(DRAFT) - Taxonomy
                                Species whale, beluga
                                 Species Id M120107
                                   Date 26 AUG 96



TAXONOMY

NAME - whale, beluga OTHER COMMON NAMES - beluga whale, belukha and white whale ELEMENT CODE - CATEGORY - Marine Mammals PHYLUM AND SUBPHYLUM - Vertebrata, CLASS AND SUBCLASS - Mammalia, ORDER AND SUBORDER - Cetacea, Odontoceti FAMILY AND SUBFAMILY - Monodontidae, GENUS AND SUBGENUS - Delphinapterus, SPECIES AND SSP - leucas, SCIENTIFIC NAME - Delphinapterus leucas AUTHORITY - Linnaeus TAXONOMY REFERENCES - 125, 166 and 178 COMMENTS ON TAXONOMY - The species is sometimes called "belukha" to differentiate it from the beluga sturgeon (the fish from which caviar is obtained) *166*. It is also sometimes called the white whale *125*. Taxonomy - 1
                                  (DRAFT) - Status
                                Species whale, beluga
                                 Species Id M120107
                                   Date 26 AUG 96



STATUS

Coded Status Marine Mammal See Comments REFERENCES FOR STATUS - 166 and 125 COMMENTS ON STATUS - Belugas have been hunted for centuries by native whalers for food, oil, and "porpoise leather". They are still being hunted in limited numbers in the Soviet Union, and by the Eskimo (Inuit) natives of Arctic Canada. Due to the high level of mercury in belugas, they are no longer considered edible, and hunting belugas for food has been banned. Some belugas have been captured for display by oceanariums. The current population is estimated to be 62,000-86,000 *166*. The average annual kill in Alaska and western Canada waters is about 570 (365 in Alaska and 205 in Canada). These estimates are from harvest figures and struck-and-lost estimates. Because data are so meager and information on the survival rate of unretrieved whales is almost completely lacking, estimates of the total annual kill are very imprecise *125*. Status - 1
                               (DRAFT) - Distribution
                                Species whale, beluga
                                 Species Id M120107
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine OW0 V 1 Estuarine AB. H 6 Estuarine AB. V 6 Riverine UB. H 6 Marine OW0 V 1 Estuarine AB. H 6 Estuarine AB. V 6 Riverine UB. H 6 Marine OW0 V 1 Marine, intertidal BB. V 1 REFERENCES FOR NWI - 125 Habitat Associations - 1
                                (DRAFT) - Food Habits
                                Species whale, beluga
                                 Species Id M120107
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE REFERENCES FOR TROPHIC LEVEL - 166 and 125 LIFESTAGE FOOD FOOD PART General Squid, Octopus See Comments General Perciformes See Comments General Clupeiformes See Comments General Salmoniformes See Comments General Lampridiformes See Comments General Osteichthyes See Comments General Pleuronectiformes See Comments General Gadiformes See Comments General Beryciformes See Comments General Crustaceans See Comments General Molluscs See Comments General Bivalve Molluscs See Comments Juvenile Crustaceans See Comments Juvenile Gadiformes See Comments Juvenile Salmoniformes See Comments Juvenile Perciformes See Comments Important Salmoniformes See Comments Important Crustaceans See Comments Important Clupeiformes See Comments Important Gadiformes See Comments REFERENCES FOR IMPORTANT FOOD - 125 REFERENCES FOR ADULT FOOD - 125 REFERENCES FOR JUVENILE FOOD - 125 COMMENTS ON FOOD - More than 100 kinds of organisms have been identified in the diet of belugas. General prey in stomachs collected along the northwest coast of Alaska include benthic invertebrates, squid, octopus, semidemersal fish (Arctic cod, saffron cod, herring, whitefish, smelt, char), and demersal fish (sculpins, suckers, eelpout). Salmon, eulachon, saffron cod, tomcod, herring, and smelt have been identified as prey items for belugas summering south of Bering Strait. Food items are swallowed whole. Octopus may be a significant spring food. Shrimp are also eaten. Of fish species consumed in spring, Arctic cod is taken in greatest numbers. Even though cod is the most commonly consumed fish, the whales feed more extensively in spring on invertebrates. Fish are the dominant food item in coastal areas of Alaska in summer. In general, belugas seem to feed on whatever fish species are most abundant and easy to catch. Thus, they prey on herring, rainbow smelt, capelin, salmon, char, eulachon, whitefish, saffron cod, and Arctic cod as these become seasonally abundant. Sculpin, flounder, sole, blenny, burbot, lamprey, Food Habits - 1 (DRAFT) - Food Habits Species whale, beluga Species Id M120107 Date 26 AUG 96 shrimp, mussels, octopus, and squid also contribute to the summer diet. In the northern Bering and southern Chukchi seas saffron cod is the most commonly consumed species. Differential feeding occurs by age and sex class. In the western Bering Sea in summer, yearlings take primarily cranonid shrimp and navaga cod; 2-, 3-, and 4-year-olds take primarily navaga cod and chum salmon; and older animals take primarily chum salmon. Other authors have noted that young animals take small prey. Yearlings supplement a diet of milk with capelin, sand lance, shrimp, and small bottom-dwelling crustaceans. Although younger and older whales of both sexes prey on the same species, older belugas eat significantly larger saffron cod than do young whales, and males eat significantly more sculpin (larger fish than saffron cod) than do females. In another study, adult males fed primarily on large fish, and females preferred food items such as sand lance, octopus, and nereid worms *125*. COMMENTS ON ADULT FOOD - Although younger and older whales prey on the same species, older belugas eat significantly larger saffron cod than do young whales *125*. COMMENTS ON JUVENILE FOOD - In the western Bering Sea in summer, yearlings take primarily crangonid shrimp and navaga cod; 2-, 3-, and 4-year-olds take primarily navaga cod and chum salmon; and older animals take primarily chum salmon. Other authors have also noted that young animals take small prey. Yearlings supplement a diet of milk with capelin, sand lance, shrimp, and small bottom-dwelling crustaceans *125*. Food Habits - 2
                         (DRAFT) - Environment Associations
                                Species whale, beluga
                                 Species Id M120107
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Water Temperature: Below 15 degrees C J Water Temperature: Specified in Comments G G Water Depth Preference: Specified in Comments G Aquatic Habitat Zonation: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 125 REFERENCES FOR LIMITING ENVIRONMENTAL ASSOC_ - 125 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 125 REFERENCES FOR RESTING ADULT ENVIRONMENTAL ASSOC_ - 125 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 125 REFERENCES FOR RESTING JUVENILE ENVIRONMENTAL ASSOC_ - 125 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - TEMPERATURE: Beluga whale distribution is generally limited to water temperatures below 15 deg C. ICE: In winter most belugas inhabit ice-permeated offshore waters. They must have access to air and therefore inhabit areas where geographic, oceanographic, or meteorologic factors cause ice motion and the formation of openings. Ice coverage affects seasonal distribution. DEPTH: During open-water months, nearly all major concentrations of belugas occur in shallow bays or in estuaries of large rivers north of 40 deg N. SALINITY: Soviet scientists have noted that belugas are most abundant in coastal shallows in years with heavy continental runoff. This association of beluga abundance with freshwater runoff is likely related to fish abundance *125*. COMMENTS ON RESTING ADULT ENVIRONMENTAL ASSOC_ - TEMPERATURE: Beluga distribution is generally limited to water temperatures below 15 deg C. However, all ages and both sexes inhabit warm estuarine waters during summer. ICE: In winter most belugas inhabit ice-permeated offshore waters. Complete ice coverage excludes use by belugas, while partial coverage Environment Associations - 1 (DRAFT) - Environment Associations Species whale, beluga Species Id M120107 Date 26 AUG 96 encourages use. Ice coverage affects seasonal distribution. DEPTH: During open-water months, nearly all major concentrations of belugas inhabit shallow bays or estuaries of large rivers north of 40 deg N. SALINITY: Soviet scientists have noted that belugas are most abundant in coastal shallows in years with heavy continental runoff. This association of beluga abundance with freshwater runoff is likely related to fish abundance *125*. COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - ICE: Prey and ice distribution often act together to influence beluga distribution. Movement of prey into coastal areas is synchronized with ice departure in spring, and association of prey with the ice edge during other seasons attracts belugas. In most coastal areas south of Bering Strait, the arrival of belugas coincides with the breakup and movement of ice offshore and the arrival of large runs of fish. DEPTH: Belugas seem to be more successful in obtaining prey in the rivers where prey are concentrated than in the bays where prey are more dispersed. Belugas in Bristol Bay feed at the mouth of the Snake River, where salmon runs are smaller than in other rivers in Bristol Bay. However, the mouth of the Snake River is shallower, and hence may concentrate the prey. TEMPERATURE: There is evidence that concentrations of belugas in warm-water estuaries in Kotzebue Sound, Kasegaluk Lagoon, areas north of Bering Strait, and eastern Canada are more closely associated with temperature than prey abundance. SALINITY: Association on beluga abundance with freshwater runoff is likely related to fish abundance *125*. COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - TEMPERATURE: Researchers have assumed that most calving takes place in coastal estuary areas. However, some calving, at least, takes place in colder offshore areas *125*. COMMENTS ON RESTING JUVENILE ENVIRONMENTAL ASSOC_ - TEMPERATURE: Noenates have thinner blubber and a higher surface-to-volume ratio than other age classes; warmer waters may favor their survival and growth *125*. Environment Associations - 2
                               (DRAFT) - Life History
                                Species whale, beluga
                                 Species Id M120107
                                   Date 26 AUG 96



LIFE HISTORY

PHYSICAL DESCRIPTION: The beluga's body is thick, muscular, and tapered at both ends, with a small head and narrow caudal peduncle (tail stock). Its head is rounded with a short beak and a prominent, protruding forehead called the "melon." The neck of the beluga is narrower than the necks of other whales, and it also differs from most whales--whose neck vertebrae are fused--in that its 7 neck vertebrae are free, which allows it to nod and turn its head. Adult belugas are almost completely white. Calves are gray to pinkish brown. At 2 years of age they become a dark gray; at 3-4 years they turn a lighter gray, and at full maturity they become white, though they retain some dark color on the edge of the flippers and flukes. Because it has no dorsal fin, the beluga can swim freely under floating ice. instead of a fin it has a narrow ridge which runs down the rear half of its back. the ridge is sometimes notched and may be a dark color. The flippers are short, broad, and may be curled up at the tip in adults. The fluke is broad, deeply notched in the center, and ornately curved. Adult males measure 5 meters (14-16 ft), with a maximum length of 5.3 m (20 ft), and weigh about 3,300 lbs. Adult females measure 3.9-4.2 m (13-14 ft), with a maximum length of 5.2 m (17 ft), and weigh about 3,000 lbs *166*. REPRODUCTION: Calving areas: Calving generally occurs throughout the beluga's circumpolar distribution between March and September, with a peak in June and July. In northwest Alaska the first post-parturient female was taken on 29 April and the last full-term fetus was recovered on 18 July. Calving peaks from mid-June to late July. Influx of animals to nearshore areas in mid-June could bias samples so that the peak in calving occurs earlier than it seems. Calving occurs in all coastal Alaska waters where belugas aggregate in summer. Neonates and afterbirths are reported from Bristol Bay, Norton Sound, Kotzebue Sound, Kasebaluk Lagoon and adjacent marine waters, and the eastern Beaufort Sea. Calving areas in Cook Inlet are not known. Calves were noted in Cook Inlet in mid-July but not in June, and it was hypothesized that calving begins between mid-June and mid-July, perhaps at the large river estuaries in the upper inlet. Researchers have assumed that most calving takes place in coastal estuary areas. However, some calving, at least, takes place in colder offshore waters. Calving sometimes occurs prior to or during the spring migration. Neonate calves are seen in the spring leads during April, May, and sometimes in March, and harvests in the leads along the northwest Alaska coast include female belugas in late pregnancy or recently in post-parturient. Here parturition must occur in very cold water. The average increase in standard length during the first year of life was 46.3 cm in a sample of 6 neonates and 4 1-year-olds from the northwest coast. Data are inadequate to show the increase in length during the first few months of life. The proportion of calves in various aggregations of belugas has been determined from aerial surveys and photogrammetry. Such methods may be reasonably accurate during June and July when neonates are small, but are much less accurate by late August when some calves are as long as yearlings. Sex ratio: Although size differences between male and female adult belugas are significant, other outward sexual dimorphisms do not commonly Life History - 1 (DRAFT) - Life History Species whale, beluga Species Id M120107 Date 26 AUG 96 distinguish males and females in the wild. Consequently, sex ratios are determined from specimens. From 533 carcasses a sex ratio of 1:1 was reported. There were, however, large deviations from unity in subsamples. Subsamples throughout the beluga's range show large differences in sex ratio, age composition, and reproductive status. Some sex and age segregation is apparent. Large deviations in pod composition pose problems in obtaining unbiased samples; sex ratios in harvest samples are also biased by hunter selectivity. Sex ratios of 1:1 have also been reported for other populations. Age at sexual maturity: The mean age at sexual maturity is slightly younger for females than for males. (Calculations of age are based on the assumption of two growth layers per year in teeth. Age at sexual maturity in females here means the age of initiation of first pregnancy.) The age of sexual maturity for females is 4-7 years, with first births at 5-8 years of age. Of 22 primiparous females from the Alaska coast, 54% conceived at age 4, 41% at age 5, and 5% at age 6. In a sample of 52 females, all animals up to age 4 were sexually immature (N=28); 33% of the 5-year-olds (N=9) and 94% of the 6-year-olds (N=16) were sexually mature. However, most of the samples were obtained at the approximate season of their birth dates; thus, 4-year-old females were not pregnant when taken during June, though some would have become so prior to their fifth birthday. Increases in testis volume and spermatogenic activity indicate sexual maturity in males. Males reach sexual maturity at 7-9 years. Pregnancy rates: A 3-year reproductive cycle is most typical in belugas. Of sexually mature females sampled from harvests along the northwest coast of Alaska, 35% were not pregnant, 35% were newly pregnant, and 30% carried full-term fetuses or had given birth. However, high pregnancy rates in 6- to 22-year-old females suggest that some conceive more frequently than once in 3 years. It's been estimated that in eastern Canada 25% of mature females have a reproductive cycle of 2 years and 75% have a cycle of 3 years. Life span and reproductive life: Belugas may live more than 30 years. Because older animals lose tooth dentine layers, maximum longevity is unknown. However, minimum ages of the oldest animals in one sample are two males 38+ years and a female 35+ years. Lifespans of 25 and 30 years have been estimated. The lifespan of males has been estimated at 40 years and lifespan of females at 32 years. Reproductive rate: The reproductive rate is the fraction of calves produced annually in the total population, without correction for mortality. Estimates are calculated from the percentage of calves seen during surveys and also from the crude birth rate and the proportion of mature females in the population. The crude birth rate is the average annual rate at which a mature female will produce a calf. Estimates of reproductive rate from calf counts range from 0.06 to 0.14 and estimates from the annual rate of calf production range from 0.09 to 0.13. Estimates of the reproductive rate derived from carcasses use the equation r=bp, where r is the reproductive rate, b crude birth rate, and p is the proportion of mature females in the population. There are no valid estimates of net reproductive rates in beluga whale populations because current data do not provide a basis for estimating natural mortality. From a sample of 265 females, crude birth rate was estimated at 0.33 Life History - 2 (DRAFT) - Life History Species whale, beluga Species Id M120107 Date 26 AUG 96 (range=0.31-0.35) and the reproductive rate at about 0.11. This matches the reproductive rate estimate of 0.10-0.12 adopted by the International Whaling Commission Subcommittee on Small Cetaceans. Possible sources of error in determining reproductive rates from calf counts include difficulty in seeing smaller, darker-colored calves; difficulty in distinguishing calves from yearlings; and possible segregation of population components. Sources of error in calculating reproductive rates extrapolated from the proportion of females in the population and reproductive rates of adult females include biases in the sex and age composition of the sample collected, and the difficulty of assessing the reproductive status of adult females. Lactation: The duration of dependent nursing may be considerably shorter than the total nursing period. The duration of dependent nursing has been reported to be 12-18 months. From observations of larger calved attending lactating females, it was concluded that lactation ends when the calf is between 1 and 2 years old. Other researchers estimated that lactation lasts about 23 months and ranges from 18 to 32 months. Weaning occurs well after the first solid food is taken. Lactation does not always inhibit ovulation and pregnancy. Milk has been found in several pregnant females. About 25% of sexually mature females become pregnant during lactation, presumably 10 months after having given birth. Gestation: Gestation is estimated to last 14-15 months. This estimate is from measurements of fetuses and neonates from Cumberland Sound and Hudson Bay. Assuming a 14.5 month gestation, and knowing that the peak of births is from mid-June to mid-July, mating should peak in April. However, specimen analyses and observations of behavior during spring suggest that most mating takes place before April. Of 13 newly pregnant females collected from late April and early May harvests, 9 had fully developed corpora lutea, 2 showed signs of recent or imminent ovulation, and 2 had embryos. No females obtained in June or July showed signs of recent ovulation. Of the 34 sexually mature males, only 2 (both taken in mid-June) were in breeding condition. the other 32 males (including 14 obtained in April and May) were in early to mid-spermatogenic retrogression. Thus, although a small proportion of belugas may be in breeding conditon in and after late April, the majority appear to breed earlier. The peak mating time may be between late February and early April. If so, either peak calving occurs prior to mid-June, or the gestation period is longer than supposed, or belugas have delayed implantation. The possibility of delayed implantation cannot be ruled out. Outside Alaska, the breeding period has been estimated from data on peak calving dates and gestation period. Peak mating is estimated to occur in Cumberland Sound in mid-May, in Hudson Bay between mid-April and mid-May, and in the St. Lawrence River estuary from April to June. Along the Siberia coast conception occurs in early May. These dates parallel mating dates which would be estimated from calving and gestation information from Alaska, and are derived in a similar manner, but they do not closely correspond with peak mating periods determined from analysis of beluga reproductive status. NATURAL MORTALITY: Data on natural mortality rates of belugas are extremely Life History - 3 (DRAFT) - Life History Species whale, beluga Species Id M120107 Date 26 AUG 96 limited. From comparison with mortality data for the pilot whale (Globicephala melaena), a natural annual mortality rate of 0.045 was suggested for belugas. The validity of this comparison, however, is questioned by the Intermational Whaling Commission Subcommittee on Small Cetaceans. Relationships between mortality, maximum length, and lifespan for various species of cetaceans were calculated. Generally, there is a high correlation among these factors in cetaceans. On the basis of body length and lifespan the natural annual mortality rate for belugas should be slightly in excess of 0.10. However, this seems too high, given that (1) recruitment appears to be in the range of 0.09 to 0.12; (2) these animals have been harvested for food for centuries, in some areas without notable declines; and (3) there are substantial losses due to ice entrapment which, although it is a cause of natural mortality, bears no likely relationship to body length and lifespan ratios. It seems unlikely that populations could have sustained harvesting and ice-entrapment with such a narrow margin of gain between natural mortality and recruitment rates. Causes of natural mortality among beluga populations include entrapment in ice and predation by killer whales (Orcinus orca) and polar bears (Ursus maritimus). Entrapment in ice has been attributed primarily to failure of belugas to migrate to open water prior to freezup in autumn, and to deep penetration of whales into pack ice during the spring migration. Recent evidence from the Soviet coast suggests that mortality due to entrapment could, at times, significantly affect the population size. However, entrapment does not always result in mortality. Predation by humans and polar bears is frequently associated with ice entrapment and is often the immediate cause of death. In some cases the entrapped whales might have otherwise survived; in others they might not have. In recent years trapped whales used by hunters have been included in catch statistics and thus death statistics from harvests and entrapment are not separable. In fact, Native settlements may have developed in areas close to locations of frequent ice entrapment. From the few sightings of entrapment in areas far from human settlements, despite increased observation oppotunities in recent years, it was concluded that the proportion of entrapped whales that die undetected is small. Polar bears occasionally prey on belugas, commonly in ice traps. Although belugas are vulnerable to polar bear predation during certain periods, even when in shallow estuaries, they are not a primary food of polar bears. Killer whales also prey on belugas. Beluga whale range in the north may have evolved partly to avoid killer whale predation. Although killer whales occur in areas inhabited by belugas, they probably cannot give birth because of the cold water, or enter heavy ice because their dorsal fin renders them poorly adapted for breaking through ice. Although parasites and their associated lesions have been described, their role in beluga mortality is not known. Of 9 belugas collected from the Churchhill area of Hudson Bay, Canada, 8 were heavily infested with Pharusus pallasii, a parasite of the hearing organs. The presence of P. pallasii in the cerebrospinal fluid suggested that infestation could produce erratic behavior, due to changes in spinal fluid pressure. These data are only preliminary, and no similar studies have been done in Alaska. Parasites are not known to cause death. EFFECTS OF HUMAN ACTIVITIES: Harvests and incidental catches have the most clear and direct effects on beluga whales. Threats posed by development and Life History - 4 (DRAFT) - Life History Species whale, beluga Species Id M120107 Date 26 AUG 96 other human activities are more difficult to evaluate, yet these activities are expanding rapidly and have the potential of significantly affecting belugas. Residents of some coastal areas assert that changes in beluga distribution in the past decades are due to habitat alteration from human activity. In particular, continued oil and gas development, development of mines and hydroelectric plants, increased commercial fishing, and changes in hunting and recreational activities will likely degrade water quality, reduce prey, and increase noise pollution. Information on the sensitivity of beluga whales to these factors is limited. Because they form dense aggregations, their population as a whole can be significantly affected by local degradation of the environment. The effects of habitat changes on belugas could take various forms, including direct mortality, displacement, increased stress or illness, changes in reproductive success, changes in fat reserves or energy expenditures, and changes in the ability to communicate, navigate, or locate prey. Marine mammals may abandon areas as their habitat deterioates, or they may exhibit high site tenacity. Belugas return to estuaries in northern Quebec every year despite severe overhunting that has reduced population levels. Although belugas in Kotzbue Sound are intensely hunted, and in the Mackenzie Delta are hunted and exposed to dramatic increases in vessel traffic and industrial noise, they return each year. In other areas where vessel traffic has increased, belugas have disappeared. Thus, tenacity may better reflect the critical value of the habitat than the level of human-imposed stress *125*. LIFE HISTORY CODES - Foraging Sites: Standing water - Limnetic zone Foraging Sites: Standing water: Profundal zone Breeding/Spawning Season: March Breeding Spawning Season: April Breeding/Spawning Season: May Breeding/Spawning Season: June Breeding/Spawning Season: July Breeding/Spawning Season: August Breeding/Spawning Season: September Display Site: Water Spawning Site: Standing Water Gestation/Incubation Period: Greater than 8 months Average Number of Offspring/Reproductive Effort: 1 Parental Care of Young: Female Dispersion: Clumped REFERENCES FOR LIFE HISTORY- 125 Life History - 5
                           (DRAFT) - Management Practices
                                Species whale, beluga
                                 Species Id M120107
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

RESULT MANAGEMENT PRACTICE Beneficial Maintaining undisturbed/undeveloped areas Beneficial Controlling pollution [thermal, chemical, physical] Beneficial Other management practices [specified in comments] REFERENCES FOR BENEFICIAL MANAGEMENT PRACTICES - 125 COMMENTS ON MANAGEMENT PRACTICES - EFFECTS OF HUMAN ACTIVITIES: Harvests and incidental catches have the most clear and direct effects on beluga whales. Threats posed by development and other human activities are more difficult to evaluate, yet these activities are expanding rapidly and have the potential of significantly affecting belugas. Residents of some coastal areas assert that changes in beluga distribution in the past decades are due to habitat alteration from human activity. In particular, continued oil and gas development, development of mines and hydroelectric plants, increased commercial fishing, and changes in hunting and recreational activities will likely degrade water quality, reduce prey, and increase noise pollution. Information on the sensitivity of beluga whales to these factors is limited. Because they form dense aggregations, their population as a whole can be significantly affected by local degradation of the environment. The effects of habitat changes on belugas could take various forms, including direct mortality, displacement, increased stress or illness, changes in reproductive success, changes in fat reserves or energy expenditures, and changes in the ability to communicate, navigate, or locate prey.*125* Primary beluga management concerns are maintenance of populations within the optimum sustainable range, protection of habitat, and prevention of interference with subsistence activities through diversion of belugas away from traditional hunting areas. Possible adverse human impacts include harvesting, noise pollution, contamination of habitat, net entanglement, and competition for-or depletion of-prey species as a result of commercial fishing. The most important management issue is to identify and protect critical habitat. It is also important to identify management units (stocks), determine abundance and productivity (sustainable yield) of each, and determine the number of animals being taken from each stock and whether there are any problems with current harvest levels. In many cases, basic information about distribution and abundance is needed before the more complex questions of causal relationships between human activities and belugas can be answered. Current subsistence harvests in Alaska and western Canada are below historic levels. In the absence of information on natural mortality, population levels, and trends, it is not possible to accurately quantify whether current harvest levels are sustainable. However, with the possible exception of the Yukon Delta, Norton Sound, and Cook Inlet (about which data are too poor to allow a reasonable appraisal), there is no indication that current harvest levels cannot be sustained. Management Practices - 1 (DRAFT) - Management Practices Species whale, beluga Species Id M120107 Date 26 AUG 96 Several habitats may have already been partially abandoned by belugas. Disturbance from human activity, primarily vessel traffic and hunting, appear to be the major reasons. This is of much concern to Kotzebue Sound residents and has been mentioned as a problem along much of the northern and western Alaska coasts. It is likely that noise pollution will continue to increase, and may become an important subject for study in protecting habitat. In one instance, overfishing of prey species by humans may have caused a temporary distributional change. This does not currently appear to be a major area of concern; however, so little is known about winter feeding that assessing impacts of foreign-fleet harvests on availability of prey is difficult. Net entanglement does not appear to be a problem anywhere. Entanglement could cause mortality in Cook Inlet or Bristol Bay in the future; however, little is known about the ability of belugas to avoid gear. Entanglement in Cook Inlet should receive special attention because the beluga population there is small and isolated. The effects of contaminants on belugas can probably be best studied in the St. Lawrence Estuary where waters are more polluted, and studies are already underway. No baseline data are available on contaminant concentrations in belugas off Alaska *125*. Management Practices - 2
                                   (DRAFT) - References
                                  Species whale, beluga
                                    Species Id M120107
                                      Date 26 AUG 96



     

References

125* Lentfer, J.W. (ed.). 1988. Selected Marine Mammals of Alaska, Species Accounts with Research and Management Recommendations. Marine Mammal Commission Washington, D.C pp 275. 166* American Cetacean Society. 1988. ACS Whale Fact Pack. American Cetacean Society San Pedro, CA. 178 * U.S. Department of Commerce. 1988. Marine Mammal Protection Act of 1972 Annual Report. National Oceanic and Atmospheric Administration (ed.):38. (Note Ref. 178 [or 179] is Ref. 195 in Marine References.) References - 1