(DRAFT) - Taxonomy
                                Species seal, bearded
                                 Species Id M120103
                                   Date 26 AUG 96



TAXONOMY

NAME - seal, bearded OTHER COMMON NAMES - bearded seal (English), square flipper (Norwegian sealers), morski zaits (Western Russian term meaning sea hare), laktak (Eastern Russian term), mukluk (Siberian Upik Eskimo) and oogruk (Inupiat Eskimo term) ELEMENT CODE - CATEGORY - Marine Mammals PHYLUM AND SUBPHYLUM - Chordata, CLASS AND SUBCLASS - Mammalia, ORDER AND SUBORDER - , FAMILY AND SUBFAMILY - , GENUS AND SUBGENUS - Erignathus, SPECIES AND SSP - barbatus, SCIENTIFIC NAME - Erignathus barbatus AUTHORITY - Erxleben TAXONOMY REFERENCES - 170 COMMENTS ON TAXONOMY - The scientific name, Erignathus barbatus, is descriptive of two very characteristic features of the bearded sea. The generic name Erignathus is Greek and refers to the deep jaw. The specific name, barbatus, is of Latin origin and refers to the relatively long and numerous moustachial vibrissae.*170* Two subspecies have been described: E. b. barbatus from the Laptev Sea, Barents Sea and North Atlantic Ocean into Hudson Bay; and E. b. nauticus from the remaining portions of the Arctic Ocean and the Bering and Okhotsk seas. The validity of the division into subspecies has been questioned, however.*125* Taxonomy - 1
                                  (DRAFT) - Status
                                Species seal, bearded
                                 Species Id M120103
                                   Date 26 AUG 96



STATUS

Coded Status Other, no FMP or PMP exists See Comments REFERENCES FOR STATUS - 125 COMMENTS ON STATUS - Currently, there are no research or management programs for bearded seals in the United States. Basic harvest monitoring is needed as well as research addressing the ecological interactions of bearded seals with competing and prey species. At a minimum, harvest monitoring should include records of the numbers taken by susistence users and Soviet sealing ships. The age and sex composition of the takes also should be recorded and analyzed for indications of temporal changes in the population structure. The age and sex biases of harvests in different regions and seasons will have to be investigated.*125* Status - 1
                               (DRAFT) - Distribution
                                Species seal, bearded
                                 Species Id M120103
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine OW0 REFERENCES FOR NWI - 125 COMMENTS ON HABITAT ASSOCIATIONS - The range of bearded seals broadly overlaps that of other ice-inhabiting pinniped, partly due to an ability to make and maintain breathing holes, and to their less discriminating selection of ice type. The proportion of bearded seals wintering in the shorefast ice in unknown but probably small. Ice is important to bearded seals as a substrate for resting, whelping, and molting. Bearded seals whelp over a wider area and longer time span than do ther phocids in the Bering Sea, possibly because bearded seals are able to use less predictable ice.*125* Habitat Associations - 1
                                (DRAFT) - Food Habits
                                Species seal, bearded
                                 Species Id M120103
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 125 LIFESTAGE FOOD FOOD PART General Crustaceans Not Specified General Molluscs Not Specified General Fish Not Specified General Bivalve Molluscs Not Specified Important Crustaceans Not Specified Important Molluscs Not Specified REFERENCES FOR GENERAL FOOD - 125 REFERENCES FOR IMPORTANT FOOD - 125 COMMENTS ON FOOD - Bearded sea;s are primarily benthic feeders, showing variations in diet with age, location, season, and perhaps with long-term changes in the composition of benthic communities. A wide variety of invertebrate and fish prey has been reported, but decapod crustaceans and molluscs make up most of the diet throughout the bearded seal's range. Major prey in the Bering, Chukchi, and Beaufort seas include crabs, clams, shrimps, and Arctic cod. No significant differences have been observed in the feeding habits of male and female bearded seals, but age-related changes have been observed. Shrimp are a larger component of the diet of first and second-year seals than of older seals, who eat more clams.*125* COMMENTS ON ADULT FOOD - Bearded seals are primarily benthic feeders utilizing mostly epibenthos (organisms occurring on, rather than in, the bottom), although organisms of the infauna and some demersal fishes are also consumed. Although the total array of food items consumed by bearded seals is large, relatively few types of organisms comprise the bulk of their diet. These include brachyuran crabs, shrimp, clams, sculpins, and sometimes arctic or saffron cod. Geographical variation in diet is largely a reflection of local faunal differences. Major prey types remain the same among areas, however, the species eatern may change.*170* COMMENTS ON JUVENILE FOOD - The nursing period in bearded seals is comparatively hsort, lasting 12-18 days. During this time the weight of the pup increases approximately 150%, mostly in the form of an accumulated blubber layer. Weaning occurs when the female abruptly deserts the pup. Some independent feeding occurs during the latter part of the nursing period; newly weaned pups are active feeders. The apparently eat more isopods, hippolytid shrimps and saffron cod than adults do, and fewer brachyuran crabs, pandalid and crangonid shrimps, sculpins, and flatfishes.*170* In the Bering Sea, hippolytid shrimp are eaten more frequently by pups, Food Habits - 1 (DRAFT) - Food Habits Species seal, bearded Species Id M120103 Date 26 AUG 96 while pandalid shrimp are more common in stomachs of older seals. Isopods and saffron cod are more important to younger seal; brachyuran crabs, sculpins, and flatfishes are more important to older seals.*125* Food Habits - 2
                         (DRAFT) - Environment Associations
                                Species seal, bearded
                                 Species Id M120103
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

Environment Associations - 1
                               (DRAFT) - Life History
                                Species seal, bearded
                                 Species Id M120103
                                   Date 26 AUG 96



LIFE HISTORY

Morphology: Bearded seals are covered with relatively short, straight hair. They have none of the distinct and diagnostic color patterns such as spots, rings or bands found on other species of seals. Most adults are basically light to dark gray, being slightly darker down the middle of the back. Coloration is sometimes tawny-brown to dark brown. Term fetuses and newborn pups have dark (usually brown), dense, slightly wavy hair with light coloration on the facial region and one to four broad, transverse light bands on the crown and back. At weaning, the pelage resembles that of older seals.*170* Maximum life expectancy of bearded seals is about 30 years. The oldest animal recorded from the Bering and Chukchi seas was 26 years of age. Based on data obtained from animals killed by coastal Eskimo hunters, about 75% of the population was ten years of age or younger, and over 95% was 20 years or younger. At some locations, samples appeared to be significantly biased toward younger aged seals, lending credence to the hypothesis that younger animals occur in larger numbers close to shore. Most male bearded seals become sexually mature, as indicated by testes and baculum size and by the onset of spermatogenic activity, at ages six to seven. Some females first ovulate at the age of three years but most do not become pregnant until the age of six years. Observed ovulation rates are: 17% at age four; 50% at age five; 80% at age six; and 100% at age eight. Life expectancy: The sex ratio of bearded seals appears to change with age. Studies conducted within the Bering-Chukchi area indicate that at birth males predominate, although the ratio is very close to one-to-one. In collections of animals older than pups, females composed 53-55%. This suggests a higher mortality rate in males although other factors such as sex-related differences in behavior, distribution, or sinking rate may influence the sex ratio of harvested animals. Maximum life expectancy of bearded seals is about 30 years. The oldest animal recorded from the Bering and Chukchi seas was 26 years of age. Based on data obtained from animals killed by coastal Eskimo hunters, about 75% of the population was ten years of age. Based on data obtained from animals killed by coastal Eskimo hunters, about 75% of the population was ten years of age or younger. At some locations, samples appeared to be significantly biased toward younger aged seals, lending credence to the hypothesis that younger animals occur in larger numbers close to shore. Predators: The major predators of bearded seals in the Bering-Chukchi region are polar bears and humans. The magnitude of polar bear predation is unknown; however, in one study undertaken in Alaskan waters, five of 71 pinnipeds (7%) killed by bears were bearded seals.*170* REFERENCES FOR LIFE HISTORY- 170 Life History - 1
                           (DRAFT) - Management Practices
                                Species seal, bearded
                                 Species Id M120103
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Existing Other management practices [specified in comments] REFERENCES FOR EXISTING MANAGEMENT PRACTICES - 170 COMMENTS ON MANAGEMENT PRACTICES - Bearded seals may be competing increasingly with walruses for clams and with commercial fisheries for crabs. At the same time they may be subjected to increasing rates of harvest. Currently, however, there are no research or management programs for bearded seals in the United States. Basic harvest monitoring is needed as well as research addressing the ecological interactions of bearded seals with competing and prey species. Currently, there are no research or management programs for bearded seals in the United States. Basic harvest monitoring is needed as well as research addressing the ecological interactions of bearded seals with competing and prey species. At a minimum, harvest monitoring should include records of the numbers taken by susistence users and Soviet sealing ships. The age and sex composition of the takes also should be recorded and analyzed for indications of temporal changes in the population structure. The age and sex biases of harvests in different regions and seasons will have to be investigated.*125* The efficacy of strip transect surveys for censusing bearded seals needs to be tested in surveys stratified primarily for bearded seal habitat. Survey methods more suitable to the low densitieds of bearded seals should be sought. Behavioral data are needed to develop correction factors for aerial surveys. The diurnal haulout cycle of bearded seals during the spring survey period should be investigated using radiotelemetry. Seasonal changes in haulout timing by different age and sex classes should be determined. Data on haulout rhythms should be used to correct for the proportion of seals submerged and uncounted during surveys. The ratios of seals on the ice to seals in the water at peak haulout hours and other times should be determined and applied as corrections to surveys flown over the course of several hours. The variance in the proportion of seals on the ice at different times of day should also be determined by a study of haulout behavior. It may be that the best time of the day to survey will be when the varience is lowest, whether or not that corresponds with the time of peak haulout.*125* Management Practices - 1
                                   (DRAFT) - References
                                  Species seal, bearded
                                    Species Id M120103
                                      Date 26 AUG 96



     

References

170 * J. Burns, K. Frost and L. Lowry (eds.)1985. (7). Alaska Department of Fish and Game: 96. 125* Lentfer, J.W. (ed.). 1988. Selected Marine Mammals of Alaska, Species Accounts with Research and Management Recommendations. Marine Mammal Commission Washington, D.C pp 275. References - 1