(DRAFT) - Taxonomy
                               Species oyster, Pacific
                                 Species Id M060003
                                   Date 26 AUG 96



TAXONOMY

NAME - oyster, Pacific OTHER COMMON NAMES - Pacific oyster ELEMENT CODE - CATEGORY - Aquatic Molluscs PHYLUM AND SUBPHYLUM - Mollusca, CLASS AND SUBCLASS - Bivalvia, ORDER AND SUBORDER - Ostreoida, FAMILY AND SUBFAMILY - Ostreidae, GENUS AND SUBGENUS - Crassostrea, SPECIES AND SSP - gigas, SCIENTIFIC NAME - Crassostrea gigas AUTHORITY - Thunberg, 1793 TAXONOMY REFERENCES - 186 COMMENTS ON TAXONOMY - Other common names include Giant oyster, Japanese oyster, and Giant Pacific oyster *10*. Taxonomy - 1
                                  (DRAFT) - Status
                               Species oyster, Pacific
                                 Species Id M060003
                                   Date 26 AUG 96



STATUS

Coded Status Commercial/consumption Commercial REFERENCES FOR STATUS - 10 COMMENTS ON STATUS - The commercial fishery for this species has grown rapidly since its introduction from Japan to the west coast of the United States in 1903. The United States now consumes almost 60 percent of the world's total oyster production. The Pacific oyster is a dominant shellfish in a growing United States aquiculture industry along the Pacific Coast. 60-70 percent of the Pacific oyster production is marketed in the Pacific Coast States. Washington State leads all others for the production of Pacific oysters which comprise 25 percent of all shellfish landings in Washington State. The landed dollar value of this species in Washington State was 3.8 million in 1982 *10*. Status - 1
                               (DRAFT) - Distribution
                               Species oyster, Pacific
                                 Species Id M060003
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine Marine, intertidal V REFERENCES FOR NWI - 10 ANIMAL/PLANT SPECIES ASSOCIATIONS - Mytilicola orientalis Cliona celata Plydora ciliata Vibrio anguillarum Protogonyalax catenella Gonyaulax catenella Ceratium fusus Gymnodinium splendices REFERENCES FOR SPECIES ASSOCIATIONS - 10 Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species oyster, Pacific
                                 Species Id M060003
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - FILTER FEEDER REFERENCES FOR TROPHIC LEVEL - 10 LIFESTAGE FOOD FOOD PART General Microorganisms Not Applicable General Phytoplankton Not Applicable General Protozoans Not Applicable General Bacteria Not Applicable REFERENCES FOR GENERAL FOOD - 10 COMMENTS ON FOOD - They consume phytoplankton, bacteria and protozoa which are filtered from the water *10*. Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species oyster, Pacific
                                 Species Id M060003
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Temperature: Between 21-27 degrees C G Water Temperature: Between 15-21 degrees C G Water Temperature: Below 15 degrees C G Water Temperature: Specified in Comments G Relation to Substrate: Attached - normally sessile G Relation to Substrate: Specified in Comments A Substrate: Specified in Comments A Turbidity: Clear water but tolerate periodic cloudiness REFERENCES FOR ENVIRONMENTAL ASSOC_ - 10 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 10 REFERENCES FOR RESTING JUVENILE ENVIRONMENTAL ASSOC_ - 10 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Pacific oysters live and grow in water temperatures from 4-24 degrees C although they are able to survive temperatures as low as -4 degrees C when exposed by high tide. They begin dying at salinities of 10 ppt. High mortalities result from siltation. Excessive turbidity can tumble the oysters and knock off their fragile shell edges or stir up bottom sediments causing silt accumulation over the oyster *10*. COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - The optimum temperature for water transport through adult oysters is 20 degrees C. The optimum salinity for water transport is 25-35 ppt. At 13 ppt little water is pumped through the gills and at 10.5 ppt harmful effects are apparent *10*. COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - The gametes are not released until the water temperature reaches 19 degrees C *10*. COMMENTS ON RESTING LARVAE ENVIRONMENTAL ASSOC_ - The larvae can survive temperatures below 15 degrees C for short periods of time and can survive temperatures as high as 30 degrees C. The larval period increases from 18 days at 72 degrees F to 30 days at 64 degrees F. Larval setting increases as the temperature increases between 15 and 30 degrees C, but declines at 35 degrees C. The larvae were unaffected by Environment Associations - 1 (DRAFT) - Environment Associations Species oyster, Pacific Species Id M060003 Date 26 AUG 96 constant salinities from 16-34 ppt. The optimum conditions for setting are 30 degrees C with salinities of about 30 ppt. Larval mortality was 27 percent at .25 g/l and 27 percent at .50 g/l *10*. Environment Associations - 2
                               (DRAFT) - Life History
                               Species oyster, Pacific
                                 Species Id M060003
                                   Date 26 AUG 96



LIFE HISTORY

Physical Description: The shell of this species is extremely rough, extensively fluted, and laminated. The lower valve is deeply cupped and the upper valve is generally flat. The shells are usually whitish with many purple streaks and spots radiating away from the umbo. The shape of the shell varies with the environment. The interior of the shell is white, with a single muscle scar that is sometimes dark, but never purple or black. The maximum length is ten inches, but normally the length is from 4-6 inches. It differs from Crassostrea virginica in never having a purple or black muscle scar, and from Ostrea lurida in its extremely large size and heavy shell. In addition, the inside of an O. lurida shell is iridescent green *10*. Reproduction: Pacific oysters are protandrous hermaphrodites. They change sex, but their timing is erratic and seasonal. The young are functionally male during their first spawning, while about half remain male for their second spawning. The adults function as seperate male or female animals in any given reproductive season, but a change of sex from male to female often occurs at some point in life. Sexual maturity is reached during the first year. They spawn annually in certain coastal waters of British Columbia when the water temperature is approximately 19.5 degrees C. Oysters carry spawn during July and August. Once the spawning temperature has been reached, usually in July or August, spawning is synchronous. Under natural conditions, simultaneous release of sperm and eggs into the water column is attained through mutual stimulation. This species is highly prolific and an average market-sized female (3 inches long) can produce 50-100 million eggs in a single spawning. The fertilized eggs are spherical, and 45-62 um in diameter. The egg is multilayered, with membranes that divide the jellylike outer coat from the small nucleus. The germinal vesicle is eccentrically located within the nucleolus. Temperature plays a major role in the development of the fertilized oyster egg. A decrease from 24.5 degrees C to 22.5 degrees C doubled the time required for the formation of trochophore larvae. Fertilized eggs reach the veliger stage in 72 hours at 14 degrees C, and in 28 hours at 22 degrees C *10*. Behavior: The pelagic larvae are planktotrophic, feeding on phytoplankton and growing over a period of 2-3 weeks. They are freeswimming in the larval stage and when they reach a length of about .30 mm they attach to the bottom or hard surface as spat. When contact is made with the foot, the larvae crawls onto the surface, and if suitable, attaches by the left valve. The adult is a filter feeder and the various particles are sorted by size using mucus secretions. They ingest bacteria, protozoa, a wide variety of phytoplankton, larval forms of other invertebrate animals, detritus and some inorganic material. Crabs and starfish can be serious predators on the young and adults. They are also preyed on by the Japanese oyster drill (Ocinebra japonica) and by the predatory flatworm (Pseudostylochus ostreophagus). Mud shrimp also dig burrows that make the beds too soft for the oyster culture *10*. Origin: This species was introduced from Japan to the western coast of the United States in 1903. Limiting factors: This species is limited by temperatures between 4-24 dregrees C, long periods of exposure to air and dinoflagellate blooms. Crabs and starfish can be serious predators on young and adults. Parasites include Mytilicola orientalis (copepod) boring sponges (Cliona celata) and sea worms (Plydora ciliata). They are not greatly affected by disease although there are occasional outbreaks of focal necrosis Life History - 1 (DRAFT) - Life History Species oyster, Pacific Species Id M060003 Date 26 AUG 96 (bacterium) and, bacillary necrosis (Vibrio anguillarum)*10*. Population parameters: This species can produce from 50-100 million eggs in a single spawning. Sexual maturity is reached during the first year *10*. Aquatic/terrestrial associations: Predators include crabs and starfish (Japanese oyster drill, Dungeness crab, rock crab and graceful crab. Mud shrimp dig burrows that retain water at low tide and make the beds too soft for oyster culture *10*. REFERENCES FOR LIFE HISTORY- 10 Life History - 2
                           (DRAFT) - Management Practices
                               Species oyster, Pacific
                                 Species Id M060003
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Adverse Wastewater disposal/assimilation Beneficial Predator control Beneficial Controlling undesirable invertebrate species Existing Applying pesticides REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 10 REFERENCES FOR EXISTING MANAGEMENT PRACTICES - 10 COMMENTS ON MANAGEMENT PRACTICES - A major long-term concern to the oyster industry in the Pacific Northwest is the present and potential future loss to water quality because of pollution (sewage, industrial waste and pulp mill effluents). Starfish can be controlled by removal. The Japanese oyster drill can be controlled somewhat by collecting the egg cases before they hatch and careful oyster farming practices. The pesticide Sevin has been used controversially as a control agent for ghost shrimp in Washington *10*. Management Practices - 1
                                   (DRAFT) - References
                                 Species oyster, Pacific
                                    Species Id M060003
                                      Date 26 AUG 96



     

References

10* Pauley, G.B., B. Van Der Raay, and D. Troutt. 1988. Species Profiles: life histories and environmental requirements of coastal fishes and invertebrates (Pacific Northwest)--Pacific oyster. U.S. Fish and Wildl. Serv. Biol. Rep. 82(11.85). U.S. Army Corps of Engineers, TR EL-82.4. 28pp. 186 * Turgeon, D.D., A.E. Bogan, E.V. Coan, W.K. Emerson, W.G. Lyons, W.L. Pratt, C.F.E. Roper, A. Scheltema, F.G. Thompson, J.D. Williams. 1988. Common and scientific names of aquatic invertebrates from the United States and Canada: mollusks. References - 1