Island *232*. Terrestrial: A wide variety of vegetation and terrain can be used by elk as escape cover. Use of escape cover varies seasonally and by sex and reproductive status. Brown bears are the only animal on the islands capable of preying on elk. Although preadation of elk by brown bears currently does not appear to be a major mortality factor, the killing of an adult female elk by a brown bear has been documented. A study has observed on two occasions brown bears that were apparently stalking elk herds. Where available, spruce-timbered areas are used as elk most during December through March, apparently because they provide extra cover and because snow depths are less *232*. COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - Feeding Locations: There is considerable overlap in the use of habitat during spring, winter, and fall, depending on snow conditions. Spring: Alder (Alnum spp.) communities. Summer: Alpine habitat. Fall: Alder communities. Winter: Spruce (Picea sitchensis); timbered areas typically on south and Environment Associations - 1 (DRAFT) - Environment Associations Species elk Species Id M050035 Date 26 AUG 96 southeast-facing slopes adjacent to beach fringes where available. Elk wintering on Tonki and Raspberry islands use heath-type habitat, which grows at or near sea level on windblown capes and bluffs *232*. COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - Most calving occurs at lower elevation in spruce timber *232*. Environment Associations - 2
                               (DRAFT) - Life History
                                     Species elk
                                 Species Id M050035
                                   Date 26 AUG 96



LIFE HISTORY

Reproductive Habitat: Most calving occurs at lower elevation in spruce timber *232*. Reproductive Seasonality: Most calving occurs in early June, although calf tracks have been observed as early as 23 May. Bulging begins in early September and continues through early October. Most breeding activity occurs during the last weeks of September *232*. Age At Sexual Maturity: Female elk can successfully breed at 1.5 years, although the frequency of successful breeding in yearling cows varies among populations and from year to year in the same population. Male elk can successfully breed at 1.5 years, but because of behavioral patterns larger bulls tend to do most of the breeding. Bulls generally are in their prime at ages 6 to 11 *232*. Fecundity: Elk generally produce one calf per year, although twinning is not unknown *232*. Home Range: Home range sizes varied between 13.7 and 39.4 mi2 and averaged 22.8 mi2. Winter home ranges were smaller, ranging from 1.2 to 7.3 mi2 and averaging 3.2 mi2. However, there appears to be some overlap between the ranges of adjacent herds, and some interchange of animals is suspected *232*. Factors Influencing Populations: Natural: Severe winter weather can cause severe losses of elk *232*. Human-related: A summary of possible impacts from human-related activities includes: pollution of water and/or food supply; reduction of food supply; alteration of habitat; disturbance/displacement; barriers to seasonal movement; and overharvest *232*. Predators: Brown bears are the only animal on the islands capable of preying on elk. Although preadation of elk by brown bears currently does not appear to be a major mortality factor, the killing of an adult female elk by a brown bear has been documented. A study has observed on two occasions brown bears that were apparently stalking elk herds *232*. Competition: Sitka black-tailed deer (Odocoileus hemionus sitkensis) feed on many of the same food plants as elk on Afognak, which could lead to increasing competition as populations increase *232*. Special Considerations: 1. Elk on Raspberry and Afognak islands are the result of transplants in 1928 *232*. 2. Drastic declines in elk numbers as a result of severe winters were documented. The decline was from 1,200-1,500 elk in 1965 to an estimated 500 animals by 1976. However, the elk populaton has since increased to over 1,200 animals *232*. Life History - 1 (DRAFT) - Life History Species elk Species Id M050035 Date 26 AUG 96 3. Much of the land occupied by elk is privately owned and may be subject to logging. The effect of logging and associated road access on elk habitat on Afognak and Raspberry islands is not clearly understood *232*. 4. State and borough land sales on Raspberry Island may be detrimental to elk where changes in land use result in habitat deterioration or loss *232*. LIFE HISTORY CODES - Foraging Strategy: Grazing Foraging Strategy: Browsing Breeding/Spawning Season: May Breeding/Spawning Season: June Breeding/Spawning Season: September Breeding/Spawning Season: October Number of Broods/Litters (Reproductive Efforts) Per Ye Average Number of Offspring/Reproductive Effort: 1 Average Number of Offspring/Reproductive Effort: 2 Home Range Size: Greater than 100 acres Home Range Size: 20 - 100 acres REFERENCES FOR LIFE HISTORY- 232 Life History - 2
                           (DRAFT) - Management Practices
                                     Species elk
                                 Species Id M050035
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Establishing/maintaining nesting and escape cover Beneficial Controlling pollution [thermal, chemical, physical] Beneficial Developing/maintaining habitat diversity Beneficial Restricting/regulating human disturbance of populations Adverse Migration barriers Beneficial Regulating harvest of species being described Beneficial Other management practices [specified in comments] Adverse Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 232 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 232 COMMENTS ON MANAGEMENT PRACTICES - Factors Influencing Populations: Human-related: A summary of possible impacts from human-related activities includes: pollution of water and/or food supply; reduction of food supply; alteration of habitat; disturbance/displacement; barriers to seasonal movement; and overharvest *232*. Much of the land occupied by elk is privately owned and may be subject to logging. The effect of logging and associated road access on elk habitat on Afognak and Raspberry islands is not clearly understood *232*. State and borough land sales on Raspberry Island may be detrimental to elk where changes in land use result in habitat deterioration or loss *232*. Management Practices - 1
                                   (DRAFT) - References
                                       Species elk
                                    Species Id M050035
                                      Date 26 AUG 96



     

References

232 * State of Alaska Department of Fish, and Game. 1986. Alaska Habitat Management Guide: Life Histories and Habitat Requirements of Fish and Wildlife. (ed.). Alaska Department of Fish and Game Juneau, Alaska:763. References - 1