(DRAFT) - Taxonomy
                              Species ciguatera(poison)
                                 Species Id M090010
                                   Date 26 AUG 96



TAXONOMY

NAME - ciguatera(poison) OTHER COMMON NAMES - ciguatera and ciguatera poisoning ELEMENT CODE - CATEGORY - Other Aquatic Invertebrate Taxa PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - , FAMILY AND SUBFAMILY - , GENUS AND SUBGENUS - Gambierdiscus, SPECIES AND SSP - toxicus, SCIENTIFIC NAME - Gambierdiscus toxicus AUTHORITY - TAXONOMY REFERENCES - 214 COMMENTS ON TAXONOMY - Ciguatera (pronounced "si gwah teh' rah) is a human illness that results from the consumption of the toxic meat of many species of large, marine, tropical and subtropical food-fish *214*. The illness is caused by at least several toxic dinoflagellates, mainly Gambierdiscus toxicus.*214* Taxonomy - 1
                                  (DRAFT) - Status
                              Species ciguatera(poison)
                                 Species Id M090010
                                   Date 26 AUG 96



STATUS

Coded Status Unclassified REFERENCES FOR STATUS - 214 Status - 1
                               (DRAFT) - Distribution
                              Species ciguatera(poison)
                                 Species Id M090010
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC REFERENCES FOR HABITAT - 214 LAND USE - Water Atlantic Ocean Coastal Waters REFERENCES FOR LAND USE - 214 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine RF. Marine RF1 REFERENCES FOR NWI - 214 COMMENTS ON HABITAT ASSOCIATIONS - One of the first documented sources of ciguatera polyether toxins in the Pacific and Atlantic Oceans was Gambierdiscus toxicus, a single-celled, benthic dinoflagellate smaller than a hundredth of an inch in diameter. Recently, however, other toxic dinoflagellates, which are usually attached to algae in coral reef environments or shallow waters, have also been implicated *214*. Preferred habitats of the organisms are usually areas sheltered from wave action with little freshwater input. The cells are most frequently found in water down to 100 feet deep. They are usually found attached to larger algae and often associated with coral reefs or other hard surfaces, such as shipwrecks *214*. ANIMAL/PLANT SPECIES ASSOCIATIONS - algae fish REFERENCES FOR SPECIES ASSOCIATIONS - 214 COMMENTS ON SPECIES ASSOCIATIONS - Ciguatera toxins tend to become concentrated in larger animals near the top of the food chain. About 400 species of fish and a few invertebrates throughout the marine tropics have been implicated in ciguatera poisonings *214*. The toxic organisms are usually found attached to larger algae *214*. Habitat Associations - 1
                                (DRAFT) - Food Habits
                              Species ciguatera(poison)
                                 Species Id M090010
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - OTHER REFERENCES FOR TROPHIC LEVEL - 214 COMMENTS ON FOOD - All of the dinoflagellates suspect as possible toxin-bearing cells are photosynthetic. Photosynthetic organisms use the sun's energy, in the presence of chlorophyll and water, to produce carbohydrates (food) and release oxygen *214*. Food Habits - 1
                         (DRAFT) - Environment Associations
                              Species ciguatera(poison)
                                 Species Id M090010
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Turbidity: Specified in Comments G Substrate: Plants G Substrate: Specified in Comments G Relation to Substrate: Specified in Comments G Water Depth Preference: Specified in Comments G Coastal Features: Reefs G Coastal Features: Specified in Comments G Human Association: Specified in Comments G Wave intensity: specified in comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 214 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - Ciguatera is a human illness that results from the consumption of the toxic meat of many species of large, marine, tropical and subtropical food-fish. Gambierdiscus toxicus and other ciguateric organisms inhabit tropical and subtropical waters of both the Atlantic and Pacific Oceans. Their abundance varies with the season, water depth, seafloor (substrate) and environmental factors, such as temperature and salinity. Preferred habitats of the organisms are usually areas sheltered from wave action with little freshwater input. The cells are most frequently found in water down to 100 feet deep. They are usually found attached to larger algae and often associated with coral reefs or other hard surfaces, such as shipwrecks *214*. The organisms seem to flourish following some major disturbance to a reef. For example, ciguatera outbreaks have been associated with construction on or dredging of reefs. Natural disturbances, such as severe storms, may also disrupt reef communities and ciguateric organisms then increase. In either case, the "clean" exposed hard surfaces are first colonized by macroalgae, which become the substrate for dinoflagellates *214*. Environment Associations - 1
                               (DRAFT) - Life History
                              Species ciguatera(poison)
                                 Species Id M090010
                                   Date 26 AUG 96



LIFE HISTORY

DEFINITION Ciguatera (pronounced "si gwah teh' rah) is a human illness that results from the consumption of the toxic meat of many species of large, marine, tropical and subtropical food-fish *214*. Ciguatera toxin(s) are compounds known as polyether substances. The disease cannot be attributed to a single toxic compound. The toxins originate from single-celled, microscopic dinoflagellates, primarily Gambierdiscus toxicus and some other benthic species, which pass through the food web from prey to predator. Ciguatera toxins tend to become concentrated in larger animals near the top of the food chain. About 400 species of fish and a few invertebrates throughout the marine tropics have been implicated in ciguatera poisonings. Some groupers, snappers, tropical mackerels, barracuda, jacks and hogfish are included among the potential sources of ciguatera. In Florida, large barracuda, grouper and snapper are most often implicated *214*. CIGUATOXIC CELLS One of the first documented sources of ciguatera polyether toxins in the Pacific and Atlantic Oceans was Gambierdiscus toxicus, a single-celled, benthic dinoflagellate smaller than a hundredth of an inch in diameter. Recently, however, other toxic dinoflagellates, which are usually attached to algae in coral reef environment or shallow waters, have also been implicated. Among the organisms identified as possible toxin-bearing cells are Prorocentrum concavum and P. mexicanum. Several other dinoflagellate species are suspect and are also under investigation. All of them are photosynthetic dinoflagellates. However, more than a thousand other species are not toxin-bearing. Photosynthetic dinoflagellates are important producers of oxygen and organic carbon, hence vital components of the marine environment *214*. REPRODUCTION As a single-celled organism, Gambierdiscus toxicus reproduces asexually by cell division. In common with other dinoflagellates, however, it probably has a sexual reproductive phase *214*. PREDATION Dinoflagellates are food for many small marine animals that browse through tropical algal and coral reef communities. In turn, small animals are food to larger crustaceans and fish, and so on through the food web. The larger predators, near the top of this chain, ingest the toxin apparently unharmed. As they continue to feed on ciguateric prey, they continue to accumulate the toxins in their bodies. If these animals are consumed by humans, the toxins in the fish flesh and organs cause the illness known as ciguatera *214*. REFERENCES FOR LIFE HISTORY- 214 Life History - 1
                           (DRAFT) - Management Practices
                              Species ciguatera(poison)
                                 Species Id M090010
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Adverse Placing artificial reefs under water Adverse Dredging Adverse Other management practices [specified in comments] Beneficial Maintaining undisturbed/undeveloped areas Beneficial Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 214 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 214 COMMENTS ON MANAGEMENT PRACTICES - NOTE: Above codes are addressed to impact on human health. The organisms seem to flourish following some major disturbance to a reef. Ciguatera outbreaks have been associated with construction on or dredging of reefs. Natural disturbances, such as severe storms, may also disrupt reef communities and ciguateric organisms then increase. In either case, the "clean" exposed hard surfaces are first colonized by macroalgae, which become the substrate for dinoflagellates *214*. A recent consideration has developed in regard to the construction of artificial reefs in areas where the potential introduction of toxic dinoflagellates may initiate ciguatera. Placement of artificial habitats should be carefully considered *214*. PUBLIC HEALTH PROBLEMS As commercial fishing for tropical reef fishes increases, a rise in the widespread occurrence of ciguatera is expected. In 1980, 12 people in Maryland were afflicted, and in 1982, one case was recorded in Boston. Both incidents involved grouper shipped from Florida *214*. Each year, ciguatera poisonings in tropical and subtropical regions worldwide are estimated to be in the tens of thousands, with a 1% mortality rate. In endemic areas, such as the Virgin Islands, where people are familiar with the disease, episodes of illness are not always reported to health officials. As a result, case numbers reported by doctors and hospitals are low. In the United States and its territories, ciguatera is the most frequently reported food-borne disease associated with a naturally occurring chemical (toxin). In the three-years from 1974-1976, 129 cases were documented in a Miami, Florida, study. Ciguatera illness is not considered to be a "reportable" disease, however, nor is the illness always rcognized as ciguatera toxicity, so statistics on its occurrence are probably low *214*. Ciguatera toxicity is separate and distinct from illnesses in humans due to bacterial contamination from improperly handled seafood. The ciguatera toxins are temperature-stable, mostly oil-soluble compounds. Cooking, freezing or smoking fish will not destroy the toxins. Management Practices - 1 (DRAFT) - Management Practices Species ciguatera(poison) Species Id M090010 Date 26 AUG 96 Unfortunately, the toxins cannot be detected in the meat by appearance or taste. Symptoms of ciguatera poisoning usually appear in three to five hours after the toxic fish has been eaten. Most victims suffer nausea and vomiting, watery diarrhea, numbness and tingling about the mouth and extremities. More severe cases may suffer muscle pains, dizziness and sensations of temperature reversal, where hot objects seem cold and cold things seem hot. Normal recovery usually requires several days to several weeks. At this time, treatment is symptomatic, easing the distress of nausea, diarrhea and pain. A specific treatment does not exist for ciguatera illness. Mortalities are extremely rare *214*. CURRENT RESEARCH Scientists are seeking a simple chemical diagnostic test to detect ciguatoxin in seafood. Current procedures include complicated laboratory techniques and animal assays, and are not practical for routine surveillance. In Hawaii, a simple test kit for distribution to fishermen is under consideration *214*. Several hundred species of fish and invertebrates have been implicated in ciguatera cases, so continued research efforts are of vital concern *214*. Management Practices - 2
                                   (DRAFT) - References
                                Species ciguatera(poison)
                                    Species Id M090010
                                      Date 26 AUG 96



     

References

214 * Stewart, V.N. 1988. Sea-Stats - A summary of information and statistics on marine organisms common in Florida waters. Ciguatera. (3) (ed.). FL Dept. of Natural Resources St. Petersburg, FL:6. References - 1