(DRAFT) - Taxonomy
                               Species oyster, Olympia
                                 Species Id M060200
                                   Date 26 AUG 96



TAXONOMY

NAME - oyster, Olympia OTHER COMMON NAMES - California oyster, native oyster, shoalwater oyster, Yaquina Bay oyster and rock oyster ELEMENT CODE - CATEGORY - Aquatic Molluscs PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - Bivalvia, ORDER AND SUBORDER - Pteroidea, FAMILY AND SUBFAMILY - Ostreidae, GENUS AND SUBGENUS - Ostrea, SPECIES AND SSP - lurida, SCIENTIFIC NAME - Ostrea lurida AUTHORITY - TAXONOMY REFERENCES - 238 COMMENTS ON TAXONOMY - Other common names include California oyster, native oyster, shoalwater oyster, Yaquina Bay oyster, rock oyster.*238* Taxonomy - 1
                                  (DRAFT) - Status
                               Species oyster, Olympia
                                 Species Id M060200
                                   Date 26 AUG 96



STATUS

Coded Status Sport Fish Commercial Commercial/consumption Subsistence REFERENCES FOR STATUS - 238 COMMENTS ON STATUS - The Olympia oyster once supported an Indian sustenance fishery and a commercial fishery, and was used to establish a maricultural industry on the west coast of the United States. Examination of Indian kitchen middens near San Francisco Bay revealed Olympia oyster shells in sufficient quantities to establish the animals as an important food item of coastal tribes.*238* Olympia oysters still form a reef and have repopulated much of Humboldt Bay, allowing a sport harvest.*238* Status - 1
                               (DRAFT) - Distribution
                               Species oyster, Olympia
                                 Species Id M060200
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - MARINE REFERENCES FOR HABITAT - 238 LAND USE - Bays and Estuaries REFERENCES FOR LAND USE - 238 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine RF2 L Marine, subtidal RF2 L Marine, intertidal RF2 L REFERENCES FOR NWI - 238 COMMENTS ON HABITAT ASSOCIATIONS - Olympia oysters survive in a broad range of habitats but are most abundant in estuaries, small rivers and streams.*238* Olympia oyster reefs are formed in the subtidal zone and are bordered by mud flats at high elevations and by eel grass beds at low elevations. They are found at depths of 0 to 71 m. Oysters may attach to the underside of rocks higher in the intertidal zone where the bottom is gravel or rock.*238* Habitat Associations - 1
                                (DRAFT) - Food Habits
                               Species oyster, Olympia
                                 Species Id M060200
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - FILTERER REFERENCES FOR TROPHIC LEVEL - 238 LIFESTAGE FOOD FOOD PART General Phytoplankton Not Applicable REFERENCES FOR GENERAL FOOD - 238 COMMENTS ON FOOD - Olympia oysters are filter feeders and rely on phytoplankton in the tidal waters as a food source. The ostia--gill openings through which water passes--are larger in the Olympia oyster than in the Japanese oyster. The large ostia result in the selection of larger food items and do not allow the Olympia oyster to consume nannoplankton.*238* Food Habits - 1
                         (DRAFT) - Environment Associations
                               Species oyster, Olympia
                                 Species Id M060200
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G G Relation to Substrate: Attached - normally sessile G Bottom Type [Aquatic]: Boulders G Bottom Type [Aquatic]: Specified in Comments G Water Depth Preference: Specified in Comments G Coastal Features: Reefs G Coastal Features: Rocky beaches REFERENCES FOR ENVIRONMENTAL ASSOC_ - 238 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Olympia oysters are sensitive to extreme high or low temperatures. Average water temperatures for southern Puget Sound are 6 to 9 degrees C in winter and 18 to 20 degrees C in summer. In Olympia oysters held at Milford, Connecticut, through the winter, mortalities were 100% at -1.0 to 5.0 degrees C and 3% at 12.0 to 13.0 degrees C.*238* Olympic oysters thrive at salinities above 25 ppt but tolerate occasional short exposure to lower salinities. Some oyster growers overcome predation from flatworms and drills by using oyster grounds which have flows of freshwater at low tide. The Olympia oysters can survive the salinity change but the flatworms and eggs of the drill cannot.*238* Environment Associations - 1
                               (DRAFT) - Life History
                               Species oyster, Olympia
                                 Species Id M060200
                                   Date 26 AUG 96



LIFE HISTORY

MORPHOLOGY: The lower (left) valve is shallowly concave and the upper (right) valve fits into the raised margin of the opposite valve. Shell shape is extremely variable. Shells on substrate will conform to the shape of the substrate. The shells of free-growing oysters are ovate to elongate. The exterior of the shell is without a periostracum, varies from white to purplish black, and may be striped with yellow or purplish brown. The interior of the valves is white to olive green and the scar of the adductor muscle is not much darker than the rest of the shell's interior.*238* REPRODUCTION: The life history of the Olympia oyster is similar to that of other oysters of the genus Ostrea. The oyster initially spawns as a male then alternates its functional gender between each spawning cycle.*238* Spawning begins at water temperatures of 13 to 16 degrees C. In the southern portion of the oysters' range, spawning occurs from spring to fall, peaking in spring and to a lesser extent in fall. In the central part of its range, spawning may be a prolonged period or may consist of one or two spawning periods in mid-summer. In the northern portion, there may be only one or two spawning periods in mid-summer.*238* Spawning begins when males release clusters of sperm into the mantle cavity. These sperm balls, which are expelled from the body by contractions of the shell, disintegrate in seawater and release the spermatozoa. Spermatozoa are brought into the mantle cavity of the female with water pumped by the gills and the eggs are fertilized. Presence of spermatozoa in seawater will stimulate synchronous spawning in Olympia oysters. The fertilized eggs develop into veliger larvae in the oyster's mantle chamber before they are discharged 10 to 12 days later. An average brood of larvae is 250,000 to 300,000. At discharge, the larvae are 185 to 187 micrometers long. The veliger stage lasts 11 to 16 days.*238* SPECIES ORIGIN WITHIN STATE: This species was introduced from Japan.*238* LIMITING FACTORS: Several introduced predators and parasites attack Olympia oysters. Two introduced invertebrate predators of Olympia oysters are the Japanese oyster drill Ocenebrajaponica and the flatworm Pseudostylochus ostreophagus. These two species were introduced with regualr sea shipments of Pacific oyster spat from Japan. The copepod Mytilicola orientalis ("red worm") lives in the anus of oysters, and oysters infected by it are usually in poor condition; this species was also introduced from Japan. In southern Puget Sound, the slipper shell Crepidula fornicata may compete with Olympia oysters for food and space and is a major fouling organism on the oyster's shell. This slipper shell was introduced from the East Coast with the shipment of American oysters. Other pests are the ghost shrimp and the blue mud shrimp. The burrowing activity of these shrimp stirs up sediment and also weakens the dykes used for culturing oysters.*238* Olympia oysters have several natural predators. Some sea ducks (scaups and scoters) feed on Olympia oysters, and the rock crab Cancer productus is an important predator over much of the oyster's range. In California, the bat ray Myliobatis californica is an oyster predator. In Yaquina Bay, Oregon, histological examination of Olympia oysters has revealed a proliferative cellular disorder characterized by large, apparently abnormal cells that Life History - 1 (DRAFT) - Life History Species oyster, Olympia Species Id M060200 Date 26 AUG 96 proliferate rapidly and cause death. the etiology of this disease is unknown. Tissue examination of oysters from Puget Sound, Washington, collected after cold-water mortality showed the presence of Hexamita and bacteria. Experimentation showed that Hexamita, a flagellated protozoan, could be transmitted to healthy oysters and act as a pathogen at low temperatures.*238* POPULATION ATTRIBUTES: The larvae develop an eyespot and foot and begin to crawl on hard substrate then metamorphose into spat. The preferred substrate is old oyster shells, but rocks, wood, metal, or any other hard material may be used as a setting surface. Olympia oyster spat is often set on the underside of horizontal surfaces. This is not a phototrophic response, but is attributed to the swimming position of the larvae in which the foot is held in a superior position. The oyster crawls with the foot on the surface and glue is secreted from the byssus gland, which attaches the shell to the substrate.*238* Artificial spat collectors that create a turbulent waterflow, disrupting the normal swimming position of larvae, collect spat on the upper and underside of horizontal surfaces. Artificial spat collectors, used to collect spat commercially, may be boards, pipes, or cardboard egg divdiders covered with a slurry of hydrated lime, portland cement, and fine sand. Olympia oysters differ from other species in that the substrate on which they settle need not be particularly clean and may be placed in the water in advance of spat settlement.*238* The Olypmia oyster grows slowly, reaching shell heights of 35 to 45 mm in three years in Washington State and has little or no growth thereafter. Shell height increased 2 mm per week at the Scripps Institute of Oceanography in southern California. Growth slows as the oysters mature but shell heights of 50 mm are sometimes reached in 30 weeks. The maximum reported size of Olympia oysters is 75 mm.*238* The maximum age of Olympia oysters has not been reported. In adults held in long-term bioassays, annual mortality was 34% in control groups.*238* LIFE HISTORY CODES - Foraging Strategy: Filtering Display Site: Water Spawning Site: Rocks Gestation/Incubation Period: 3-4 weeks Average Number of Offspring/Reproductive Effort: Grea Number of Broods/Litters (Reproductive Efforts) Per Ye Number of Broods/Litters (Reproductive Efforts) Per Ye REFERENCES FOR LIFE HISTORY- 238 Life History - 2
                           (DRAFT) - Management Practices
                               Species oyster, Olympia
                                 Species Id M060200
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Existing Put-and-take stocking Beneficial Placing artificial reefs under water Beneficial Constructing/maintaining bulkheads, seawalls, and dikes Adverse Other management practices [specified in comments] Adverse Other management practices [specified in comments] Adverse Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 238 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 238 REFERENCES FOR EXISTING MANAGEMENT PRACTICES - 238 COMMENTS ON MANAGEMENT PRACTICES - Early husbandry of the oysters was practiced by returning oyster shells to oyster reefs which provided substrate for new generations of oysters. In the early 1900's, the construction of dyked oyster beds, which held water at low tide and had gravel bottoms, marked a major development in oyster culture. Oyster shells were placed in locations favored for spat collection. Cultch containing Olympia oysters was spread in the dyked beds and left for 3.5 to 5 years until most of the oysters had reached a marketable size or 35-40 mm. These oysters were marketed as shucked meats rather than in the shell.*238* The decline in cultured Olympia oyster production was attributed to urbanization and domestic pollution and industrial pollution. The discharge of sulphite waste liquor from pulp mills has been especially destructive to oysters. Major growing grounds previously used for Olympia oysters are used to grow Pacific oysters thus further reducing production of Olympia oysters.*238* The assemblage of invertebrates in bays of the Pacific coast is currently cosmopolitan, owing to the introduction of exotic species. Possibly some displacement of native species by invading exotic species has occurred.*238* Silt from highway construction has caused high mortality by smothering oysters in shallow areas of southern Puget Sound. Olympia oyster populations have never recovered to their preexploitation levels.*238* Management Practices - 1
                                   (DRAFT) - References
                                 Species oyster, Olympia
                                    Species Id M060200
                                      Date 26 AUG 96



     

References

238 * Couch, David, T.H. Hassler. 1989. Species Profiles and Environmental Requirements of Coastal Fishes and Invertebrates (Pacific Northwest) --Olympia oyster. Species Profile Series 82(11.124) (ed.). U.S. Fish and Wildlife Service Washington, DC:8. References - 1