(DRAFT) - Taxonomy
                                Species seal, ribbon
                                 Species Id M120104
                                   Date 26 AUG 96



TAXONOMY

NAME - seal, ribbon OTHER COMMON NAMES - ELEMENT CODE - CATEGORY - Marine Mammals PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - , ORDER AND SUBORDER - , FAMILY AND SUBFAMILY - , GENUS AND SUBGENUS - Phoca, SPECIES AND SSP - fasciata, SCIENTIFIC NAME - Phoca fasciata AUTHORITY - TAXONOMY REFERENCES - 125 and 170 COMMENTS ON TAXONOMY - Although once considered a member of a separate genus, Histriophoca, recent studies have shown that the ribbon seal is a member of the genus Phoca. This indicates the close relationship between ribbon seals and other members of the genus Phoca including ringed, spotted, harbor, and harp seals.*170* Taxonomy - 1
                                  (DRAFT) - Status
                                Species seal, ribbon
                                 Species Id M120104
                                   Date 26 AUG 96



STATUS

Coded Status Marine Mammal Furbearer Subsistence See Comments REFERENCES FOR STATUS - 170 and 125 COMMENTS ON STATUS - Only Native subsistence hunters harvest ribbon seals in Alaskan waters.*125* Present provisions of the MMPA (Marine Mammal Protection Act) preclude actual management of the ribbon seal population and regulation of its users. *170* No major problems are anticipated with regard to future State (Alaska) management of ribbon seals. In the absence of large commercial or subsistence harvest, and given the secure status of the population, the amount of biological data gathered would be small. Emphasis would continue to be placed on monitoring harvest levels.*170* Status - 1
                               (DRAFT) - Distribution
                                Species seal, ribbon
                                 Species Id M120104
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - AQUATIC REFERENCES FOR HABITAT - 125 and 170 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine COMMENTS ON HABITAT ASSOCIATIONS - Although they spend many months in ice-free waters, sea ice is vital to the life cycle of ribbon seals. For pupping, ribbon seals select areas of moderately thick large floes commonly 10-60 km north of the ice fringe and open sea.*170* Ribbon seals seem to select an ice type that seldom occurs near shore, and tend to be 50-250 km offshore during the pupping and nursing periods and move to within 20-100 km of shore during the molt. Ribbon seals have been reported in highest densities in areas of 60-80% ice coverage. In April they have been found throughout the ice front, but most abundant south of the continental shelf over deep water.*125* Habitat Associations - 1
                                (DRAFT) - Food Habits
                                Species seal, ribbon
                                 Species Id M120104
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 170 LIFESTAGE FOOD FOOD PART General Squid, Octopus Not Specified General Crustaceans Not Specified General Fish Not Specified General See Comments; Food Not Applicable Adult Squid, Octopus Not Specified Adult Fish Not Specified Adult See Comments; Food Not Applicable Juvenile Crustaceans Not Specified Juvenile See Comments; Food Not Applicable Important Crustaceans Not Specified Important Squid, Octopus Not Specified REFERENCES FOR GENERAL FOOD - 170 REFERENCES FOR IMPORTANT FOOD - 170 REFERENCES FOR ADULT FOOD - 170 REFERENCES FOR JUVENILE FOOD - 170 COMMENTS ON FOOD - The foods of ribbon seals are known only for the months of February to June. Most samples are from April to June, a time of reduced feeding, but the types of foods in ribbon seal stomachs during spring undoubtedly shed some light on foods eaten at other times of year. In general, ribbon seals eat a variety of fishes, cephalopods (octopus and squid), and crustaceans (primarily shrimp and mysids).*170* COMMENTS ON ADULT FOOD - Based on food remains in stomachs of animals taken in spring, fishes and cephalopods comprise the bulk of the adult diet.*170* COMMENTS ON JUVENILE FOOD - Soviet researchers have suggested that crustaceans are the main food of young seals.*170* Food Habits - 1
                         (DRAFT) - Environment Associations
                                Species seal, ribbon
                                 Species Id M120104
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Substrate: Specified in Comments G Relation to Substrate: Specified in Comments G Aquatic Habitat Zonation: Open water [pelagic zone] G Aquatic Habitat Zonation: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 125 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - In Alaskan waters ribbon seals are found in the open sea, on the pack ice, and only rarely on shorefast ice. Clean ice that is not discolored by sediment or algae seems to be preferred. Except for occasional moribund individuals, they do not haul out on land. It has been noted that ribbon seals seem to select an ice type that seldom occurs near shore.*125* Environment Associations - 1
                               (DRAFT) - Life History
                                Species seal, ribbon
                                 Species Id M120104
                                   Date 26 AUG 96



LIFE HISTORY

Physical description: Adult males are easily recognized by the characteristic banding of their pelage, from which the species' name is taken. Four nearly white bands contrast with a brown to black background. One band encircles the neck and base of the head, another encircles the posterior portion of the trunk, and each forflipper in encircled by an oval extending generally from the lower neck to the midsection of the trunk. These "ribbons" are most distinct in adult males; most females show much less contrast between the markings and background coloration. The pattern is indistinct in young seals and becomes progressively clearer with successive molts up to the age of marturity.*125* In addition to their distinctive coloration, ribbon seals can be recognized by their relatively short snout, long neck, slender trunk, and seemingly long foreflippers. The ribbon seal skull is characterized by a short rostrum, broad cranium, and small, widely spaced teeth. Notable in soft tissue anatomy is an air sac extending from the posterior end of the trachea over the ribs on the right side. The inflatable sac is well developed only in adult males, and its function is unknown. It has been speculated that the air sac functions as a buoyancy device, for storage of air during diving, and in sound production.*125* Mature ribbon seals average 1.50-1.75 m in standard length and weigh up to 148 kg. Weight declines substantially in spring when the blubber thichness decreases by 50-60%.*125* Reproduction: Mating takes place from late April to early May in the Bering and Okhotsk seas. At that time of year the seals inhabit the southern ice front, where they apparently prefer to haul out on floes about 20 m in diameter.*125* Implantation of the blastocyst occurs approximately 2.5 months after fertilization and gestation lasts 8.5 months, so parturition occurs 11 months after mating. A single pup is born on the ice in April or May; the majority of births take place between April 5 and 15.*125* Population size: The most recent estimate of the worldwide population of ribbon seals is 240,000, from surveys conducted in the mid-1970s. Ribbon seals are difficult to enumerate because they occur in a remote and harsh environment and spend much time out of sight, underwater. Estimates of their numbers are based on aerial surveys conducted in spring when the breeding seals haul out on the ice.*125* Population trends: A decline in the numbers of ribbon seals in the Bering Sea was believed to be the result of overharvest by Soviet commercial sealers in the early 1960s. The catch per unit of effort dropped by more than 50% between the first commercial harvest in 1961 and 1963. The Bering Sea population was believed to have decreased from 80,000-90,000 in 1964 to 60,000 in 1969.*125* Predation: Potential predators of ribbon seals include man, polar bears, walruses, killer whales, eagles, and gulls.*125* Life History - 1 (DRAFT) - Life History Species seal, ribbon Species Id M120104 Date 26 AUG 96 Parasites: Ribbon seals harbor low numbers of anopluran lice. It has been suggested that the scarcity of the ectoparasites is related to the seal's low level of gregariousness.*125* More than a dozen species of helminths occur in ribbon seals. Helminth damage to the intestines, stomach, lungs, and possibly liver of ribbon seals has been reported, but the contribution of helminth infestations to ribbon seal mortality is not known.*125* Other causes of morbidity and mortality: Dermatitis is fairly common in ribbon seals. Molting seals especially may be vulnerable to microbial infections of the skin.*125* Ribbon seals are occasionally found traveling over ice and land after losing access to the water because of sudden freeze-ups or ice movements. Undoubtedly, some ribbon seals die as a result.*125* Ribbon seals are incidentally taken in groundfish trawls and salmon gill nets. The numbers of seals killed in this manner are unknown, but the incidence may be increasing, as is thought to be the case for other pinnipeds.*125* Organochlorine compounds and heavy metals have been found in high concentrations in some phocids, but the occurrence of pollutants in ribbon seals has not been investigated. The tendency of ribbon seals to remain away from coastal waters may decrease the risks of encountering some pollutants.*125* Offshore oil development in the Bering and Chukchi seas presents several potential causes of morbidity or mortality. Well blowouts or shipping accidents could bring ribbon seals into direct contact with crude oil. The effects on ribbon seals have not been studied. There is some evidence that seals may be most adversely affected by contact with oil during periods of naturally occurring stress such as the molt, reproduction, scarcity of food, or illness.*125* LIFE HISTORY CODES - Breeding Spawning Season: April Breeding/Spawning Season: May Gestation/Incubation Period: Greater than 8 months Average Number of Offspring/Reproductive Effort: 1 Number of Broods/Litters (Reproductive Efforts) Per Ye REFERENCES FOR LIFE HISTORY- 125 and 170 Life History - 2
                           (DRAFT) - Management Practices
                                Species seal, ribbon
                                 Species Id M120104
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Regulating harvest of species being described Beneficial Restricting/regulating human use of habitats Beneficial Other management practices [specified in comments] Adverse Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 125 REFERENCES FOR ADVERSE MANAGEMENT PRACTICES - 125 COMMENTS ON MANAGEMENT PRACTICES - Management of ribbon seals should be directed toward monitoring and regulating harvests and reducing potential threats from commercial fishing and offshore industrial development activities. The population and ecological data necessary for developing effective management plans currently are lacking. Harvest monitoring should be a routine part of management activities and should include analysis of the age and sex composition of the kill and the reproductive history of harvested seals.*125* Ribbon seals are incidentally taken in groundfish trawls and salmon gill nets. The numbers of seals killed in this manner are unknown, but the incidence may be increasing, as is thought to be the case for other pinnipeds.*125* Organochlorine compounds and heavy metals have been found in high concentrations in some phocids, but the occurrence of pollutants in ribbon seals has not been investigated. The tendency of ribbon seals to remain away from coastal waters may decrease the risks of encountering some pollutants.*125* Offshore oil development in the Bering and Chukchi seas presents several potential causes of morbidity or mortality. Well blowouts or shipping accidents could bring ribbon seals into direct contact with crude oil. The effects on ribbon seals have not been studied. There is some evidence that seals may be most adversely affected by contact with oil during periods of naturally occurring stress such as the molt, reproduction, scarcity of food, or illness.*125* Management Practices - 1
                                   (DRAFT) - References
                                   Species seal, ribbon
                                    Species Id M120104
                                      Date 26 AUG 96



     

References

125* Lentfer, J.W. (ed.). 1988. Selected Marine Mammals of Alaska, Species Accounts with Research and Management Recommendations. Marine Mammal Commission Washington, D.C pp 275. 170* Burns, John J., K.J. Frost, L.F. Lowry (ed.)Marine Mammal Species Accounts. 1985. (7). Alaska Department of Fish and Game:96. References - 1