(DRAFT) - Taxonomy
                                Species conch, queen
                                 Species Id M060165
                                   Date 26 AUG 96



TAXONOMY

NAME - conch, queen OTHER COMMON NAMES - queen conch and pink conch ELEMENT CODE - CATEGORY - Aquatic Molluscs PHYLUM AND SUBPHYLUM - Mollusca, CLASS AND SUBCLASS - Gastropoda, Prosobranchia ORDER AND SUBORDER - Mesogastropoda, FAMILY AND SUBFAMILY - Strombidae, GENUS AND SUBGENUS - Strombus, SPECIES AND SSP - gigas, SCIENTIFIC NAME - Strombus gigas AUTHORITY - Linnaeus TAXONOMY REFERENCES - 186 and 213 COMMENTS ON TAXONOMY - The common names, queen or pink conch, come from the beautiful color and graceful lines of the shell. There are 55 species of Strombus conchs, six of which occur in Caribbean and Florida waters *213*. Taxonomy - 1
                                  (DRAFT) - Status
                                Species conch, queen
                                 Species Id M060165
                                   Date 26 AUG 96



STATUS

Coded Status Commercial Existing, FMP exists Commercial/consumption Commercial/bait Other federal or state designation See Comments REFERENCES FOR STATUS - 213 COMMENTS ON STATUS - Conch is one of the most important export fisheries in the Caribbean *213*. In 1985, Florida banned all harvest, killing, molestation, harming or mutilation of queen conch in Florida waters. Beginning in December 1986, the rule was extended so now harvest by all persons in Florida-registered vessels in federal waters contiguous to Florida waters is banned. As of 1989, it is unlawful to take or harvest any queen conch from the land or waters of Florida or to possess or transport any queen conch so taken or harvested *213*. The soft parts of the body are used for fish bait or are discarded *213*. The well-developed foot muscle of the queen conch provides the sweet white meat so highly prized. Excluding the shell, 35% of the animal's weight is marketable meat *213*. Status - 1
                               (DRAFT) - Distribution
                                Species conch, queen
                                 Species Id M060165
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC REFERENCES FOR HABITAT - 213 LAND USE - Water Atlantic Ocean Coastal Waters REFERENCES FOR LAND USE - 213 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine AB. Marine RF. Marine RF1 Marine OW0 REFERENCES FOR NWI - 213 COMMENTS ON HABITAT ASSOCIATIONS - Adults are found most often in seagrass meadows and hard-bottom communities, but may also be seen on the sandy sea floor, coral rubble and reefs. The larval stage drifts and swims in the ocean currents, feeding on phytoplankton *213*. ANIMAL/PLANT SPECIES ASSOCIATIONS - pink cardinal fish slipper limpets REFERENCES FOR SPECIES ASSOCIATIONS - 213 COMMENTS ON SPECIES ASSOCIATIONS - Other sea creatures often share the large shell of the conch. Pink cardinal fish, or conchfish, may live in the space between the shell and the animal. As many as five may be present. A small crab also lives commensally with the conch, making its home wedged between the shell and the foot. Slipper limpets are sometimes found securely attached to the shell or operculum *213*. Habitat Associations - 1
                                (DRAFT) - Food Habits
                                Species conch, queen
                                 Species Id M060165
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - DETRITIVORE HERBIVORE/GRAZER REFERENCES FOR TROPHIC LEVEL - 213 LIFESTAGE FOOD FOOD PART General Microorganisms Not Applicable General Bacteria Not Applicable General Algae Not Applicable General See Comments; Food Not Applicable Adult Microorganisms Not Applicable Adult Algae Not Applicable Adult Detritus - Organic Not Applicable Adult See Comments; Food Not Applicable Juvenile Microorganisms Not Applicable Juvenile Algae Not Applicable Juvenile Detritus - Organic Not Applicable Juvenile See Comments; Food Not Applicable Larva Microorganisms Not Applicable Larva Phytoplankton Not Applicable Larva See Comments; Food Not Applicable REFERENCES FOR GENERAL FOOD - 213 REFERENCES FOR ADULT FOOD - 213 REFERENCES FOR JUVENILE FOOD - 213 REFERENCES FOR LARVAE FOOD - 213 COMMENTS ON FOOD - The snout-like mouth can be extended to graze on small algae growing on rocks and dead coral, or to ingest algae- or bacteria- covered sand grains *213*. All species of stromboid conchs in the Caribbean are herbivore and detritus feeders *213*. COMMENTS ON ADULT FOOD - Adults feed on algae and algal detritus *213*. COMMENTS ON JUVENILE FOOD - Juvenile conchs, called "rollers", graze on the algae attached to seagrasses and detritus *213*. COMMENTS ON LARVAE FOOD - Veligers feed on phytoplankton drifting in the sea. Their ciliated lobes capture tiny, single-celled plants *213*. Food Habits - 1
                         (DRAFT) - Environment Associations
                                Species conch, queen
                                 Species Id M060165
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Depth Preference: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 213 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Queen conchs are found in depths down to nearly 500 feet, but are most abundant in waters ranging from the shallows to 70 feet *213*. Environment Associations - 1
                               (DRAFT) - Life History
                                Species conch, queen
                                 Species Id M060165
                                   Date 26 AUG 96



LIFE HISTORY

DESCRIPTION The body consists of a black-speckled, long and narrow foot, a "head" with colorful yellow eyes on the end of two protruding stalks, and a snout-like probosis between them, plus a pair of slender, sensitive tentacles. The orange or yellow mantle resembles a close-fitting cover enclosing the foot and head. The respiratory organs are feathery gill combs in the mantle which remove oxygen from the water. Eye development in queen conch is more complex and keener than most other gastropods *213*. The foot bears a horny, sickle-shaped operculum that digs into the bottom as the conch hurtles itself forward. It moves about half its body length with each leap. The operculum is too small and curved to seal the shell opening completely when the animal retreats into its shell, so the conch is not able to "close" its shell as well as other snails do. The operculum is used mainly to assist in locomotion and righting the shell when it is turned over *213*. An adult conch has a "heavy" shell with a broad, flaring lip. A blunt crown of spines projects from each whorl of the spiral shell as they descend from the apex. The shell coils to the right when viewed with the apex nearest the eye *213*. The broad, flaring lip of a mature conch has a "stromboid notch," which is a deep, upward ripple of shell at the end opposite the apex. When the animal emerges from its shell, one eyestalk protrudes from this notch and the other from the opening at the end of the spiral column. The living shell is covered with a brown papery covering called the periostracum, which scales off when the shell is dried. Ther periostracum, which collects silt, bacteria and algae, contributes to the mossy-rock look of conchs resting on the sea floor *213*. Young conch shells, called "rollers", do not have the flaring lip of adults. The shell is more fragile, the spines are well-developed, more pointed and not yet blunted. When the lip is first laid down, it is thin and delicate. Boring sponges and other creatures can cause the lip to crumble. The conch then directs its energies to renew and repair the lip, so it becomes thicker. As time goes by, the animal continues to strengthen its shell and flaring lip from the inside of the shell, increasing the shell thickness and decreasing the shell volume. The shell is composed mainly of calcium carbonate, excreted by the mantle as a fluid which hardens as crystals*213*. GROWTH AND MATURATION Queen conchs are among the largest gastropods in the sea, though males tend to be slightly smaller than females. Shells measuring up to 12 inches have been found. The shell length increases at an average rate of nearly three inches (75 mm) a year in its active growing stage. With maturity, the shell thickens and the increase in length slows or ceases *213*. FEEDING The snout-like mouth can be extended to graze on small algae growing on rocks and dead coral, or to ingest algae- or bacteria- covered sand grains *213*. EGGS Egg masses are excreted as tightly wound gelatinous strings of capsules Life History - 1 (DRAFT) - Life History Species conch, queen Species Id M060165 Date 26 AUG 96 containing more than a half-million embryos. The tube is folded back and forth on itself, producing a long, slightly curved mass. From 70 to more than 129 feet of tube strands may be extruded, at a rate of about five feet per hour. Conchs spawn mainly in the warmer months, although egg masses have been observed throughout the year. Larval veligers emerge from the egg case after three to five days *213*. LARVAE The newly hatched , microscopic larva, called a veliger, has two ciliated "wings" (lobes) used for swimming. The veliger has a tiny, transparent shell (protoconch) which will eventually form the familiar shell. By the sixth day of life, the veliger has four wing-like lobes; by the twelfth, six. It drifts and swims in the ocean currents, feeding on phytoplankton Velagers may travel great distances, moved by ocean currents. The currents disburse the larvae throughout their geographic range *213*. After 21 days, the veliger settles to the sea floor, the lobes disappear, and the "foot" and proboscis development continues. It takes about a month to develop into a shelled animal that begins to resemble the adult form. JUVENILE Queen conchs measure about three inches long at one year and five inches by the second year. Juvenile conchs, called "rollers", graze on the algae attached to seagrasses and detritus *213*. The broad, flaring lip of the shell begins to develop after two-and-a-half to three years, when it is about eight or ten inches long *213*. ADULTS Adults may wander for miles. Tagged adults have moved as much as one mile over a two month period, and more than 700 yards in a week. They gather in large colonies in shallow water in the summer through the fall months to spawn. The animal reaches sexual maturity only after the lip is well-developed; in terms of meat production, this stage is the optimum time for harvest. From this point, the shell does not grow significantly in size and length; the conch energies are directed to strengthening the shell and lip, so it becomes thicker and heavier and the space occupied by the animal actually becomes smaller. Older, heavier conchs were once believed to be a different species and were called "sambas" *213*. REPRODUCTION Adults gather in large colonies in shallow water in the summer through the fall months to spawn. Queen conchs mate in shallow waters in sandy areas behind the reefs. The sexes are separate. Fertilization is internal. During mating the male sits behind the female, depositing sperm via the black, spade-like verge which is inserted into the female. The female retains sperm for several weeks, releasing it to fertilize the eggs when they are laid. Some may spawn as many as six or eight times a season *213*. PREDATION In the larval stages, queen conchs are preyed on by other small, swimming creatures. Juveniles are eaten by crabs, sharks and rays, loggerhead turtles, groupers and snappers, and octopus. After two years man is probably the most important predator, fishing for conch to the point that some Caribbean populations are threatened *213*. Life History - 2 (DRAFT) - Life History Species conch, queen Species Id M060165 Date 26 AUG 96 FEEDING HABITS The conch digestive system includes a crystalline style, a flexible rod about five inches long composed of a microprotein gel. As the style rotates against the gastric shield of the digestive tract, enzymes are released that are capable of digesting the cellulose of the plants consumed. In addition, a salivary gland and esophageal pouches digest cellulose *213*. LIFE HISTORY CODES - Foraging Strategy: Grazing Foraging Sites: Herbaceous vegetation Average Number of Offspring/Reproductive Effort: Grea Number of Broods/Litters (Reproductive Efforts) Per Ye REFERENCES FOR LIFE HISTORY- 213 Life History - 3
                           (DRAFT) - Management Practices
                                Species conch, queen
                                 Species Id M060165
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Regulating harvest of species being described Beneficial Prohibiting harvest of species being described Existing Regulating harvest of species being described Existing Prohibiting harvest of species being described Existing Mariculture activities Existing Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 213 REFERENCES FOR EXISTING MANAGEMENT PRACTICES - 213 COMMENTS ON MANAGEMENT PRACTICES - MANAGEMENT PROBLEMS There have been many management problems in the efforts to protect this significant fishery. In the Bahamas, the fishery is a diffuse, "cottage" fishery, with many small, independent businesses involved, difficult to oversee. Shell disposal is a problem; cleaned shells, dumped at the docks, can clog harbors. (Nassau, Bahamas, imposes fines for shell dumping.) In other areas, the fishery is run by fishing cooperatives with large processing plants. All conch are cleaned at sea and only the body brought ashore *213*. The animal reaches sexual maturity only after the lip is well-developed; in terms of meat production, this stage is the optimum time for harvest *213*. FISHING GEAR In the Caribbean, queen conchs were once fished from dinghies by scanning the bottom with glass-bottom buckets, then retrieving the animals with long, hooked poles. Now divers use SCUBA and snorkeling gear to find conchs. They are often brought to port live. They are cleaned by knocking a hole between the third and fourth whorl of the spire, inserting a knife to cut the animal free from the shell, and then pulling it out. The soft parts of the body are used for fish bait or discarded *213*. REGULATIONS Venezuela, Belize and Puerto Rico now have seasons closed to conch harvest. These shellfish have been so extensively harvested that their numbers have severely declined. In 1978, Florida declared a moratorium on commercial fishing. In 1985, Florida banned all harvest, killing, molestation, harming or mutilation of queen conch in Florida waters. Beginning in December 1986, the rule was extended so now harvest by all persons in Florida-registered vessels in federal waters contiguous to Florida waters is banned. As of 1989, it is unlawful to take or harvest any queen conch from the land or waters of Florida or to possess or transport any queen conch so taken or harvested *213*. FARMING POTENTIAL There has been great interest throughout the Caribbean in breeding and rearing conchs in aquaculture systems, especially since 1982. Several Management Practices - 1 (DRAFT) - Management Practices Species conch, queen Species Id M060165 Date 26 AUG 96 hatcheries have been established, but mass-rearing of conchs has had mixed success *213*. STOCKING In 1986, the Florida Department of Natural Resources field station in Marathon, Florida, initiated a stock assessment program and a research effort to determine the feasibility of queen conch stocking in Florida as a management tool. To date, small-scale efforts reported in the literature have indicated massive mortality of hatchery-reared juveniles released in the field. During 1988-1989, to test out-door facilities for grow-out and production of larger juveniles, hatchery-reared conchs were raised to a size of five inches. Much work is yet to be done to determine whether stock enhancement is, in fact, necessary and feasible. To protect native stock, permits will eventually be required before any hatchery animals can be released into the wild. *213*. Management Practices - 2
                                   (DRAFT) - References
                                   Species conch, queen
                                    Species Id M060165
                                      Date 26 AUG 96



     

References

186 * Turgeon, D.D., A.E. Bogan, E.V. Coan, W.K. Emerson, W.G. Lyons, W.L. Pratt, C.F.E. Roper, A. Scheltema, F.G. Thompson, J.D. Williams. 1988. Common and scientific names of aquatic invertebrates from the United States and Canada: mollusks. American Fisheries Society Special Publication 16. (ed.):277. 213 * Stewart, V.N. 1990. Sea-Stats - A summary of information and statistics on Florida's marine organisms and the marine environment. Queen Conch. (14) (ed.). FL Department of Natural Resources St. Petersburg, FL:10. References - 1