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N x, y, z, t
The individual animal is:
a p, q, r, s, x, y, z, t
where p is sex; q is age class or stage; r is a dominance, behavioral, or health coefficient; x is a pregnancy coefficient (including zero); x,y,z is location in 3-D space (z expressing the timely quality of the map location for the animal); and t is time.
A population is the sum of the individuals.
A population resource is the population products or services aided or hampered by the number and characteristics (size, health, fur primeness) of the animals in the population. Thus, if n is the observed population, and f is a coefficient expressing enhancement due to density of animals and positive (or negative relations), then the population resource is:
Nt+1 = n + nf or n(1.0 + f)
Fundamental or basic models of population dynamics or change are:
Nt+1 = Nt + Ntr
Nt+1 = Nt + Nt(B-D) (B-D = birth minus death)
Nt+1 = Nt (1 + r) (difference equation)
Nt+1 = Nt + Mt (migration unit M each year)
Nt = No (1 + r)t (exponential)
Note that r, a rate of population change, may be considered to be B - D.
Plot the values of N over time. Plot an upper and lower constraint. Low numbers may be "minimum viable population."
Nt+1 = Nt +ct(B-D) (allometric model)
compare the above equation to S = a + CAz (the species-area curve)
Nt+1 = Nt + r Nt (1-(Nt/K)) (sigmoid)
Nt = Noert (logistic)
The previous population is a very key element of any population analysis.
A fundamental model recommended for use by managers, one highlighting negative feedback and one closely related to adaptive management, is:
Nt+1 = N* - (1-K)(Nt - N*)
Where K is the level of managerial control over the difference between where the system is now (N)and where it ought to be (N*) (the desired population based on stated objectives).
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Last revision January 17, 2000.