(DRAFT) - Taxonomy
                                   Species pinfish
                                Species Id TSNL010605
                                   Date 26 AUG 96



TAXONOMY

NAME - pinfish OTHER COMMON NAMES - bream, pin perch, sand perch, salior's choice, spined bream, chopa espina (Sp.), Sargo salema (Sp.), Sar saleme (Fr.), poisson beurre (Cajun) and butterfish ELEMENT CODE - CATEGORY - Fish PHYLUM AND SUBPHYLUM - , CLASS AND SUBCLASS - Osteichthyes, ORDER AND SUBORDER - Perciformes, FAMILY AND SUBFAMILY - Sparidae, GENUS AND SUBGENUS - Lagodon, SPECIES AND SSP - rhomboides, SCIENTIFIC NAME - Lagodon rhomboides AUTHORITY - Linnaeus TAXONOMY REFERENCES - 266 Taxonomy - 1
                                  (DRAFT) - Status
                                   Species pinfish
                                Species Id TSNL010605
                                   Date 26 AUG 96



STATUS

Coded Status Commercial/industrial Migrant Commercial/bait See Comments Commercial/consumption REFERENCES FOR STATUS - 266 and 267 COMMENTS ON STATUS - Because of its small size, the pinfish lacks widespread appeal. If prepared while fresh, however, the fish is acceptably palatable. Commercial catch statistics include the pinfish in the unclassified or industrial fish category. Pinfish has a potential value as a baitfish and is preyed upon many species of commercial and sport fishes. It is also a potential source of fish meal. Pinfish have been used extensively as test fish in pesticide bioassays and physiological experiments *266*. The pinfish has little food value, is edible when cooked whole, or has a fine flavor. Anglers have condemned pinfish for stealing bait from their hooks. Pinfish caught by anglers for food and bait along the Mississippi coast contributed less than 17% of the total catch in a July through December sportfishing survey in the Mississippi Sound. Most of the pinfish caught by headboat fishermen in Texas coastal bays and the Gulf of Mexico are included in the sport catch. Pinfish are not listed in commercial fishery statistics because they are combined with unclassified species or industrial fish. They are caught incidentally in gill nets, trammel nets, beach seines, traps, hook and line, and purse seines in the menhaden fishing off the Mississippi River. Pinfish yield a high grade of oil but few, if any, are now used for that purpose. Gillnet selectivity for pinfish has been reported from St. Andrew Bay, Florida *266*. Pinfish frequents grassy bays in warm weather and migrates to deeper water in the colder months *267*. The pinfish has a reputation as a "bait stealer" because of its nibbling habit. It is commonly used for live bait, especially for tarpan. Pinfish is edible but rarely reaches sufficient size to warrant keeping unless it is cooked whole *267*. Status - 1
                               (DRAFT) - Distribution
                                   Species pinfish
                                Species Id TSNL010605
                                   Date 26 AUG 96



DISTRIBUTION

Distribution - 1
     

HABITAT ASSOCIATIONS

HABITAT - aquatic marine REFERENCES FOR HABITAT - 266 and 267 LAND USE - Water Bays and Estuaries REFERENCES FOR LAND USE - 266 and 267 NATIONAL WETLAND INVENTORY CODES NWI NWICLS NWIMOD NWISPEC Marine Estuarine REFERENCES FOR NWI - 266 and 267 COMMENTS ON HABITAT ASSOCIATIONS - The pinfish is an abundant estuarine dependent fish that usually inhabits vegetated marine bottoms, rocky reefs, jetties, and mangrove swamps. It is so abundant and predaceous that it is believed to alter the composition of estuarine epifaunal seagrass communities. When the larvae are about 11 mm long, they begin moving into the estuaries. Juveniles migrate into estuaries in the spring and summer. Juvenile pinfish in northwest Florida marshes first appeared in estuaries in March. Small juveniles were most abundant in vegetated, shallow flats of estuaries. Juvenile occasionally have been collected in fresh waters. Young pinfish rarely venture outside of seagrass-covered habitat into sand patches, except at night, when they inhabitat open sandy bottoms. Adult pinfish have been reported to be abundant in channels and vegetated flats, but they prefer open water. They also prefer vegetated substrate *266*. Pinfish frequents grassy bays in warm weather and migrates to deeper water in the colder months. Pinfish are abundant in inshore waters. They have been found to spawn in offshore waters. Small pinfish may be found far inside bays *267*. ANIMAL/PLANT SPECIES ASSOCIATIONS - Seagrass Ladyfish Porpoise Spotted seatrout Alligator gar Gulf flounder Streptococcus sp. Isopod (Lironeca ovalis) Haematozoan (Haemogreyarina bigemina) Speckled trout REFERENCES FOR SPECIES ASSOCIATIONS - 266 and 267 COMMENTS ON SPECIES ASSOCIATIONS - It is so abundant and predaceous that it is believed to alter the composition of estuarine epifaunal seagrass communities *266*. Habitat Associations - 1 Known predators on pinfish are ladyfish, porpoises, spotted seatrout alligator gar and gulf flounder *266*. Streptococcus sp. is the major disease of pinfish along the Alabama and Florida coast. Major parasites are the isopod Lironeca ovalis and the haematozoan Haemogreyarina bigemina *266*. Spotted seatrout and speckled trout are predators of the pinfish *267*. Habitat Associations - 2
                                (DRAFT) - Food Habits
                                   Species pinfish
                                Species Id TSNL010605
                                   Date 26 AUG 96



FOOD HABITS

TROPHIC LEVEL - CARNIVORE HERBIVORE/GRAZER OMNIVORE OTHER REFERENCES FOR TROPHIC LEVEL - 266 and 267 LIFESTAGE FOOD FOOD PART Adult Poaceae Not Specified Larva Crustaceans Not Specified Larva Copepods Not Specified General Malacostraca Not Specified Juvenile Malacostraca Not Specified General Detritus Not Specified General Plants Not Specified Juvenile Branchiopods Not Specified General Plants See Comments General Algae Not Specified General Osteichthyes Not Specified General Poaceae Not Specified General Molluscs Not Specified General Bivalve Molluscs Not Specified Important Plants Not Specified REFERENCES FOR GENERAL FOOD - 266 and 267 REFERENCES FOR IMPORTANT FOOD - 267 REFERENCES FOR ADULT FOOD - 267 REFERENCES FOR JUVENILE FOOD - 266 REFERENCES FOR LARVAE FOOD - 266 COMMENTS ON FOOD - In laboratory experiments, feeding intensity was greatest at a water temperature of 24 oC. Feeding stopped at temperatures above 35 oC and below 6 oC. Stomach evacuation time at 24 oC was 28 to 37 h. The consumption rate was about 0.05 calorie per milligram of fish per day. Pinfish change their diet as they increase in size and tooth structure changes. At 16 to 20 mm long, when notched incisiform teeth usually appear, juvenile pinfish fed on shrimp, mysids, and amphipods. The selection of amphipods by pinfish 16 to 80 mm long appears to be a function of macrophyte density. The five major ontogenetic stages in pinfish diets appears to be primarily a function of mouth size and changes in incisiform teeth. Pinfish demonstrated planktivory, omnivory, strict carnivory, and strict herbivory at different times, locations, and stages of development. An increase in mouth width and height with an increase in pinfish body size enables pinfish to capture a larger size of prey *266*. Food Habits - 1 (DRAFT) - Food Habits Species pinfish Species Id TSNL010605 Date 26 AUG 96 The intensity of pinfish predation on amphipods in seagrass communities in spring and summer probably limit amphipod abundance there. The consumption of plant material and detritus by pinfish contributes to the export of organic materials in estuaries, especially where eelgrass contributes up to 64% of the total primary production *266*. Pinfish are carnivores and herbivores. They eat seagrass, algae, macro- and microcrustaceans, mollusks, and fish. They are an a voracious feeder well noted for its bait-stealing activities. The stomach contents of 8 pinfish were examined. One of them contained razor clam shells and the other 7 contained plant material in the form of algae or grass. Lack of plant material on the Gulf beach probably accounts for the absence of the pinfish in that environment. Vegetable debris seems to be the most important food. Evidently this fish is largely a grazer *267*. COMMENTS ON ADULT FOOD - Adults feed on grass *267*. COMMENTS ON JUVENILE FOOD - Juvenile pinfish 20 to 80 mm long selectively feed on amphipods in Apalachee Bay, Florida. Pinfish, 15 to 19 mm long, in the Newport River estuary of North Carolina, fed more heavily near mid-day when tidal currents were low *266*. At 16 to 20 mm long, when notched incisiform teeth usually appear, juvenile pinfish fed on shrimp, mysids, and amphipods. The selection of amphipods by pinfish 16 to 80 mm long appears to be a function of macrophyte density *266*. Young feed on the bottom *267*. COMMENTS ON LARVAE FOOD - Larval pinfish feed mainly on calanoid copepods *266*. Food Habits - 2
                         (DRAFT) - Environment Associations
                                   Species pinfish
                                Species Id TSNL010605
                                   Date 26 AUG 96



ENVIRONMENTAL ASSOCIATIONS

G = General A = Adult LIM = Limiting RA = Resting Adult J = Juvenile FA = Feeding Adult RJ = Resting Juvenile BA = Breeding Adult FJ = Feeding Juvenile P = Pupae L = Larvae E = Egg RL = Resting Larvae FL = Feeding Larvae
LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Bottom Type [Aquatic]: Rooted aquatic vegetation G Bottom Type [Aquatic]: Specified in Comments G Coastal Features: Reefs G Coastal Features: Specified in Comments G Coastal Wetlands: Mangrove swamps G Coastal Wetlands: Specified in Comments BA Water Depth Preference: 100-200 ft. BA Water Depth Preference: Specified in Comments BA Water Temperature: Specified in Comments L Estuarine habitat zone: specified in comments J Estuarine habitat zone: specified in comments J Coastal Wetlands: Specified in Comments J Aquatic Vegetation [specified type]: Specified in comments J Bottom Type [Aquatic]: Sand J Bottom Type [Aquatic]: Specified in Comments A Aquatic Vegetation [specified type]: Specified in comments A Bottom Type [Aquatic]: Rooted aquatic vegetation A Bottom Type [Aquatic]: Specified in Comments A Substrate: Plants A Substrate: Specified in Comments A Aquatic Habitat Zonation: Open water [pelagic zone] A Aquatic Habitat Zonation: Specified in Comments J Water Depth Preference: Specified in Comments J Water Temperature: Greater than 27 degrees C J Water Temperature: Between 21-27 degrees C J Water Temperature: Between 15-21 degrees C J Water Temperature: Below 15 degrees C J Water Temperature: Specified in Comments FJ Water Temperature: Greater than 27 degrees C FJ Water Temperature: Between 21-27 degrees C FJ Water Temperature: Between 15-21 degrees C FJ Water Temperature: Below 15 degrees C FJ Water Temperature: Specified in Comments G Water Temperature: Greater than 27 degrees C G Water Temperature: Between 21-27 degrees C G Water Temperature: Between 15-21 degrees C G Water Temperature: Below 15 degrees C G Water Temperature: Specified in Comments J G Dissolved Oxygen: Specified in Comments Environment Associations - 1 (DRAFT) - Environment Associations Species pinfish Species Id TSNL010605 Date 26 AUG 96 LIFESTAGE ENVIRONMENTAL ASSOCIATIONS G Bottom Type [Aquatic]: Rooted aquatic vegetation G Bottom Type [Aquatic]: Specified in Comments G G Estuarine habitat zone: bay G Estuarine habitat zone: specified in comments G Aquatic Vegetation [specified type]: Specified in comments J Water Depth Preference: Specified in Comments G Water Depth Preference: Specified in Comments G Aquatic Vegetation [specified type]: Specified in comments J Aquatic Vegetation [specified type]: Specified in comments J Estuarine habitat zone: bay J Estuarine habitat zone: specified in comments G Water Depth Preference: 10-25 ft. G Water Depth Preference: 25-50 ft. G Water Depth Preference: Specified in Comments G Water Temperature: Greater than 27 degrees C G Water Temperature: Between 21-27 degrees C G Water Temperature: Between 15-21 degrees C G Water Temperature: Below 15 degrees C G Water Temperature: Specified in Comments G Turbidity: Specified in Comments G Human Association: Specified in Comments G Bottom Type [Aquatic]: Rooted aquatic vegetation G Bottom Type [Aquatic]: Specified in Comments G G G G G G G G A A A A A A A A J J J J J J L L L L L L Environment Associations - 2 (DRAFT) - Environment Associations Species pinfish Species Id TSNL010605 Date 26 AUG 96 LIFESTAGE ENVIRONMENTAL ASSOCIATIONS L L FJ Aquatic Habitat Zonation: Specified in Comments BA Water Temperature: Specified in Comments REFERENCES FOR ENVIRONMENTAL ASSOC_ - 266 and 267 REFERENCES FOR ADULT ENVIRONMENTAL ASSOC_ - 266 and 267 REFERENCES FOR FEEDING ADULT ENVIRONMENTAL ASSOC_ - 266 REFERENCES FOR BREEDING ADULT ENVIRONMENTAL ASSOC_ - 266 and 267 REFERENCES FOR JUVENILE ENVIRONMENTAL ASSOC_ - 266 and 267 REFERENCES FOR FEEDING JUVENILE ENVIRONMENTAL ASSOC_ - 266 and 267 REFERENCES FOR LARVAE ENVIRONMENTAL ASSOC_ - 266 and 267 COMMENTS ON ENVIRONMENTAL ASSOCIATIONS - POLLUTANTS: In bioassays, pinfish were highly sensitive to the pesticide Antimycin A at 7 ppb, as well as PCB's and mirex. Petrochemical wastes from a Corpus Christi turning basin depressed respiratory rates of the pinfish and caused up to 10% mortality *266*. WATER TEMPERATURE: Pinfish in estuaries tolerate water temperatures of 10 to 35 oC. The upper temperature tolerance of pinfish is about 33 oC. In fall, pinfish migrate from estuaries to offshore spawning sites. Water temperature has been suggested as the major factor triggering emigration. Pinfish are relatively active at 7.6 oC. Large pinfish were killed on the east coast of Florida when surface water temperatures dropped to 4 oC. Larger numbers of pinfish died during a severe cold spell in Copano Bay, Texas *266*. Higher water temperatures increase erythrocyte abundance and hematocrit in pinfish. High temperatures increased hemoglobin concentration and red blood cell counts but decreased hematocrit and mean erythrocyte volume *266* Pinfish were collected in water temperatures of 13.8 - 27.5 oC. Another study found the fish in a range of 27.2 to 31.6 oC. Another study found them in 9.1 - 34.9 oC range *267*. Pinfish can tolerate lower temperatures. Pinfish may burrow in the bottom when exposed to cold, and presumably this habit has survival value *267*. SALINITY: Pinfish live in waters with salinities as low as 1 ppt and as high as 75 ppt, and, in Florida, some enter freshwater. In the northern Gulf of Mexico, pinfish tolerate salinities of 0 to 37.5 ppt *266*. Common all year round in moderate to high salinity waters. Pinfish have Environment Associations - 3 (DRAFT) - Environment Associations Species pinfish Species Id TSNL010605 Date 26 AUG 96 been found in salinities ranging from 25.0 - 32.3 ppt, with a mean of 29.3 ppt. Pinfish were caught in water ranging from 2.1 to 37.2 ppt salt. This fish is rather indifferent to the salinity of water. No relation between the size of the fish and the salinity of the water was detected *267*. Pinfish are known to enter freshwater. They are found in all salinities in summer. In winter, they die quickly when trapped in hypersaline waters *267*. Pinfish have taken below a salinity of 5.0 ppt *267*. In attempting to predict monthly average weight of the pinfish, regression analysis showed that salinity was the most improtant variable *267*. DISSOLVED OXYGEN: Respiration studies on pinfish weighing 0.013 to 240 g revealed lower rates of oxygen consumption in smaller fish and indicated that the relation between body size and respiration rate depended on the life history stage of the fish. The metabolism-weight regression (K) for fish of different weight was 0.99 for fish weighing less than 0.2 g, 0.15 for fish weighing 0 to 10 g, and 0.73 for fish above 10 g. Oxygen consumption rates were related also to temperatures and swimming velocities. Oxygen-carrying capacity of pinfish blood increases in response to lowered environmental oxygen, increased exercise, and increased salinity but does not compensate entirely for the increased respiratory demand caused by these changes *266*. LIGHT: Diel distribution of pinfish in experimental cages in a thermal effluent channel was directly related to low ambient light conditions governed by the time of day and turbidity. Pinfish fed during daylight *266*. SUBSTRATE: Pinfish prefer vegetated bottoms in Gulf of Mexico estuaries and along the southeast Atlantic coast. Most fin-clipped pinfish in mark-recapture studies inhabited grass flats until they migrated out of the estuary in late summer and fall. Pinfish are usually active foragers and make no attempt to conceal themselves, but can burrow into sand when startled. The abundance and type of submerged vegetation in estuaries depends on bottom type, turbidity, salinity, water temperatures, bottom slope, and tidal range. Nearly 50% of the estuarine bottoms along the southern Florida gulf coast are covered by submerged vegetation which may account for the high abundance of pinfish there. The abundance of submerged vegetation decreases northward and covers only 5% of the bottom in coastal waters of the Florida panhandle and the northern gulf coast. Of about 800,000 acreas of submerged estuarine vegetation along northern Gulf of Mexico coastline, 63% is in Florida and 31% in lower Texas *266*. PRESSURE: Pinfish exhibit a yawn reaction within 0.5 to 2.0 minutes in response to a 0.5 cm3 volume pressure increase; consequently, they can be expected to react to tidal changes to maintain buoyance drift. Pinfish adapted to gradient chambers established a base reference point to which they reacted within 0.1 sec to pressure changes of less than 0.5 cm water pressure. In thermal effluents pinfish detected and avoided supersaturated (total gases) surface waters by seeking deeper waters *266*. Environment Associations - 4 (DRAFT) - Environment Associations Species pinfish Species Id TSNL010605 Date 26 AUG 96 SOUND: When held out of the water, pinfish make a scraping sound by sliding the upper and lower incisor teeth, but no evidence of underwater sound attributable to pinfish were detected. The critical masking ration for auditory analysis falls within the band width measured for mammals, including man *266*. HABITAT: The pinfish is an abundant estuarine dependent fish that usually inhabits vegetated marine bottoms, rocky reefs, jetties, and mangrove swamps *266*. Pinfish frequents grassy bays in warm weather and migrates to deeper water in the colder months *267*. One of the most common inshore fishes except in the highly turbid brackis wters of western Louisiana, the pinfish is usually found around wharves and pilings and over grass flats *267*. Pinfish are commonly abundant in Thalassia beds. Changes in densities of large pinfish from area to area coincide with changes in grass density from area to area *267* DEPTH: In a study conducted in the spring and summer, all pinfish were taken in 3 and 6 fathoms, indicating a shallow Gulf habitat *267*. COMMENTS ON ADULT ENVIRONMENTAL ASSOC_ - Oxygen consumption of adult pinfish in laboratory respirometers increases with an increase of water temperature from 5 to 30 oC, especially above 20 oC. Metabolism increases with water temperatures up to 33 oC *266*. Adult pinfish have been reported to be abundant in channels and vegetated flats, but they prefer open water. They also prefer vegetated substrate. Pinfish were most abundant at night when tidal currents were low and where aquatic vegetation was abundant. Pinfish select cover or rest on the bottom at night under offshore platforms but they are free-swimming during the day *266*. Adults occur in 0.1 - 75 ppt. Juveniles and adults occur together in 2.1 - 37.2 ppt *267*. COMMENTS ON FEEDING ADULT ENVIRONTAL ASSOC_ - Food evacuation rates in pinfish decreased as water temperature declined from 30 oC to 12 oC; all feeding ceased at 6 oC *266*. COMMENTS ON BREEDING ADULT ENVIRONMENTAL ASSOC_ - Three large schools with ripe females were observed near the surface in waters about 38 m deep. Spawning location is probably related more to depth and water temperature than to distance offshore *266*. Pinfish breed at the time of falling or low temperatures *267*. COMMENTS ON JUVENILE ENVIRONMENTAL ASSOC_ - Oxygen consumption of juvenile pinfish in laboratory respirometers increases with an increase of water temperature from 5 to 30 oC, especially Environment Associations - 5 (DRAFT) - Environment Associations Species pinfish Species Id TSNL010605 Date 26 AUG 96 above 20 oC. Metabolism increases with water temperatures up to 33 oC. The swimming speed of juvenile pinfish increased up to 11 times their body length per second in relation to increased temperature *266*. Juvenile pinfish leave northwest Florida marshes before surface water temperatures reached 41 oC in June *266*. No relation was found between salinity or temperatures and the arrival of juvenile pinfish in estuaries. Juvenile pinfish are less able to withstand thermal shock if salinity also changes, i.e., they suffered from a temperature-salinity synergistic interaction effect. For example, at a salinity of 15 ppt, no mortality was reported for juveniles after being exposed 40 min to water temperature increased to 32 oC, but a mortality of about 20% was observed when the water temperature was 12 oC and the salinity increased from 20 to 30 ppt *266*. Juvenile pinfish approached sandy bottom substrate only at night, whereas larger pinfish rested at night in cover and on the bottom at offshore platforms. Juvenile pinfish fed most heavily at mid-day *266*. Juveniles migrate into estuaries in the spring and summer. Juvenile pinfish in northwest Florida marshes first appeared in estuaries in March. Small juveniles were most abundant in vegetated, shallow flats of estuaries. Young pinfish rarely venture outside of seagrass-covered habitat into sand patches, except at night, when they inhabit open sandy bottoms *266*. Juveniles usually inhabit the shallow estuaries but when surface water temperatures exceed 32 oC, most seek the deeper, cooler water of channels *266*. Young are found in grass flats in all Texas bays and in the Gulf of Mexico. Small pinfish may be found far inside bays *267*. Juveniles occur in 11 - 75 ppt. Juveniles and adults occur together in 2.1 - 37.2 ppt *267*. COMMENTS ON LARVAE ENVIRONMENTAL ASSOC_ - Larval pinfish mortality is high at temperatures below 4 oC in North Carolina. Pinfish larvae withstood 12 oC temperature shocks at acclimation temperatures of 5, 10, 15 oC, but 18 oC was fatal *266*. No nocturnal-diurnal differences in catches of larval pinfish in experimental gear were reported *266*. In a study, larval pinfish were collected near the surface offshore from Beaufort, North Carolina. When the larvae are about 11 mm long, they begin moving into the estuaries *266*. Larvae occur in 1 - 45.3 ppt salinity *267*. Environment Associations - 6
                               (DRAFT) - Life History
                                   Species pinfish
                                Species Id TSNL010605
                                   Date 26 AUG 96



LIFE HISTORY

MORPHOLOGY/IDENTIFICATION AIDS: Dorsal spines XII, soft rays 11; anal spines III, rays 11; scales 65-70 in lateral line; 10 rows above and 17 rows below the lateral line. Twelve dorsal rays and 53 to 68 scales in the lateral line. Body oval and compressed. Mouth comparatively small, 2.75 to 3.1 times in head length; maxilla reaches only to below the anterior margin of the eye. Both jaws have eight broad, deeply notched, jaws have eight broad, deeply notched, incisorlike teeth anteriorly and have 2.5 rows of molarlike teeth laterally. A single small foward-directed spine precedes the 12 spines of the dorsal fin. The body is olivaceous above; the bluish-silver sides have yellow longitudinal stripes broader than the interspaces. A dark shoulder spot is near the origin of the lateral line. Six dark, diffused, vertical bars mark the body. Anal and forked caudal fins are yellow with broad light-blue margins. A montypic genus *266*. SPAWNING: The time and location of spawning of the pinfish described here are inferred by the abundance and location of ripe females and newly hatched larvae taken by experimental gear near the surface in offshore waters. For example, in one area off the Mississippi coast, three large schools of more than 1,000 pinfish each were observed near the surface in waters about 38 m deep. All females sampled with a cast net were ripe and may have been spawning. Spawning location is probably related more to depth and water temperature than to distance offshore. Small pinfish have been collected in January along the Texas coast near passes in the open gulf. Although spawning concentrations of pinfish were observed off the Mississippi coast in winter, spawning there peaked in the fall. Most studies in the gulf have indicated offshore spawning in the fall and early winter. In Florida coastal waters, most spawning is in February through March *266*. The spawning season has been well established as occurring in late fall and winter offshore. A study reports that the spawning season is from October to March. They may also spawn at the snapper banks. Ripe males have been taken in the Port Aransas vicinity in late October. The fish probably spawns in the open Gulf near the passes *267*. EGGS AND FECUNDITY: The diameter of pinfish eggs in different samples ranged from 0.90 to 0.93 mm and 0.99 to 1.05 mm. The eggs usually have a single oil globule and a very narrow perivitelline space. In the laboratory immature and infertile eggs sank to the bottom of finger bowls; whereas, fertile eggs were semibuoyant. Larvae that hatched after a 48-h incubation at 18 oC were 2.3 mm. Eggs with more than one oil droplet present did not hatch. A 157-mm long female collected in Florida in late November contained an estimated 90,000 eggs. A study estimated 7,700 to 39,200 (average 21,600) eggs in eight pinfish 111 to 152 mm standard length *266*. LARVAE AND POSTLARVAE: The yolk sac was visible 24 h after hatching and was completely absorbed by the time the larvae were about 2.7 mm long. The mouth began to develop after 96 h, when the larvae were about 3 mm long. Soft rays first appear when the larvae are 5 to 7 mm long and spines begin to differentiate when the larvae are 8 to 10 mm long. The caudal fin becomes rounded when the larvae are 8 to 10 mm, square-ended at 12 mm, and concave at 14 mm TL. Teeth are formed by the time the larvae are 10 mm long Life History - 1 (DRAFT) - Life History Species pinfish Species Id TSNL010605 Date 26 AUG 96 and scales first form when the larvae are 15 mm long. When the larvae are about 11 mm long, they begin moving into the estuaries. Larvae 11 to 18 mm long were collected from December through March in plankton tows at Dog Keys Pass off Horn Island in the Mississippi Sound and from late November through January near Cedar Key, Florida *266*. Larvae are widespread both alongshore and offshore in February and March, but no larvae have been found in May *267*. JUVENILES: Juveniles (15-100 mm long) migrate into estuaries in the spring and summer. When the fish are 16 to 20 mm long, notched incisiform teeth appear outside a row of conical teeth and gradually replace the conical teeth. When the fish are 30 mm long, the color pattern is similar to that of the adult, and when they are about 44 mm long, the caudal fin becomes more deeply forked and the pectoral fin increases disproportionately in length. Most pinfish become sexually mature at lengths of 80 to 100 mm. In the Mississippi Sound, first year juveniles begin to migrate into the estuaries in March and increase in abundance there until May. Juvenile pinfish in northwest Florida marshes first appeared in estuaries in March and increased in numbers until June when all juveniles moved offshore. Juveniles first appeared in January in the inshore waters of Cedar Key, Florida, and in January to March, juveniles had moved into Aransas Bay and Copano Bay, Texas. Small juveniles (20-80 mm long) were most abundant in vegetated, shallow flats of estuaries. Juveniles occasionally have been collected in fresh waters. Young pinfish rarely venture outside of seagrass-covered habitat into sand patches, except at night, when they inhabit open sandy bottoms *266*. ADULTS: Adult pinfish have been reported to be abundant in channels and vegetated flats, but they prefer open water. Another study reported that adult pinfish prefer vegetated substrate. Trawl catches of pinfish in Whitewater Bay, Florida, were highest at night when tidal currents were low and where aquatic vegetation was abundant. Pinfish select cover or rest on the bottom at night under offshore platforms near Panama City, Florida, but they are free-swimming during the day. Most pinfish in Florida mature in their second year of life (the smallest mature female was 128 mm long) and first spawn in third year (age 2), but pinfish in northwest Florida spawned late in their first and second year of life and all mature fish were 110 mm SL or longer. Most pinfish mature during the fall offshore spawning migration or at offshore spawning sites. Catches of pinfish with experimental fishing gear give some idea of their size range. Those caught in gill nets near Panama City, Florida, ranged from 115 to 240 mm FL and averaged 160 mm. In trawl samples in shallow water, only about 1% were longer than 100 mm SL and none exceeded 128 mm SL *266*. MIGRATION: After hatching in offshore waters in fall and winter, larval pinfish migrate into the estuaries where they grow in the summer. Larval pinfish exhibited higher relative abundance than diurnal abundance. Juveniles usually inhabit the shallow estuaries but when surface water temperatures exceed 32 oC, most seek the deeper, cooler water of channels. Pinfish migrate out of the estuaries in late fall to their spawning grounds where they congregate into size groups. Large pinfish left shallow flats when water temperatures dropped below 10 oC, whereas some juveniles remained Life History - 2 (DRAFT) - Life History Species pinfish Species Id TSNL010605 Date 26 AUG 96 inshore. Adults were reported to be abundant in January and February and in December through February in St. Andrew Bay, Florida *266*. GROWTH CHARACTERISTICS: Pinfish embryos develop rapidly in the egg. Yolksac larvae are 2.3 mm long 48 h after emergence and 2.9 mm long when their yolk sac is absorbed. The larvae begin to migrate shoreward when they are about 11 mm long and grow from 18 mm in March to 52 mm by June in northwest Florida. In northwest Florida, the daily growth of age 0 pinfish was 0.32 mm in spring, 0.23 mm in summer, and 0.1 mm in fall. Daily growth of age 1 fish was 0.32 mm in spring, 0.21 mm in the summer, 0.04 mm in the fall, and 0.02 mm in the winter. The annual daily growth rate was 0.12 mm. The growth increment was 65 to 110 mm at the end of their first year (age 0); 55 mm at the end of the second year (age 1), and 45 mm at the end of the third year (age 2). The annulus formed on scales in April of the second year of life (age 1). Age 1 pinfish in Texas were 103 to 143 mm long in May. Maximum lengths of pinfish reported in the literature were 400 mm and 343 mm; 365 mm TL in Louisiana; 330 mm and 245 mm in North Carolina; 250 mm in Texas-Louisiana; and 240 mm FL in Florida. Conversion factors calculated from 100 pinfish were standard length (SL) = 0.85 fork length, and 0.78 total length. A length-weight equation is calculated Log Wt(g) = -4.3734 + 2.9136 Log L(SL); and another equation is Log Wt(g) = -4.353 + 2.903 Log L(SL). Larval pinfish length-weight relationship was Wt(g) = 0.0089 TL (exp 2.81) *266*. The average length of pinfish is 6 to 8 inches but they sometimes reach 14 inches. They have been found up to 437 mm *267*. The smaller specimens in a study first came into the catch in January and continued to come in until March. They were from 13 to 33 mm long. A few large fish from 103 to 148 mm long were also taken in January and throughout the summer. The size of the fish increased from April to October. During the latter month the size of fish caught was from 63 to 203 mm. In May the length of the fish were found to be 103 to 143 mm, and, in June, from 43 to 103 mm. By July these two groups had practically fused and the smallest fish was 58 mm long. The upper size limit began to decrease in September, while the lower continued to increase and by November, the fish caught measured from 93 to 128 mm long *267*. FOOD HABITS: The high abundance of pinfish in many inshore coastal waters and estuaries is certain to have broad ecological effects on the aquatic flora and fauna. Pinfish are numerically dominant among fish in seagrass habitat in shallow subtidal area of Gulf of Mexico and the southeast Atlantic coast. The intensity of pinfish predation on amphipods in seagrass communities in spring and summer probably limit amphipod abundance there. The consumption of plant material and detritus by pinfish contributes to the export of organic materials in estuaries, especially where eelgrass contributes up to 64% of the total primary production. On the basis of routine metabolism, pinfish use 1.7% of the total yearly energy available to secondary consumers in the Newport River estuary in North Carolina. Pinfish are valuable forage for larger fishes in estuaries, and are useful as bait *266*. The pinfish is a grazer *267*. Life History - 3 (DRAFT) - Life History Species pinfish Species Id TSNL010605 Date 26 AUG 96 HABITAT: Pinfish are commonly abundant in Thalassia beds. Changes in densities of large pinfish from area to area coincide with changes in grass density from area to area. Juveniles are slightly nocturnal in its occurrence in the grass beds. This habit changed in larger individuals to a strong diurnal tendency *267*. OCCURRENCE: The fish was taken in least numbers in the winter and spring months, especially in December, and found to be most abundant in the summer and early fall, especially in August and September. The fish were not caught in Copano Bay from December to March inclusive and were scarce in Aransas Bay in December. The pinfish was taken only in the Gulf in November and January *267*. POPULATIONS: Populations of the pinfish were small in late fall, winter, and spring, with large populations occurring during the warm months of July, August, and September. Average water temperature and cloud cover were the most important variables in predicting monthly populations of pinfish. The pinfish was found to be more abundant durig the day than during the night *267*. POLLUTION: By means of multiple regression equations the observed oxygen consumption rates were related to body weights, temperatures, and swimming velocities for fish acclimated in both unpolluted and slightly polluted waters over the temperature ranges of 10-20 C and 20-30 C and at the single temperature of 20 C. Under normal conditions the pinfish seems to have quite high rates of oxygen consumption compared to temperate species. In all cases the slightly polluted water caused a depression of respiratory levels. Altered oxygen consumption levels are primarily under the influence of pollution and not salinity or oxygen levels. It is generally recognized that low dissolved oxygen levels associated with pollution depress locomotor activity and growth, with the relatively greater depression at the higher temperatures. For the smaller fish there is a rather uniform and linear rise in oxygen consumption with temperature for active and inactive levels and for polluted and unpolluted waters; while for the larger fish there is a much more obvious depression in the polluted waters at both the 30 C and 10 C extremes *267* The abundance of pinfish on a grass flat are of Redfish Bay, Texas, increases rapidly in late winter, reaches a spring peak and declines through the remainder of the summer and fall to low levels in winter. Pinfish are absent altogether during brief periods in mid-summer and mid-winter when temperatures exceed lethal levels. Nearby deeper channels provide refuge at these times. Metabolic rates of the pinfish are effectively temperature compensated. Metabolic depression is evident at temperatures near lethal limits and depression was observed at periods when movements to other more favorable areas occurred. The period of rapid growth corresponds to times of maximum pinfish density on the grass flats and growth proceeds at a low rate during winter *267*. LIFE HISTORY CODES - Breeding/Spawning Season: February Breeding/Spawning Season: March Life History - 4 (DRAFT) - Life History Species pinfish Species Id TSNL010605 Date 26 AUG 96 LIFE HISTORY CODES - Average Number of Offspring/Reproductive Effort: 1,00 Average Number of Offspring/Reproductive Effort: Grea Breeding/Spawning Season: October Breeding/Spawning Season: November Breeding/Spawning Season: December Breeding/Spawning Season: January Periodicity: Active in day Foraging Strategy: Grazing REFERENCES FOR LIFE HISTORY- 266 and 267 Life History - 5
                           (DRAFT) - Management Practices
                                   Species pinfish
                                Species Id TSNL010605
                                   Date 26 AUG 96



MANAGEMENT PRACTICES

Management Practices - 1
                                   (DRAFT) - References
                                     Species pinfish
                                  Species Id TSNL010605
                                      Date 26 AUG 96



     

References

266 * Muncy, Robert J. 1984. Species Profiles: Life histories and environmental requirements of coastal fishes and invertebrates (Gulf of Mexico) --Pinfish. Species Profile Series 82(11.26) (ed.). U.S. Fish and Wildlife Service Washington, DC:18. 267 * Texas System of Natural Laboratories. 1991. Species Profiles - Pinfish. (ed.). Texas System of Natural Laboratories Austin, Texas. Additional references available at the Austin Office. References - 1